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-PHYTOLOGIA

_ Designed to expedite botanical pi blication

VoL. 2 f io August, 1941 No. 1

CONTENTS

c. L. Lunvert: Diagnoses of new American plaMismed Sie e mi oigie oce 2

Merritt: A new species of Premna from the Philippines ......+5

ON,

Mo.penxe: Novelties in the Eriocaulaceae and Verbenaceae ....06

Published by H. A. Gleason and Harold N. Moldenke
The New York Botanical Garden
Bronx Park, New York, N. Y.

Price of this number, 50 cents; per volume, $5.00 in advance

Vol. 1, No. 15, was issued January 28, 1941

PHYTOLOGIA

A journal designed to expedite botanical publication

Published by H. A. Gleason and Harold N. Moldenke
New York Botanical Garden, Bronx Park, New York, N. Y.

| Volume Two

DIAGNOSES OF NEW AMERICAN PLANTS -- I (a)
CO. Le Lunde 11

STRUTHANTHUS ESCUINTLENSIS Lundell, sp. nov.

Frutex epiphyticus omino glaber, ramlis subteretibus,
brunnescentibus. Folia petiolata, petiolo ad 1.5 cm. longo;
lamina coriacea, elliptica, 2.2--5 om. longa, 1.7--3.2 cm.
lata, apice rotundata, minute apiculata, basi rotundata,
nervis lateralibus inconspicuis, venulis obsoletis. Inflor-
escentiae subspicatae, axillares, solitariae, ad 12 cm. lon-
gee, floribus in spiculas trifloras subsessiles dispositis,
bracteolis acutis. Petala linearia, 6.5 mm. longa. Stylus
oa. 4.5 mm. longus, haud contortus. --MEXICO: Chiapas, Mt..
Ovando, alt. 1800 Me, ON tree, July 1--16, 1940, Eizi Matuda
4185 (Univ. Michigan Herb., type). --Allied to S. macro-
stachyus Lundell and S. belizensis Lundell.

CALLIANDRA SILTEPECENSIS lundell, sp. nov.

Arbor parva, 3--4 m. alta, ramlis subteretibus dense
hirtellis. Stipulae ca. 4 mm. longae. Folia petiolata, peti-
olo ad 2.5 em. longo, hirtello, rachi usque ad 5.5 cm. longa;
pinnis 5--7-jugis; foliolis chartaceis, 21--53-jugis, line-
ari-oblongis, ad & mm. longis, 1.5 mm. latis, sessilibus,
adpresse hirtello-ciliatis, apice acutis, basi obliquis.
Capitulae axillares, pedunculis ad 5.5 om. longis, parce
hirtellis vel glabris. Legumina glabra, ad 10.5 om. longa,
0.9 cm. lata, apice rotundata et minute apiculata, basi
stipitata. --MEXICO: Chiapas, Barranca Honda, Siltepec, riv-
erside, October--November, 1940, Eizi Matuda 4040 (Univ.
Michigan Herb., type).

2 PHYTOLOGIA Vol. 2, noe l

HELIOCARPUS BELIZENSIS Lundell, sp. nov.

Arbor, 13 om. diam., ommino minute rufo-glandulosa. Rami-
li paroe hirsuti, glabrescentes. Folia petiolata, petiolo
1.3--4.5 cm. longo; lamina integra, late ovata, ovata, vel
ovato~lanceolata, 7--13 cm. longa, 2.5--6.5 cm. lata, apice
caudato~acuminata, basi rotundata, serrulata, supra parce et
breviter hirsuta, subtus stellato—pilosa. Infructescentia
magna, rhachibue hirsutis. Pedicelli fructiferi 4.3--5 m.
longie Fructus crasse stipitatus. --BRITISH HOMDURAS: El
Cayo District, Vaca, on hilltop, Mer. 2, 1938, Percy H.
Gentle 2273 (Univ. Michigan Herb., type), vernacular name
moho. -~The clothing of minute red glands suggests H.
glanduliferus Robinson, but that species has sessile fruits.

HELIOCARPUS CUSPIDATUS Lundell, sp. nov.

Arbor, 20 cm. diam., ramulis parece et minute stellato-
puberulentibus. Folia petiolata, petiolo 2.5--7 cm. longo;
lamina ovato-lanceolata, 7--15 om. longa, 3.6--6.7 cme lata,
apice caudato~acuminata, basi late obtusa vel rotundata,
serrulata, glabrescentia. Inflorescentia parva, rhachibus
minute stellato—tomentulosis; pedicellis 5--6 mm. longis.
Sepala 4, linearis, 6 mm. longa, extus minute stellato-
tomentulosa. Petale 4, anguste spatulata, 3.8-—4 mm. longa,
baso breviter pilosa. Stamina 14 vel 16. Stylus bifidus, ov-
erio longior. Pedicelli fructiferi 3.5-—-4.5 mm. longi.
Fructus stipitatus. --BRITISH HONDURAS: El Cayo District,
Vaca, on hilltop, Mar. 4, 1938, Percy H. Gentle 2297 (Univ.
Michigan Herb., type), vernacular name moho; near Camp 6, on
hillside, Mar. 15, 1938, Gentle 2355. --From description H.
cuspidatus appears to be close to H. Donnell-Smithii Rose,
but differs in the narrower serrulate leaves with caudate-
acuminate tips, longer sepals, plumose hairs on faces of
young fruits, and in the longer fruit stipes. The flowers
appear to be pseudohermaphroditic which may account for the
differences in pedicel lengths.

HELIOCARPUS FLORIBUNDUS lamdell, sp. nov.

Arbor, 15 m. alta, 25 cm. diam. Ramli glabrescentes.
Folia petiolata, petiolo 3.5--11.5 cm. longo; lamina integ-
ra, late ovata vel suborbicularia, 10--22 cm. longa, 7--12.5
cm. lata, apice acuminata, basi rotundata vel subcordata,
serrulata, supra minute stellato-puberula, subtus parce tom
entosa. Infructescentia magna, usque ad 45 om. longa, rhach-
ibus fulvis, breviter stellato-tomentulosis. Pedicelli
fructiferi 2--4 mm. longi. Fructus stipitatus. --BRITI&H
HONDURAS: Belize District, Gracie Rock, Sibum River, Mar.
28, 1955, Percy H. Gentle 1534 (Univ. Michigan Herb., type),
vernacular name mountain moho. El Cayo District, Little Coc-
quericot, Belize River, Mare 27> 1933; Ce Le Lundell 4177 °

1941 Lundell, Diagnoses 3

Stann Creek District, 22 Mile, Feb. 17, 1952, W. A. Schipp
872, vernaculer name broad leaf moho. ~-Essential character-
istics of H. floribundus are as follows: leaf blades mimte-
ly appressed stellate-puberulent above and tomentose beneath,
huge panicles with branches and pedicels clothed with short
fulvous stellate tomentum, short pedicels, long fruit
stipes, faces of fruit rugose and persistently hirsute but
not bearing plumose hairs, and plumose bristles of fringe as
much as & mm. long. It is related to H. tomentosus Turez.,
which has smaller leaves and coarsely hirsute branchlets and
infructescence.

HELIOGARPUS GENTLEI lundell, sp. nov.

Arbor, 10 cm. diam., ramulis minute stellato-tomentulos-
is. Folia persistenter stipulata, stipulis 16 mm. longis vel
minoribus, petiolata, petiolo 4.5--14 om. longo; lamina sub-
orbicularia, 12--26 cm. longa, 9-17 com. lata, trilobata,
lobo terminali acuminate, lobis lateralibus acutis vel rot-.
undatis, basi cordate, serrulata, supra glabrescens, nervis
parce et minute stellato-puberulentis, subtus parce stellato
-puberulens, nervis breviter villosis. Inflorescentia magna,
rhachibus stellato-tomentulosis, pedicellis 4.5--4.5 m.
longis. Sepala 4, linearis, 5.5-—-6 mm. longa, obtusa, exter-
iora stellato—tomentulosa. Petala 4, spatulata, 4--4.5 m.
longa. Stamina 16. Stylus ad mediam bilobus, ovario duplo
longior. Fructus ignotus. --BRITISH HONDURAS: Belize Dis-
trict, Gracie Rock, Sibun River, on hilltop, Jan. 30, 1936,
Percy H. Gentle 1787 (Univ. Michigan Herb., type), vernacu-
lar name white mohe. --In the absence of fruits, the exact
relationship is difficult to determine. The persistent stip-
ules suggest an alliance with H. stipulatus Hochr. which,
according to description, has sordid-tomentose branches,
flowers almost sessile, tuberculate sepals, and the androgy-
nophore surrounded by a conspicuous pilose ring. H. Gentlei
differs in these and other significant characteristics. The
large 3-lobed leaves, short villous beneath along the prim-
ary and secondary veins, the minute gray stellate tomentum
of branchlets and inflorescence, and the nodose flowers, in
addition to the persistent stipules already mentioned, dis-
tinguish the species.

DAPHNOPSIS FLAVIDA landell, sp. nov.

Arbor parva, 5 m. alta, ramulis rubris, juvenilibus parce
strigosis vel glabrescentibus. Folia petiolata, petiolo ad 5
mm. longo; lamina glabra, subcoriacea, flavida, oblonge-
elliptica, 5.5--10 em. longa, 2--4 cm. lata, apice attenue-
ta, obtusa, basi acuta, nervis venisque prominulis et retic-
ulatis, nervis lateralibus utroque latere ca. 8. Inflores-
centiae 6 axillares vel laterales, paucifloras, pedunculis

A PHYTOLOGIA Vol. 2, no. l

minute et parce strigillosis, ca. 5 mm. longis, floribus
flavidus apice umbellatis, pedicellis minute et parce strig-
illosis, ad 2 mm. longis; perianthii tubus ca. 11 mm. longus, —
lobis 4, ovato-orbicularibus, 2--2.5 mm. longis, minute pub-
erulis, extus fere glabris. Ovarium glabrum. Styli 1.5 mm.
longi. —-MEXIO0O: Chiapas, Mt. Ovando, in forest, alt. 2000
me, July 1--16, 1940, Eizi Matuda 4157 (Univ. Michigan Herb.,

type).

EUGENIA ESOUINTLENSIS Lundell, sp. nov.

Arbor, 15 m. alta, glabra, ramulis gracilibus, internod-
iis elongatis. Folia petiolata, petiolo canaliculato, ad 7
mm. longo; lamina chartatea, flavida, obovata, obovato-ell-~
iptica, vel oblongo-elliptica, 5--10.5 om. longa, 1.7--4.7
cm. lata, apice subacuminata, acumine obtuso, basi acuta,
nervis lateralibus 7--l0-jugis, utrinque prominulis. Flores
breviter racemosi, racemis ad 5 mm. longis, fasciculatis,
glebris, axillearibus; pedicellis 4—7 mm. longis; bracteolis
triangulari-ovatis, ca. 0.8 mm. longis, parce puberulis. Se-
pala 4, late ovata, 1.2--1.5 ym. longa, parce ciliolata.
Petala suborbiculeria, cae 3.5 mm. longa. Stylus 4.5 m.
longus. ~-MEXICO: Chiapas, Finca Esperanza, Escuintla, Feb.
28, 1940, Eizi Matude 4144 (Univ. Michigan Herb., type).

VACCINIUM MATUDAI Lundell, sp. nov.

Frutex; ramulis rubris, juvenilibus parce pilosellis, mox
glabrescentibus. Folia petiolata, petiolo ad 2.5 mm. longo,
crasso; lamina sublucida, crasse coriacea, ovato-cordata,
2--4.5 om. longa, 1.5--3.5 cm. lata, apice obtusa, basi late
cordata, obscure crenulata, utrinque glabra costa ad basin
puberula excepta, venulis reticulatis. Inflorescentia rubra,
glabra vel parce pilosella, pauciflora, racemosa, ca. 2 com.
I-nga, pedicellis 3.5--5 mm. longis. Calyx glaber, calycis
lobi 5, late deltoidei, ca. 1 mm. longi, acuti. Corolla ur-
ceolata, ca. 7 mm. longa, extus glebra, lobis 5, ovato=
oblongis, ca. 1.5 mm. longis, obtusis, intus parce pilosis.
Filamenta pilosa. Antherae 5 mm. longae, ecalcaratae. Ovari-
um glabrum, ut videtur 10-loculare. Fructus ca. 5 mm. diam.
~-MEXICO: Chiapas, Barranca Honda, Siltepec, growing on
rocks, alte 2600 m., Oct.—-Nov., 1940, Eizi Matuda ord
(Univ. Michigan Herb., type). --Allied to V. Selerianum
(Loes.) Sleumer.

ARDISIA (Grephardisia) PAQUITENSIS lundell, sp. nov.

Ramuli crassiusculi, glabri. Folia petiolata, petiolo
marginato, ad 1.8 em. longo; lamina membranacea, glabra, in-
tegra, elliptica vel obovato-elliptica, 23.5--30 om. longa,
11--15.5 cm. lata, apice subabrupte acuminata, basi anguste-
ta. Inflorescentiae glabrae, terminales, ca. 3.5 cm. longae,

1941 Lundell, Diagnoses 5

bracteae foliolaceae. Pedicelli usque ad & mm. longi. Sepala
oblonga, 4.5--5 mm. longa. Petala elliptica, ca. 7 mm. lon-
ga. Stamina 4 m. longa, filamentis glandulosis, ca. 1.2 mm.
longis. Ovarium glabrum. -—-COSTA RICA: San José Province,
low hills above Rfo Paquita, alt. 5--50 m., Aug. 15, 1936,
GO. W. Dodge & V. Fe Goerger 9885 (Herb. Field Mus., No.
885,447, type). --A. paquitensis, allied to A. opegrapha
Oerst., is remarkable for its large entire leaves up to 50
em. long and 13.5 om. wide, and the small inflorescence
scercely 3.5 om. long.

A NEW SPECIES OF PREMNA FROM THE PHILIPPINES

Ee De Merrill

PREMNA ATRA Merrill, sp. nov. éPremnos.

Arbor parva, inflorescentiis exceptis glabra vel subglab-
ra, ramis pallidis, subteretibus, ramulis brunneis, glabris
vel leviter pubescentibus; foliis in sicco atris, utrinque
concoloribus nitidisque, oblongo-ovatis vel late lanceolato-
ovatis, integris, firmiter chartaceis, 8--15 cm. longis, 4--
6 om. latis, sursum angustatis, longe acuminatis, basi sub-
rotundatis, utrinque glabris vel supra ad costam nervosque
et subtus in axillis inferioribus breviter pubescentibus;
nervis primariis utrinque 5--6, perspicuis, utrinque plus
minusve elevatis, plerumque pallide brunneis, laxe arcuato-
anastomosantibus, reticulis primariis sublaxis, ultimis sub-
obseuris, subcatervis; petiolis 1.5--4 cm. longis, supra
_ gubplanis vel leviter canaliculatis, hic breviter pubescent-
ibus, ceteroquin glabris; inflorescentiis terminalibus, cor-
ymboso=cymosis, plerumque pedunculatis (pedunculo 1--2.5 cm.
longo), 5--7 cm. longis latisque, breviter subadpresse pub-
escentibus (praesertim partibus junioribus); floribus circi-
ter 5 mm longis, in sicco atris, pedicellis circiter 1 mm.
longis, bracteis oblongis, obtusis, circiter 2 m. longis,
bracteolis brevioribus vix 1 mm. longis; calycibus subobli-
quis, distincte 2-labiatis, cupuletis, glabris vel extus
parcissime breviter pubescentibus, labio majore 3-lobato,
minore 2-lobato, lobis late rotundatis, brevibus; corolla
extus glabra, intus villosa, 2-labiata, labio inferiore 5-
lobato, lobis centralibus ad 1.5 mm. longis, suborbiculari-

6 PHYTOLOGIA Vol. 2, now l
ovatis, rotundatis, lateralibus paullo brevioribus, labio
superiore late truncato-rotundato vel obscure retuso; ovario
globoso, glabro; stylis filamentisque glabrie; fructibus im-
maturis subglobosis, glabris, 4-locellatis, circiter 4 mm.
diametro.

PHILIPPINES: Luzon, Rizal Frovince, Mount Irig and Mount
lumaten, Bur. Sei. 41873 (type, herb. Arnold Arboretum)
42171 Ramos, February and April, 1923.

A species clearly belonging to the section Premos,
strongly characterized by its nearly glabrous, entire, long
acuminate, shining leaves, these and the inflorescences
characteristically black when dry. By the cheracters of Dr.
Lam's key to the Malaysian species it falls in the group
with Prema benguetensis Merr., a species totally different
in all respects.

NOVELTIES IN THE ERIOCCAULACEAE AND VERBENACEAE
Harold N. Moldenke

SYNGONANTHUS VAUPESANUS Moldenke, sp. nov.

Herba acaulis; foliis caespitosis reflexis mmerosis
linearibus obtusis utrinque glabris nitidisques pedunculis
gracillimis aggregatis bicostatis dense albido-tomentellis;
vaginis glabris.

Acaulescent herb; leaves basal, tufted, reflexed or. ap-
pressed to the ground, numerous, linear, 1--3.35 cm. long,
about 1 mm. wide at the middle, blunt at apex, glabrous and
shining on both surfaces; peduncles very slender, aggrega-
te, 4--6 or more per plant, 15--24 om. long, 2-costate,
slightly twisted, densely white-tomentellous throughout;
sheaths narrow, closely appressed, equaling or surpassing
the leaves, 1.5-—-3 em. long, slightly twisted, glabrous, o-
bliquely split at apex, the blade appressed and bluntish;
heads hemispheric, 3--9 mm. in diameter; involucral bract-
lets numerous, very conspicuous and showy, white, their
margins subhyaline, obovate or oblanceolate, the outermost
ones slightly stramineous, about 2.5 mm. long and 1.5 mm
wide, the inner ones about 4.5 mm. long and 2 m. wide,
rounded at apex, more or less naviculer-cucullate, glab-
rous; receptacle densely villous with tenuous translucent
hairs; steminate florets: sepals 3, separate, hyaline,
translucent, oblanceolate, about 2 mm. long and 0.5 m.
‘wide, rounded at apex, glabrous; petal—tube very pale-
stramineous, translucent, 1.3--1.4 mm. long, glabrous, amp-
liate and 3-lobed at apex, the lobes very short, rounded,

?

1941 Moldenke, Novelties 7

and involute; stamens 33; enthers white, versatile; pistill-
ate florets: sepals 3, separate, broadly elliptic, 1.8--1.9
mm. long, about 0.9 mm. wide, rounded at apex, hyaline and
translucent, glabrous; iggy 3, connate at apex, oblanceo-
late, about 1.9 mm. long and o4 mm. wide at apex, densely
pilose with long antrorse po hairs; style very short, in-
fundibular-eampliate toward apex, glabrous; stigmas 5; style-
appendages longer than the stigmas; ovary J-oelled, 3-seeded
The type of this species was collected by José Cuatrecas-
as (no. 6 at Yuruperi, alt. 220 me, about 350 km. above
Mita, Vaupes, Oolombia, on September 24, 1939, and is depos-
ited in the United States National Herbarium at Washington.
The species is obviously related to S. niveus (Bong.) Ruhl.

AEGIPHILA CUATRECASASI Moldenke, sp. nov.

Arbor parva; ramis percrassis tetragonis adpresso-puberu-
lis vel furfuraceis glabrescentibus valde medullosis; foliis
oppositis permagnis; petiolis crassis densissime adpresso-
puberulis; leminis chartaceis vel submembranaceis late ell-
ipticis utrinque puberulento-pulverulis, ad apicem acutis
vel breviter acuminatis, ad basin acuminatis.

Small tree, about 5 m. tall; branches very coarse and
stout, tetragonal, more or less densely appressed-puberulent
or furfuraceous with very minute sordid furf, glabrous in
age, marked with scattered corky elongated lenticels, very
medullose with large white pith; nodes slightly flattened;
principal internodes 3--5 om. long; leaves decussate-opposi-
te, very lerge; leaf-scars very large, prominent, and corky;
petioles stout, 3.5--5 om. long, very densely appressed-
puberulent with grayish-brown hairs; blades chartaceous or
submembranous, broadly elliptic, lighter and more grayish~-
green beneath, 32--35 cm. long, 15--17 om. wide, acute or
short-acuminate at apex, acuminate at base, entire, densely
puberulent-pulverulent with very minute appressed pulveru-
lence above, somewhat more conspicuously and densely puberu-
lent beneath with sordid yellowish or grayish puberulence;
midrib very steut, flat or slightly prominulous above, very
mach rounded-prominent beneath, decreasing rapidly in diam
eter as the apex is approached, censely puberulent; second-
aries slender, 13--20 per side, ascending, not much arcuate
except near the margins where they are arcuately joined,
flat above, prominulous beneath; veinlet reticulation rath-
er abundant, obscure or indiscernible above, the larger -
portions slightly prominulous beneath; inflorese-nce axil-
lary, glomerate, apparently borne on the older wood; flowers
not seen; peduncles none; fruiting-pedicels very stout, a-
bout 5 mm. long, densely furfuracevus—puberulent, verrucul-
ose; fruiting-ealyx very large and incrassate, cupuliform,
about 1 cm. long and 1.5 cm. wide, verruculose, glabrate,

8 PHYTOLOGIA Vol. 2, noe l

the rim subtruncate; fruit drupaceous, oblong, about 15 mm.
long, 12--14 mm. wide, glabrous, shiny, with a conspicuous
corky scar at the apex; seeds 4, elongate-oblong.

The type of this remarkable species was collected by Jose
Cuatrecasas (no. 8566) below Gabinete, alt. 2100—-2250 m.,
Quebrada del Rio Hacha, eastern slope of the Cordillera Ori-
ental, Caqueta, Colombia, on March 23, 1940, and is deposit-
ed in the United States National Herbarium at Washington. It
is obviously related to A. Gleasonii Moldenke and A. sessil
iflora Moldenke. :

AEGIPHILA HAUGHTII Moldenke, sp. nov.

Frutex gracilis; ramlis gracilibus sparsissime pilosulis
glabrescentibus; foliis oppositis; petiolis minutissime pub-
erulentis vel glabris; laminis membranaceis obovatis vel ob-
ovato-ellipticis longe acuminatis integris, ad basin cuneato
~attenuatis, utrinque minutissim puberulis glebrescentibus;
inflorescentiis axilleribus cymosis paucifloris.

Slender shrub, about 2 m. tall; branchlets slender, gray,
very sparsely pilosulous on the’ nodes and younger parts,
glabrous in age; nodes not annulate; principal internodes
1.5=-10 cm. long; leaves decussate-opposite, usually about 2
pairs clustered near the apex of the season's growth; petio-
les rather slender, 5--10 mm. long, very minutely puberulent
or glabrous; blades membranous, obovate or obovate-elliptic,
11—-24 om. long, 3.7--8.3 cm. wide, rather long-acuminate at
apex, entire, cuneate-attenuate at base, very minutely and
obscurely puberulent on both surfaces, glabrescent in age;
midrib slender, flat above, prominent beneath; secondaries
slender, about 10 per side, arcuate-ascending, prominulous
beneath and slightly so above, plainly anastomosing near the
mergins; veinlet reticulation very abundant, slightly prom-
imulous on both surfaces; inflorescence axillary, cymose;
cymes solitary in the uppermost axils, 5-~6.5 om. long, 3-4
ome wide, few- (about 7-) flowered, mich shorter than the
subtending leaves; peduncles very slender or subfiliform,
about 53.5 cm. long, glabrous or obscurely puberulent at a-
pex; pedicels filiform, 5--8 mm. long, glabrous; calyx cupu-
liform, about 2.7 mm. long and 5 mm. wide, glabrous, black=-
ening in drying, its rim truncate and entire; corolla hypo-
crateriform, white and rather showy, nigrescent in drying,
ites tube slender, about & mm. long, glabrous, its lobes 5,
oblong-lingulate, 4--5 mm. long, glabrous.

The type of this distinctive species was collected by
Oscar Haught (no. 2904) -- in whose honor it is named —— at
the foot of Cerro Cimalon, alt. about 50 m., on Hacienda
Vainillo, Guayas, Ecuador, on October 7, 1939, and is depos-
ited in the United States National Herbarium at Washington.
It cannot be confused with any other species in the group.

1941 Moldenks, Novelties 9

AEGIPHILA STEINBACHII Moldenke, sp. nove

Frutex vel arbor; ramlis tetragonis adpresso~puberulis;
sarmentis gracilibus obtuse tetragonis densissime tomentell-
4s velutinis; foliis oppositis; petiolis gracilibus; laminis
membranaceis ovatis acutis vel acuminatis integris, ad basin
acutis vel subtriumcatis, supra densissime velutinis, subtus
dense breviterque pubescentibus; inflorescentiis terminali-
bus paniculatis; cymis multifloris.

Shrub or tree, to 5 m. tall; branches tetragonal, often
decussately flattened, more or less appressed-puberulent;
twigs slender, obtusely tetragonal, very densely tomentell-
ous with grayish-brown tomentum, velvety to touch; nodes not
anmilate; principal internodes 1.5--5.5 cm. long; leaves
decussate-opposite; petioles slender, 5--8 mm. long; blades
membranous, uniformly green on both surfaces or somewhat
lighter beneath, ovate, 7—-12 om. long, 3--6.5 om. wide, a-
cute or acuminate at apex, entire, acute or subtruncate at
base, very densely velutinous with more or less subappressed
multicellular hairs above, densely short-pubescent beneath
with sordid-grayish hairs; inflorescence terminal, panicu-
late; peduncles and rachis densely sordid-tomentellous like
the branches; cymes small, abbreviated, 1.5--4 om. long,
many-flowered; bractlets subulate, to 7 mm. long, densely
strigose-tomentellous; pedicels very slemer, about 1 m.
long, densely appressed-pubescent; calyx infundibular, 3--4
mm. long and wide, rather densely appressed=-pubescent, its
rim deeply 4—-lobed, the lobes broadly triangular, about 1
mm. long, acute; corolla hypocrateriform, its tube narrow-
cylindric, about 4 mm. long, glabrous, its limb 4~parted,
the lobes oblong-lingulate, 2--2.5 mm. long; stamens 4,
long-exserteds filaments filiform, 7--& mm. long, glabrous.

The type of this species was collected by José Steinbach
(no. 3168) -- in whose honor it is named — at Bosquecitos
San Javier, Sara, alt. 450 m., Santa Cruz, Bolivia, on Nov-
ember 16, 1916, and is deposited in the Britton Herbarium at
the New York Botanical Garden. Thies collection was errone~
ously cited by me in Brittonia 1: 406 (1934) and Phytologia
l: oho (1937) as A. mollis H.B.Ke, which has a subtruncate
calyx-rim and therefore belongs to an entirely different
section of the genus. It is very probable that all the other
Bolivian specimens cited by me as A. mollis are also this
new species and that A. mollis does not occur in Bolivia.

ALOYSIA ALOYSIOIDES Loes. & Moldenke, sp. nov.

Frutex; ramis mediocriter gracilibus obtuse tetragonis
glabris suberosis; sarmentis brevibus parce pilosis; petiol-
is gracilibus parce pilosis; laminis membranaceis ovatis ob-
tusis vel rotundatis, ad basin truncatia vel subtruncatis,
crasse dentatis, subrevolutis, supra seabris, subtus puberu-

10 PH Y 7:0 b:0'G:i 2; Vol. 2, no. 1

lis; inflorescentiis axillaribus dense multifloris.

Shrub; branches medium-slender, obtusely tetragonal, gla-
brous, gray, with very large and elevated leaf-scars pro-
jecting 2--3 mm. in divaricate fashion from the branches,
corky; nodes not flattened nor annulate; twigs short, apar-
sely pilose with scattered hairs, the nodes often rather ob-
securely anmlate with a line of hairs; leaves decussate-
opposite; principal internodes 1-35 om. long; petioles
slender, 1-2 mm. long, sparsely scattered—pilose; blades
membranous, rather uniformly light-green on both surfaces or
somewhat lighter beneath, ovate, 1.5=—5 om. long, 1.5--2.5
cm. wide, obtuse or rounded at apex, truncate or subtruncate
at base or slightly prolonged into the petiole when young,
coarsely dentate from almost the base to the apex with
rounded broadly triangular teeth, the margins slightly revo-
lute, scabrous above, densely or sparsely puberulent be-
neath; midrib, secondaries, and veinlet reticulation conspi-
cusus on both surfaces, subimpressed above, prominulous and
dark beneath; inflorescence axillary; spikes 5--8.5 om.
long, about 1 em. wide in anthesis, densely many-flowered;
peduncles very slender, 2--3 om. long, rather sparsely
pilose-puberulent with spreading hairs like the twigs and
petioles; rachis more densely spreading-piloee; prophylla
lanceolate, 2--3 mm. long, long-acuminate at apex, attenuate
at base, pilose-ciliate; calyx about 1.5 mm. long and wide,
very densely villous; corolla-tube about 5 mm. long, glab-
rous outside, its limb about 3 mm. wide.

The type of this species was collected by August Weber-
bauer (no. 5206) below Surco, dept. Lima, Peru, alt. 1800
me, in February, 1909, and is deposited in the herbarium of
the Field Museum of Natural History at Chicago. The cheiro-

nym, Lippia aloysioides Loes., appears on the label.

ALOYSIA HERRERAE Moldenks, spe nove

Frutex; ramulis graciusculis tetragonis glabrescentibus;
petiolis gracillimis perbrevibus vel obsoletis pilosulo-
puberulentis; laminis chartaceis oblongis vel oblongo-ellip-
ticis acutis vel subacutis integris, ad basin acutis, supra
scaberrimis, subtus scabris et dense resinoso-punctatis; in-
florescentiis axillaribus terminalibusque.

Shrub; branches rather slender, tetragonal, stramineous
or brownish, glabrous and shiny in age, finely and very ob-
scurely scattered-puberulent on the youngest parts; nodes
annulate; principal internodes 2=-5.5 cm. long; twigs short
and very slender; leaves decussate-opposite; petioles very
slender, 1—2 mm. long or obsolete, pilosulous=puberulent;
blades chartaceous, sini Ser above, lighter beneath, ob-
long or oblong-elliptic, 0.9--4.5 om. long, 3--10 mm. wide,
acute or subacute at apex, acute at base, entire, very scab-

1941 Moldenke, Novelties ll

rous above with very short stiff bulbous~based hairs, scab-
rous bensath with shorter hairs and also densely resinous-
punctate; veinlet reticulation impressed above on smaller
leaves, obscure on larger ones, conspicuous but flat be-
neath; inflorescence axillary and terminal, the spikes abb-
reviated, 7--12 mm. long, many-flowered; peduncles 1--2 mn.
long, very slender, densely puberulent; calyx tubular, about
2 mme long, densely puberulent, not hirsute, its rim slight-
ly flaring and triangular-toothed; corolla-tube about 4 m.
long, densely short—pubescent outside, its limb about 4 m.
wide.

The type of this remarkable species was collected by
Fortunato L. Herrera (no. 1534) -- in whose honor it is
named -- at an altitude of 53000 m. in the Urubamba Valley,
Peru, in July, 1927, and is deposited in the herbarium of
the Field Museum at Chicago. A common name recorded by the
collector is "cedronsillo".

ALOYSIA LEPTOPHYLLA Loes.& Moldenke, sp. nov.

Frutex (7); ramis gracilibus sparsiuscule albo-hirsutis
medullosis; foliis sessilibus amplexicaulibus; laminis mem
branaceis ovatis ad apicem rotundatis, at basin cordatis,
crasse dentatis subrevolutis utrinque plusminus hirsutis;
inflorescentiis axillaribus terminalibusque dense multi-
floris.

Shrub (?)3 branches slender, tetragonal, rather sparsely
and irregularly hirsute with white hairs, more densely so
toward the apex, medullose; nodes not flattened nor anml-
ate; principal internodes 3--4 cm. long, not lenticellate;
leaves decussate-opposite, sessile and more or less clasping
at base; blades membranous, ovate, rather uniformly green on
both surfaces or somewhat lighter beneath, 2--5 cm. long,
1.5-—4 om. wide, rounded at apex, cordate at base and clasp-
ing the stem, coarsely dentate from base to apex with broad-
ly triangular teeth, the margins slightly revolute, more or
less densely hirsute above with weak whitish bulbous—based
hairs, somewhat hirsute and also more or less densely puber-
ulent beneath, the larger venation often subimpressed above,
prominulous beneath; inflorescence axillary and terminal;
spikes 5--8 cm. long, densely many-flowered, erect or ascen-
ding; peduncles very slender, 2--5 cm. long, rather densely
hirsute-pubescent like the branches, often surmounted by a
pair of foliaceous bracts about 1 cm. long and 7 mm. wide,
dentate, sessile, hirsute; bractlets large and conspicuous,
lanceolate, 5--6 mm. long, 1——-1.2 mm. wide, acuminate-atten-
uate at both ends, hirsute; calyx about 3 mm. long, densely
spreading—hirsute, its rim long-toothed with subulate-atten-
uate teeth; corolla-tube 5--6 mm. long, entirely glabrous
outside, its limb about 4 mm. in diameter.

12 PHYTOLOGIA Vol. 2, noe l

The type of this remarkable species was collected by Aug-
ust Weberbauver (no. 5374) somewhere in Peru between 1909 and
1914 and is deposited in the herbarium of the Field Museum
of Natural History at Chicago.

ALOYSIA MINTHIOSA Moldenke, sp. nov.

Frutex; ramis ramulisque grucilibus tetrazonis densely
pulverulento-puberulis resinoso-granulosis; internodiis abb-
reviatis; foliis sessilibus vel subsessilibuss; laminis sub-
coriaceis elliptico-ovatis supra pernitidis, ad apicem obtu-
sis, regulariter serrulatis subrevolutis utrinque dense pul-
verulento-puberulis et resinoso-glandulosis; inflorescentiis
axillaribus densissime multifloriss calyee non villoso.

Shrub, with a mint-like fragrance; branches and branch-
lets slender, obtusely tetragonal, more acutely so when
young, densely but obscurely pulverulent-puberulent and res~
inous-granular when young, less so in age; nodes not anmle-
te; principal internodes abbreviated, 5--20 mm. long; leaf-
scars small but very prominent, divaricate-raised; leaves
decussate-opposite, sessile or practically so; leaf=blades
subcoriaceous, uniformly bright~green on both surfaces, el-
liptic-ovate, very shiny above, 7--19 mm. long, 5--12 m.
wide, obtuse at apex,:uniformly serrulate from almost the
base to the apex with blunt and subrevolute teeth, densely
but obscurely pulverulent-puberulent on both surfaces, less
densely so above in age, ani resinous—glandular; midrib and
vendtion somewhat impressed above, flat beneath; inflores-
cence axillary, abundant, 4--13 cm. long, spicate, very
densely many-flowered; peduncles and rachis very slender,
densely puberulent, the former 4--15 mm. long; prophylla
numerous, lanceolate, 1--1.5 mm. long, acuminate, puberu-
lent; calyx about 2 mm. long, densely puberulent, not vill-
ous, its rim unequally 4-lobed; corolla about 5 mm. long,
its limb 4~lobed, the lobes subequal, the tube short, pubsr-
ulent within; stamens 4; anthers subsessile; style terminal;
stigma very minutely 2—lobed; ovary 2-celled, each cell with
a single basal ovule.

The type of this species was collected by J. Francis Mac-
bride & Featherstone (no. 2564) in a cliff crevice, alt. ab-
out 2000 feet, Yautan, Peru, on October 9, 1922, and is dep-
osited in the herbarium of the Field Museum at Chicago.

ALOYSIA NAHUIRE Gentry & Moldenke, sp. nov.

Frutex; ramulis elongatis gracilibus debilibus fere sub-
teretibus glabrescentibus; sarmentis substrigoso-puberulis;
petiolis gracillimis breviter pubescentibus vel strigosia;
laminis chartaceis lanceolato-ellipticis acutis vel breviter
acuminatis regulariter serrulatis, ad basin acutis, supra
scabris bullatis, subtus puberulis dense resinoso-punctatis.

1941 Moldenke, Novelties 13

Very slender shrub, 1--4 m. tall, with licorice-like od-
or; branches elongate, slender, weak, very obscurely tetrag-
onal or almost subterete, glabrous in age; twigs substrigose
-puberulent; leaf-scars large, concave, corky, rather prom
inent on the branches; nodes not annulate nor flattened;
principal internodes 1--5.5 cm. long; leaves decussate-oppo-
site; petioles very slender, 2--6 mm. long, short-pubescent
or strigose; blades chartaceous, bright-green on both surf-
aces, lanceolate-elliptic, 3--12.5 cm. long, 1.2--3.7 cme
wide, acute or short~acuminate at apex, acute at base, regu-
larly serrulate from almost the base to the apex, scabrous
above with very minute whitish bulbous-based hairs and bull-
ate, puberulent and densely resinous—punctate beneath; mid-
rib, secondaries, and veinlet reticulation deeply impressed
above, prominent beneath; inflorescence axillary, nutant,
53--4.5 om. long, densely many-flowered, hop-like; peduncles
filiform, densely strigose-puberulent; rachis densely
spreading-puberulent; bracts foliaceous, hop-like, elliptic,
about 8 mm. long and 4 mm. wide, acute or short-acuminate at
apex, rounded at base, densely silky-pubescent with long ap-
pressed whitish hairs, very conspicuous; calyx about 2.5 mm.
long, very densely hirsute; corolla-tube very slender, about
5 mm. long, sparsely spreading-pilose outside, ite limb ab-
out 3.4 mm. wide.

The type of this remarkable species was collected by HKaw-
ard Scott Gentry (no. 5721) in Croton Monte, in a coastal
thorn forest, Cerro Tecomate, west of Pericos, alt. 100 feet,
Sinaloa, Mexico, on February 27, 1930, and is deposited in
the Britton Herbarium at the New York Botanical Garden. Tea
is made locally from the foliage and the vernacular name is
"nahuire".

CARYOPTERIS INCANA var. BRACHYODONTA (Hand.—Mazz.) Moldenke,
comb. nov.

Qaryopteris tangutica var. brachyodonta Hand.—Mazz., Acta
Hort. Goth. 9: 68. 1934.

CITHAREXYLUM DRYANDERAE Moldenke, Spe nNOVe

Arbor; ramlis crassis acute tetragonis densiuscule far-
inaceo-puberulis; foliis oppositiss; petiolis crassiusculis
densiuscule farinaceo-puberulis in sicco corrugatis; laminis
coriaceis ellipticis acutis integris, ad basin plerumque ac~-
utis, supra glabris et nitidis, subtuse densely adpresso-
tomentellis, ad basin biglandulosis; inflorescentiis racemo-
so-spicatis multifloris, rhachide dense adpresso-furfuraceo.

Tree, about 5 m. tall; branchlets stout, acutely tetrag-
onal, rather densely farinaceous—puberulent with sordid-
brownish furf; nodes flattened, not annulate; leaf-scars
very large, ampliate, corky; principal internodes 2--5 cm.

¢

“14 .° EY 2 Orso eile Vole 2, noe 1

long; leaves decussate-opposite; petioles stoutish, 3--4.5
cme long, rather densely farinaceous-puberulent like the
branchlets, wrinkled-striate in drying, ampliate at base;
blades coriaceous, gray-green and shiny above, yellow-green —
beneath, elliptic, 11--19 om. long, 35--7 cm. wide, acute at
apex, entire, usually acute (sometimes rounded) at base,
glabrous above, densely appressed-tomentellous or furfurace-
ous with yellowish furf beneath, bearing two large and prom
inent glands parallel to the petiole at the very base; mid-
rib stout, impressed above, very prominent beneath; second-
aries slender, 10-—-15 per side, arcuate-ascending, flat and
rather inconspicuous above, very prominent and glabrous be-
neath; veinlet reticulation abundant, flat and often rather
inconspicuous above, the larger portions promimulous and
glabrous beneath; inflorescence racemose-subspicate, 6-14
em. long, many-flowered, solitary in the upper axils, erect;
flowers not seen; fruiting peduncles stout, 1--2.5 om. long,
more or less densely appressed-furfuraceous; rachis in
fruit stout and wrinkled-striate, densely appressed-furfura-
ceous with brownish furf; fruiting-pedicels stout and in-
crassate, about 1 mm. long or less, densely appressed-furf-
uraceous; fruiting-calyx indurated, 5--6 mm. long, 9-—-10 m.
wide, venose, glabrate, the rim irregularly lobed; fruit
drupaceous, fleshy, oblong-elliptic, 7--12 mm. long, 5--9
mn. wide, glabrous, shiny, red.

The type of this species was collected by Editha Dryander
(no. 2362) -- in whose honor it is named -—- at an altitude
of 2000 m. in El Valle, Colombia, in May, 1939, and is dep-
osited in the United States National Herbarium at Washington

CITHAREXYLUM ROSEI var. DURANGENSIS Moldenke, var. nove

Haec varietas a forma typica speciei recedit foliis minu-
te obscureque puberulis, pilis brevissimis adpressis.

This variety differs from the typical form of the species
in having its leaves only very minutely and obscurely puber=
ulent on both surfaces with very short appressed grayish
hairs.

The type of this variety was collected by Forrest Shreve
(no. 9123) on outwash plains near Pasaje, alt. 4650 feet,
Durango, Mexico, on August 23, 1939, and is deposited in his
herbarium at Tucson, Arizona. He describes the plant as a
shrub 6 feet tall, with its mature fruit red in color.

CITHAREXYLUM STEYERMARKII Moldenke, sp. nov.

Frutex; ramis tetragonis brunneis glabris nitidis; sarm-
entis mimte puberulis; laminis maturis subcoriaceis ellip-
ticis, acuminatis integris, ad basin acutis vel acuminatis,
utrinque glabris vel obscure pulverulento-punctatis; in-
florescentiis terminalibus racemiformibus dense mulfifloris.

1941 Moldenke, Novelties 15

Shrub, to 10 feet tall; branches tetragonal, brownish,
glabrous, medium-slender, shiny; youngest twigs minutely
puberulent; nodes annulate; principal internodes 1.5--10.5
em. long; leaves decussate-opposite; petioles stout, 5--18
mm. long, glabrous; leaf-secars large, corky, prominent, di-
vergent, 3--4 mm. long; blades chartaceous when young, sub-
coriaceous when mature ("firmly membranaceous" according to
the collector), elliptic, 6--18 om. long, 2--7.8 cm. wide,
acuminate at apex, entire, often slightly undulate along the
margins, acute or acuminate at base, glabrous or very mim-
tely and obscurely pulverulent-punctate on both surfaces,
very minutely and obscurely short-puberulent along the mid-
rib above; midrib slender, flat or subimpressed above, very
prominent beneath; secondaries slender, 7 or & per side, ar-
cuate-ascending, flat or subprominulous above, very sharply
prominent beneath, joined in many loops near or at the marg-
ins beneath; veinlet reticulation very abundant, conspicu-
ously promimuleus above, sharply prominulous beneath; in-
florescence terminal, racemiform; racemes simple or the
large ones branched at base, 6--15 cm. long, densely many-
flowered; peduncles (2—2.5 cm. long) and rachis slender,
minutely puberulent; pedicels very slender, 1-2 mm. long,
puberulent, in fruit to 5 mm. long and glabrescent; calyx
camparnulate, about 3 mm. long and wide, light, very shiny,
glabrous, the rim truncate and entire, short-ciliolate;
corolla hypocrateriform, sweet-scented, its tube 5 mm.
long, its lobes spreading, slightly squarrose, densely pub=-
escent within; fruiting-calyx slightly indurated, cupuli-
form, about 3 mm. long and 5 mm. wide, glabrous, light-
colored, very shiny, its rim entire and truncate; immature
fruit subglobose, about 5 mm. long and wide, glabrous,
shi ° °
The type of this species was collected by Julian A. Stey-
ermark (no. 31,453) on shaded cloud-forest slopes on top of
Volcan Quezaltepeque, 3--4 miles northeast of Quezaltepeque
at an altitude of 1500--2000 m., Chiquimla, Guatemala, on
November 8, 1939, and is deposited in the herbarium of the
Field Museum at Chicago. The type is in fruit. An isotype at
Chicago is in anthesis and is remarkable in having mech lar-
ger and thinner leaves, only chartaceous in texture and to
18 cm. long and 7.8 om. wide. The type has ite leaves sub-
coriaceous in texture and only 4.5--11.5 om. long and 1.8--
4.2 om. wide.

CITHAREXYLUM VALLENSE Moldenke, sp» nove

Arbor; ramulis percrassis acute tetragonis marginatis
dense puberulo—farinosis glabresocentibus; foliis oppositis;
petiolis crassis pulverulentomfarinosis glabrescentibus;
laminis coriaceis ovatis acutis vel breviter acuminatis in-

tegris, ad basin acutis, utrinque sparsissime pulverulentis
glabrescentibus, ad basin biglandulosis; inflorescentiis ax-
illaribus spicatis dnese multifloris; rhachide percrasso.

Tree, to 8 m. tall; branchlets very coarse and heavy,
sharply tetragonal, decussately flattened and ampliate at
the nodes, margined, densely pulverulent-farinose when
young, glabrescent in age; nodes plainly annulate with a
circumferential ridge; principal internodes 5-—-6 cme long
(at tips of branchlets); leaves decussate-opposite; petioles
heavy, about 5 cm. long, pulverulent-farinose, glabrescent
in age; blades coriaceous, dark-green above, lighter be-
neath, ovate, about 30 cm. long, 10--12 om. wide, acute or
short-acuminate at apex, entire, acute at base and there
bearing 2 large black glands parallel to the midrib, very
sparsely pulverulent along the midrib and larger veins on
both surfaces, glabrescent in age; midrib heavy, flat or
subimpressed above, very prominent beneath; secondaries
slender, about 15 per side, flat above, sharply prominent
beneath, arcuate-ascending, conspicuously joined in many
loops near the margins; veinlet reticulation obscure or in-
discernible above, prominulous beneath; inflorescence spic-
ate, axillary, 8-15 om. long, densely many-flowered; pedun-
cles (1--2 om. long) and rachis very stout, very densely
furfuraceous with sordid grayish or buff-colored furf, less
densely so in ages pedicels obsolete; prophylla tiny, scale-
like, 1--1.5 mm. long, densely furfuraceous or pulverulent-
farinose; calyx tubular, heavy and coriaceous, 6--S mm.
long, 4-5 m. wide, densely furfuraceous-farinose with sor-
did grayish or buff-colored furf; corolla white, barely pro-
truding from the calyx, its limb Sparted, the lobes ellip-
tic-lingulate, about 3 mm. long, densely pilose at base.

The type of this species was collected by Ellsworth Paine
Killip and Hernando Garcfa y Barriga (no. 33,940) in a dense
forest, San Antonio, west of Cali, near the summit of the
Cordillera Occidental, alte 1900--2350 m., between February
26 and March 2, 1939, and is deposited in the Britton Herb-
arium at the New York Botanical Garden.

DURANTA MACRODONTA Moldenks, sp. nove

Frutex; ramis gracilibus inermis plusmimus tetragonis
breciter adpresso-pubescentibus medullosis; foliis opposit-
is; petiolis gracillimis submrrginatis dense adpresso-pubes—
centibus vel strigosis; laminis membranaceis ovatis vel sub-
rotundis, ad basin et apicem breviter acuminatis, crasse
dentatis, utrinque sparsissime pilosulis; inflorescentiis
paniculatis foliosis multifloris.

Shrub, 4~—5 feet talls branches slender, unarmed, more or
less tetragonal, the younger parts decussately flattened at
the nodes, shortly appressed-pubescent with sordid-grayish

1941 Moldenke, Novelties 17

hairs, corky-lehticellate, brunnescent, medullose; nodes not
annulate, flattened; buds very densely villous-pubescent
with sordid-canescent hair; leaves decussate-opposite; peti-
oles very slender, 4--6 m. long, deeply canaliculate above,
submargined, densely appressed-pubescent or strigose with
sordid-canescent hair; blades membranous, uniformly dark=
green on both surfaces, brunnescent in drying, ovate or the
youngest subrotund, 2--5.8 om. long, 1--4.8 cm. wide, short=
acuminate at apex and base, coarsely dentate with broadly
triangularteeth from below the middle to the base of the
terminal acumination, very sparsely and obscurely pilosulous
on both surfaces with widely scattered hairs, more densely
so on the midrib; inflorescence supra-axillary and terminal,
the supre-exillary ones aggregated near the tips of the
brenches, forming a loose and leafy terminal panicle; race-
mes 9--18 cm. long, about 2 em. wide, rather densely many-
flowered, not secund, erect or recurved; peduncle slender,
1=-2.5 om. long, brunnescent, more or less appressed-pilose
like the branches; rachis similar, but more densely apprese~
ed-pilose with sordid-canescent hairs; bracts often leafy,
1--6 at the base of the racemes, ovate, 5-15 m. long, acu-
minate at apex and base, stipitate, pilosulous; prophylla
linear-setaceous, 3--5 mm. long, densely strigose, persist-
ent, conspicuous; pedicels about 2 mm. long, densely canes-
cent-pubescent; calyx tubular—campanulate, about 3 mm. long,
about 5 mm. wide at the apex, uniform, densely strigose with
sordid-canescent hairs like the branches and rachis, its rim
long-apiculate, the apiculations 1 mm. long and densely
strigose, erect; corolla blue, its tube about 5 mm. long,
very densely puberulent above the calyx, its limb about l
om. wide.

The type of this species was collected by Mohamed Nur bin
Mohamed Ghose in the Botanic Gardens at Singapore, Federated
Malay States, on October 21, 1924, and is deposited in the
herbarium of the Bailey Hortorium at Ithaca. It was cultiva-~
ted under the name of "Duranta plumieri Jacq."

DURANTA REPENS var. GRANDIFLORA Moldenke, var. nov.

Haec varietas a forma typica speciei recedit floribus
majoribus, corollae limbo usque ad 1.8 cm. diametro.

This variety differs from the typical form of the species
in its larger flowers, the corolla-limb being to 1.8 cm.
wide, its margins more or less crisped.

The type of this variety was collected by Frank F. Gander
in cultivation at 4681 50th Street, San Diego, California,
on May 28, 1936, and is deposited in the herbarium of the
Bailey Hortorium at Ithaca. Dr. Bailey states that the flow-
ers of this variety may attain a diameter of 3/4 inch (app-
roximately 2 com.). They are violet-blue in color.

18 mAs PHYTOLOGIA Vol. 2, no. l

DURANTA SPRUCEIL var. COLOMBIENSIS Moldenke, var. nov.

Haec varietas a forma typica speciei recedit calyce levi-
ter puberulo vel strigilloso.

This variety differs from the typical form of the species
in its calyx being only lightly puberulent or strigillose.

The type of this variety was collected by Brother Alberto
[Apolinar-Marfa 263] at San Pedro, Antioquia, Colombia, on
July 25, 1938, and is deposited in the herbarium of the
Field Museum of Natural History at Chicago.

LANTANA CAMARA ver. HYBRIDA (Neubert) Moldenke, comb. nov.
Lantana hybrida Neubert, Deutsch. Gart. Mag. 10: 98.
1857; Lantana chrysantha Schmdger ex Neubert, loc. cit., in

syne
This is the dwarf yellow-flowered garden form.

LANTANA CAMARA var. MULTIFLORA (Otto & Dietr.) Moldenke,
combe nOVe
Lantana multiflora Otto & Dietr., Allg. Gartenz. 9: 370.
1841.

LANTANA SCANDENS Moldenke, spe nov.

Frutex alto-volubilis; ramis gracilibus inermis acutius-
cule tetragonis hirsutulis; foliis oppositis nigrescentibus;
petiolis gracillimis glanduloso=punctatis hirsutulis; lamin-
is membranaceis ovatis breviter acuminatis, ad basin acutis,
regulariter arguteque serratia utrinque plusminus strigoso-
pilosis, maturitate supra scabris.

High-climbing vine; stems slender, unarmed, rather acute-
ly tetragonal, more or less abundantly hirsutulous with
stiff spreading short hairs and with shorter gland-tipped
hairs beneath; nodes annulate, usually marked with a denser
band of long-hirsute hairs; principal internodes 1.5--7.8
em. long; leaves decussate-opposite, nigrescent in drying;
petioles very slender, 4--6 mm. long, glandular-punctate and
rather abundantly hirsutulous; blades membranous, ovate, 3.5
--7 cm. long, 1.6--4.3 cm. wide, short~-acuminate at apex, a-
cute at base and often somewhat prolonged into the petiole,
regularly sharp~serrate from the apex almost to the base,
scattered strigose-pilose along the larger venation beneath,
more uniformly so on the lamina above with bulbous-based
hairs, causing the mature leaves to be quite scabrous above;
midrib very slender; secondaries very slender, about 7 pairs
inflorescence axillary, capitate; peduncles very slender,
1.5--6 cm. long, very sparsely hirsutulous with scattered
white hairs and more abundantly pilosulous with much shorter
gland-tipped hairs; heads hemispheric, 1--2.5 cm. wide,
many-flowered; bractlets rather large, often foliaceous, ac-
ute, variable in size and shape, the inner ones lanceolate,

1941 Moldenke, Novelties 19

4--5 mm. long, the outer ones spatulate or elliptic, to 10
mm. long and 3.5 mm. wide, more or less strigillose on both
surfaces; corolla "red and yellow or all red or all yellow",
its tube about 10 mm. long, very narrow, dense ly puberulent
outside, its limb 5--6 m. wide.

The type of this species was collected by George B. Hint-
on (no. 12,315) at Villa Victoria, Pto de Aire, alt. 1480
Mey Coalcomen, Michoacan, Mexico, on October 3, 1938, and is
deposited in the Britton Herbarium at the New York Botanical
Garden. The species is obviously closely related to L. Cama-
ra L. of the West Indies, but differs in being a high-
climbing vine and in its very thin nigrescent leaves, gland-
ular pubescence, and foliaceous bractlets.

LIPPIA ANTAICA Loes. & Moldenke, sp. nov.

Frutex; ramis gracilibus strictis virgatis acute tetrag-
onis adpresso-puberulis; foliis parvis; petiolis parvissimis
vel obsoletis; laminis firme charteceis vel subcoriaceis
flabelliformibus vel subrotundis (juventute obovatis), ad
apicem rotundatis, ad basin maturitate truncatis vel sub-
truncatis (juventute subcuneatis), reguleriter crenato~
serratis revolutis, supra scaberrimis bullatis, subtus dense
pubescentibus.

Shrub; branches slender, apparently strict and virgate,
acutely tetragonal, brownish, appressed-puberulent through-
out; principal internodes 2--45 cm. long; nodes nct annulate;
leaves decussate-opposite, small; petioles very slender, 1l--
2 mm. long or obsolete, densely short-pubescent; blades
firmly chartaceous or subcoriaceous (when mature), flabelli-
form or subrotund, obovate when immature, to abouy 2 cm.
long and wide when mature, with numerous smaller thinner and
more obovate ones in their axils, rounded at apex, subtrunc-
ate or truncate at base (the immature and smaller ones acute
or subcuneate at base), regularly and uniformly crenate-
serrate from base to apex with rounded teeth, the margins
revolute, very scabrous and bullate above, puberulent on the
venation, densely pubescent over the entire surface beneath;
midrib, secondaries, and veinlet reticulation deeply impres-
sed above, the larger parts prominent beneath; inflorescence
axillary, a pair at each node, capitate; peduncles very
slender, erect, 5--10 mm. long, densely appressed-puberulent
with greyish hair like the branches; heads densely many-
flowered, about 8 mm. long and 10 mm. wide; bracts ovate,
2.5--3 mm. long, subacute at apex, densely short—pubescent;
corolla 4—-5 mm. long, its limb about 3 mm. wide.

The type of this species was collected by August Weber-
beaver (no. 5918) somewhere in Peru between 1909 and 1914,
and is deposited in the herberium of the Field Museum at
Chicago. It is most unfortunate that the label on the type

20 PHYTOLOGIA Vol. 2, noe lL

specimen does not give the exact place and date of collec-
tion, but it was probably in the neighborhood of Ante in
Cuzcoe

LIPPIA FRANCENSIS Moldenke, spe nove

Frutex (7); ramis ut videtur simplicibus rectis gracilib-
us acutiuscule tetrahonis dense hirsutis glanduliferis velu-
tinis; foliis oppositis; petiolis gracilibus dense albo-
hirsutis; laminis coriaceis elliptico-subrotundis ad basin
et apicem rotundatis regulariter serratis utrinque dense
hirsutis subvelutinisque, subtus dense resinoso-punctatis.

Shrubby (7); stems apparently simple, erect, slender,
rather acutely tetragonal, densely hirsute with stiff whit-
ish hairs and shorter gland—-tipped hairs, velutinous to
touch; nodes anmlate with a band of denser hirsute hairs;
principal internodes 1--4.5 cm. long; leaves decussate-oppo-
site; petioles slender, 2--5 mm. long, densely white-hirsu-
te; blades coriaceous, elliptio-subrotund, somewhat lighter
beneath, 1.2--4 em. long, 1--2.8 cm. wide, rounded at apex
and base, regularly serrate from the apex almost to the base
with rounded revolute-margined teeth, densely hirsute on
both surfaces, subvelutinous and densely resinous-punctate
beneath; midrib, secondaries, and veinlet reticulation deep-
ly impressed above, very prominent beneath; secondaries 5 or
6 per side, ascending, not much arcuate; inflorescence axil-
lary, borne at the tips of the stems, usually 2 pairs, capi-
tate; peduncles very slender, 0.8-—3.5 om. long, very dense-
ly hirsutulous with stiff white gland-tipped hairs; heads
hemispheric, about 2 cm. in diameter, many-flowered; bracts
large, foliaceous, red, ovate, to about 10 mm. long and 8
mm. wide, blunt at apex, densely pubescent with short silky
mostly gland-tipped hairs, ciliate-margined; corolla yellow.

The type of this handsome species was collected by Guil-
herme Gehrt [Herb. Inst. Biol. S& Paulo 4037] in fields at
Franca, Sao Paulo, Brazil, on April 11, 1920, and is depos-
ited in the Britton Herbarium at the New York Botanical Gar-
den. It was originally distributed as L. lupulina Cham., to
which the species is closely related.

LIPPIA PINETORUM Moldenke, sp. nove

Frutex; ramulis graciusculis obtuse tetragonis obsolete
pilosis vel glabrescentibus; sermentis nigrescentibus pilos-
ie; petiolis gracilibus piloso-hirsutulis; laminis chartace-
is brunnescentibus ellipticis acutis vel obtusis, ad basin
acutis vel subacuminatis, regulariter serratis revolutis,
supra bullatis et scabris et hirsutulis, subtus sparse pilo-
sis; inflorescentiis axillaribus perspicue involucratis.

Shrub; branchlets rather slender, obtusely tetragonal,
grayish, obsoletely scattered-pilose or glabrescent; twigs

1941 Moldenke, Novelties 21

nigrescent in drying, more abundantly pilose with scattered
short spreading hairs; nodes annulate; principal internodes
2--6 om. long; leaves decussate-opposite; petioles slender,
5--10 mm. long, sparsely or rather densely pilose-hirsutul-
ous with stiff spreading hairs; blades chartaceous, dark-
green above, lighter beneath, brunnescent in drying, ellip-
tic, 2--7.5 cm. long, 1.6-—4 om. wide, acute or obtuse at a-
pex, acute or subacuminate at base, regularly serrate from
apex almost to base with blunt revolute-margined teeth, bul-
late and scabrous above, rather abundantly hirsutulous with
bulbous—based whitish hairs above, very sparsely and obscu-
rely pilose beneath; the slender midrib and 4--7 arcuate-
ascending secondaries impressed above, sharply prominent be-
neath; veinlet reticulation abundant, subimpressed above,
prominulous beneath; inflorescence simple, axillary, borne
at the tips of the twigs, capitate, conspicuously involucra-
te, about equaling or shorter than the subtending leaves;
peduncles very slender, 1--2 cm. long, densely hirsutulous
and brownish~pubescent with gland-tipped hairs; heads 5--17
mn. in diameter; involucral bractlets large and foliaceous,
ovate, to & m. long and 5 mm. wide, acute at apex, densely
short=pubescent with brownish gland-tipped hairs and
scattered-hirsutulous with longer white hairs.

The type of this species was collected by Eizi Matuda
(no. 3925) in pine land, Mt. Ovando, Chiapas, Mexico, be-
tween November 14 and 18, 1939, and is deposited in the
Britton Herbarium at the New York Botanical Garden. It was
originally distributed as L. cardiostegia Benth., to which
the species is obviously related.

LIPPIA TAYACAJANA Moldenke, sp. nov.

Frutex; ramis gracilibus tetragonis costatis, juventute
dense breviterque pubescentibus, senectute glabrescentibus;
internodiis valde abbreviatis; petiolis brevissimis vel ob-
soletis; laminis firme chartaceis oblanceolato-ellipticis,
ad apicem rotundatis vel acutis, ad basin cunsato-attenuat-
is, revolutis serratis, supra scabris substrigosis, subtus
dense tomentellis.

Shrub, about 1 m. tall; branches slender, tetragonal,
ribbed, densely short—pubescent when young, glabrescent in
age and then with peeling shreddy bark, brown, somewhat
twiggy below; nodes not annulate; principal internodes much
abbreviated, 1--3 cm. long; leaves decussate-opposite;
twigs very short, leafy; petioles slender, 1--2 mm. long
and densely short-pubescent or obsolete; blades firmly
chartaceous, uniformly gray~green on both surfaces, oblanc-
eolate-elliptic, 0.8--1.9 cm. long, 3--9 mm. wide, rounded
or acute at apex, cuneate-attenuate at base, revolute along
the margins and serrate from about the middle to the apex,

22 PHT? 0-50 G2 Vol. 2, noe l

- geabrous and substrigose above, densely tomentellous be-
neath; midrib and slender secondaries deeply impressed ab-
ove, very prominent beneath; inflorescence axillary, solit-
ary in each axil, 1--1.5 cm. long, capitate, rather few- or
submany~flowered; peduncles very slender or filiform, 10--12
mm. long, densely appressed-pubescent with antrorse canes
cent or yellowish hairs; heads small, about 5 mm. long and
wide; bractlets lanceolate, about 4 m. long and 1.5 mm.
wide, densely appressed-strigose—pubescent, sharply acute;
corolla about 5 mm. long, its limb about 2 mm. wide.

The type of this species wae collected by August Weber-
bauer (no. 6510) in the valley of the Mantaro, northeast of
Pampas, prov. Tayacaja, dept. Huancavelica, Peru, at an alt-
itude of 1800—-1900 m., in March, 19135, and is deposited in
the United States National Herbarium at Washington. The
species is related to L. ferruginea H.BeK.

PHYLA NODIFLORA var. LONGIFOLIA Moldenke, var. nov.

Haec varietas a forma typical speciei recedit foliis val-
de elongatis oblanceolato-cuneatis usque ad 5.5 cm. longis
et 10 mm. latis, versus apicem argute patento-dentatis.

This variety differs from the typical form of the species
in its much more uniformly elongate leaves, the blades being
oblanceolate-cuneate, to 5.5 cm. long, 4--10 mm. wide, and
sharply spreading-dentate toward the apex.

The type of this variety was collected by T. G. Yuncker,
Je Me Koepper, and K. A. Warner (no. 8327) in sandy soil on
the beach at Salado, in the vicinity of La Ceiba, Atlantida,
Honduras, on July 10, 1938, and is deposited in the Britton
Herbarium at the New York Botanical Garden.

PHYLA NODIFLORA var. ROSEA (D. Don) Moldenke, comb. nov.
Zappania nodiflora var. rosea De Don in Sweet, Brit. Fl.
Gard. 6: pl. 2256 1834.

STACHYTARPHETA SCHAUERII Moldenke, nome nove

Stachyterpheta villosa (Pohl) Schau. in A. DC., Prodr.
11: 570. 1847 [not S. villosa Oham., Linnsea 7: 247. 1832]
-- Melasanthus villosus Pohl, Pl. Bras. Ic. 1: 76, pl. 60.
1827.

VERBENA BAJACALIFORNICA Moldenke, spe nove

Herba annua; remis rectis simplicibus vel pauci-brachiat-
is obtuse tetragonis sparse hirsutulis glabrescentibus; pet-
iolis gracillimis dense vel sparse hirsutulis submarginatis;
laminis chartaceis ovatis pinnatifido-incisis vel obscure 3-
partitis subrevolutis utrinque sparse hirsutulo-pilosis.

Annual herb; stems erect, simple or sparsely branched,
8--15 om. long, obtusely tetragonal, sparsely hirsutulous

1941 | Moldenke, Novelties 23

with mostly scattered, stiff, whitish, non-glandular hairs
about 1 mm. long, glabrescent in age, sometimes decumbent at
the very base and throwing out roots from the lower nodes;
leaves decussate-opposite, petiolate; petioles distinct,
very slender, 1--10 mm. long, densely or sparsely hirsutul-
ous with stiff, white, nomglandular hairs like the stems,
slightly margined; blades chartaceous, uniformly green on
both surfaces, ovate in outline, 0.8--2.7 om. long, 0.41.8
em. wide, sparsely hirsutulous—pilose with rather short and
subappressed whitish hairs on both surfaces, more densely so
along the midrib and larger veins beneath, abundantly pinna~
tifid-incised, sometimes obscurely 3—parted with the divis-
ions again abundantly pinnatifid-incised, the lobes rounded,
subrevolute along the margins; inflorescence erect, long-
pedunculate, 5--15 or more cm. long; peduncles slender, ob-
tusely tetragonal, 2--6.5 cm. long, sparsely hirsutulous-
pilose with rather scattered nom-glanduler whitish hairs;
rachis densely many-flowered, more densely hirsutulous, not
glandular, the flowers close together and densely imbricate
before, during, and even after anthesis or the 2 or 3 lower-
most to 5 mm. apart in fruit; bractlets very small, lanceol-
ate, 2--5 mm. long, about half the length of the calyx, att-
enuate at apex, glabrate except for the long-cilisate margin;
oalyx tubular, 4--5 mm. long, irregularly short-pubescent
with whitish spreading hairs, obscurely (if at all) glandul-
ar; corolla 7--8 mm. long, slightly projecting from the cal-
yx, its tube slightly puberulent at apex outside, its limb
about 4 mm. wide.

The type of this species was collected by Forrest Shreve
(no. p69) eighteen miles north of El Refugio, Baja Califor-
nia, Mexico,on March 16, 1935, and is deposited in the herb-
arium of the University of Michigan. It is closely related
to V. Shrevei Moldenke, but differs in its ovate abundantly
incised=pinnatifid leaves, sparser non-glandular pubescence
on stems and peduncles, densely flowered spikes with closely
imnrivate flowers even after anthesis, and very short non-
glandulose bractlets.

VERBENA CLOVERI var. LILACINA Moldenke, var. nov.

Haec varietas a forma typica speciei recedit rhachide
calycibusque bracteolisque brevissime pubescentibus, pilis
glanduliferis, et corollis lilacinis.

This variety differs from the typical form of the species
in its much shorter and densely glandular pubescence on the
rachis, calyx, and bractlets and in its lavender (instead of
purple) corollas.

The type of this handsome variety was collected by Cyrus
Longworth Lundell and Amelia A. Lundell (no. 10,142) off U.
S. Highway 81 near Millett, La Salle County, Texas, on April

24 PHYTOLOGIA : Vol. 2, now. l

9, 1941, and is deposited in the Britton Herbarium at the
New York Botanical Garden.

VERBENA LUNDELLIORUM Moldenks, sp. nov.

Herba; ramis gracilibus obtuse tetragonis albido-hirsutu-
lis; petiolis 5--10 mm. longis valde hirsutulis submarginat-
is; laminis ovatis acutis, ad basin subtruncatis et in peti-
olum subprolongatis, ad marginem crasse et irregulariter in-
ciso-dentatis, utrinque sparse adpresso-pilosis, pilis albi-
dis; inflorescentiis spicatis abbreviatis dense multifloris;
pedunculis acute tetragonis valde hirsutulis; bracteolis
lineari-lanceolatis ca. 6 mm. longis dense puberulis, ad
marginem longe ciliatis.

Herb, about 18 inches tall; stems slender, obtusely tet~
ragonal, rather abundantly hirsutulous with stiff white
hairs about 1 mm. long; branches numerous, very slender, e-
rect or ascending, obtusely tetragonal, more densely hirsut-
ulous; leaves decussate-opposite, numerous; petioles very
slender, 5--10 mm. long, abundantly hirsutulous like the
branches, slightly margined; blades thin-chartaceous or mem
branous, ovate, somewhat lighter=—green beneath, 1.1--3.5 om.
long, 7-22 mm. wide, acute at apex, subtruncate at base and
slightly prolonged into the petiole at the center, coarsely
and irregularly incised-dentate along the margins with blunt
er subacute teeth, sparsely scattered-pilose on both surf-
aces with appressed whitish hairs; inflorescence spicsate,
abbreviated, 2--6 cm. long, densely many-flowered, the flow-
ers closely imbricate before and during anthesis, somewhat
more separated in fruit; peduncles (8-30 mm. long) and ra-
chis very slender or filiform, more acutely tetragonal, ab-
undantly hirsutulous like the branches, often with shorter
glandular hairs interspersed; bractlets linear-lanceolate,
about 6 mm. long, slightly shorter or longer than the calyx,
densely puberulent, long-ciliate along the margins with
stiff white hairs; calyx tubular, swollen, 5--6 mm. long,
somewhat puberulent and also sparsely hirsutulous with long=-
er white hairs, not glandular; corolla small, inconspicuous,
barely protruding from the calyx, about 7 mm. long, purple,
its limb about 2 mm. wide.

The type of this curious woodland species was collected
by C. L. Lundell and A. A. Iundell (no. 8698) -- in whose
joint honor it is named -- in a clearing at the Palm Grove,
south of Brownsville, Cameron County, Texas, on May 4, 1940,
and is deposited in the Britton Herbarium at the New York
Botanical Garden.

VERBENA PLICATA var. DEGENERI Moldenke, var. nov.
Haec varietas a forma typica speciei recedit bracteolis
firmis rigidis late ovatis usque ad 9 mm. longis et 6 mm.

1941. Moldenke, Novelties 25

latis stramineis siccis abruptissime longeque acuminatis.

This variety differs from the typical form of the species
in its bractlets being very firm and rigid, broadly ovate,
dry, stramineous, to 9 mm. long and 6 mm. wide, and very ab-
ruptly long-acuminate.

.The type of this desert variety was collected by my good
friend and co-worker, Otto Degener (no. 5184), near Fort
Stockton, Pecos County; Texas, on August 2, 1953, and is de-
posited in the Britton Herbarium at the New York Botanical
Garden. It affords me exceptional pleasure to dedicate this
variety to so careful and indefatigable a botanical collect-
or, whose monumental "Flora Hawaliensis" is one of the most
important and valuable floras now being written. _

VERBENA RUNYONI Moldenks, sp. nov.

Herba alta anrmma; caulis reotis crassiusculis argute tet-
ragonis sparse hirsutulis glabrescentibus; foliis sessilibus
amplexicaulibus plusminus tripartitie, segmentis pinnatifido
=incisis, utrinque valde albido-hirsutulus, pilis bulbosis
deciduis, laminis senectute scabris; inflorescentiis spicat-
is compositis, ramis gracilibus rectis densiuscule miltiflo-
ris, floribus densissime imbricatis; pedunculis rhachideque
gracilibus argute tetragonis patento-pilosis vel breviter
pubescentibus, pilis glandulosis brevissimis.

Tall annual herb; stems erect, green, rather stout, sharp
ly tetragonal, sparsely hirsutulous with short whitish di-
vergent hairs especially on the angles and at the nodes,
glabrescent in age, more or less scabrellous on the angles;
internodes elongated; leaves decussate-opposite, sessile,
clasping, 2~--6 om. long, 0.8--3 cm. wide, more or less 3-
parted, each division pinnatifid-incised with broad acute
teeth, abundantly hirsutulous on both’ surfaces with rather
short whitish hairs which are bulbous~based on the upper
surface and wear off there, leaving the upper surface scab-
rous on older leaves; inflorescence spicate, compound, the
branches slender, erect, 14--25 cm. long, rather closely
many-flowered, often bearing 1——3 pairs of much reduced
leaves near the base, the flowers with a faint odor, very
densely imbricate before and during anthesis, rather uni-
formly separated in fruit; peduncles (2--6 cm. long) and re-
chis slender, sharply tetragonal, rather densely or sparse-
ly spreading-pilose or -pubescent, glandular, the pubescence
very short; bractlets linear-lanceolate, about 3 mm. long,
equaling the calyx, sharply attenuate, rather sparsely pub-
erulent and glandular, the margins sparsely and irregularly
ciliolate toward the base; calyx tubular, about 3 mm. long,
glandular-pilose with short spreading hairs; corolla blue,
about 6 mm. long, its tube puberulent at the apex outside,
its limb about 4 mm. wide.

26 | PHYTO,GeLE sz Vol. 2, now l

The type of this hitherto neglected species was collected
by my good friend, Robert Runyon (no. 2485) in clay soil at
10 m. altitude in open moist ground and ditches, El Jardin
tract, Cameron County, Texas, on April 2, 1941, and is dep-
osited in the Britton Herbarium at the New York Botanical
Garden. It is with considerable satisfaction that I dedicate
this fine species to Mr. Runyon, who has done such note-
worthy work in botanizing so thuroughly the region of Camer-
on and Hidalgo Counties, Texas, and in collecting such ample
and excellent material to substantiate his records through
the years. Museum and herbarium workers are deeply indebted
to field workers like this, to whom so mich of the credit in
the discovery of novelties is due. The species is related to
and has hitherto been confused with V. xutha Lehm., which
differs notably in its dense long-strigose or hirsute non=-
glandular pubescence throughout, especially on the bractlets
and calyx, and which inhabits dry instead of uniformly moist
ground.

VERBENA SHREVEL Moldenks, sp. nov.

Herba annua; ramis decumbentibus gracilibus obtuse tetra-
gonis dense patento-pubescentibus; foliis petiolatis vel
subsessilibus; petiolis marginatis dense hirsutulis vel pat-
ento=pubescentibus; laminis chartaceis ellipticis utrinque
dense strigosis plerumque plusminus tripartitis, partibus
pauce inciso-lobatis, lobis rotundatis.

Anmal herb; stems decumbent at base, slender, obtusely
tetragonal, more or less densely spreading-pubescent with
whitish often glandular hairs, often many=branched with er-
ect or ascending branches, which are usually somewhat more
densely spreading=-pubescent; leaves decussate-opposite, pet-
iolate (or the uppermost subsessile); petioles very slender,
1--10 mm. long, more or less winged, densely hirsutulous or
spreading-pubescent; blades chartaceous, rather uniformly
green on both surfaces, elliptic in outline, 1=-2 cm. long,
9--17 mm. wide, rather densely strigose on both surfaces,
usually more or less 3—parted, the divisions sparingly in-
cised-lobed, the lobes rounded at apex; inflorescence spic-
ate, elongating to 10 cm. or more, densely many-flowered,
the rachis elongating even during anthesis and thus separat-
ing the individual flowers by 4——13 mm. toward the base of
the spike; peduncles (1--4 cm. long) and rachis slender, ob-
tusely tetragonal, densely spreading-pubescent or hirsutul-
ous with whitish often glandular hairs; bractlets lanceola-
te, about 4 mm. long, shorter than the calyx, attenuate at
apex, densely glandular=pubescent on the back, densely long-
ciliate with longer stiff white non-glandular hairs on the
margins; caiyx tubular, 5--6 mm. long, rather densely gland-
ular-pubescent and also more or less scattered white-hirsut-

1941 Moldenke, Novelties 27

ulous; corolla small, 7--8 mm. long, slightly projecting
from the calyx, its tube minutely puberulent at the apex
outside, its limb about 4 mm. wide.

The type of this speoies was collected by my esteemed
friend, Dr. Forrest Shreve (no. 7119) -- in whose honor it
is named -- at an elevation of 1900 feet, 19 miles northeast
of Oomondén, Baja California, Mexico, on March 16, 1935, and
is deposited in his herbarium at Tucson, Arizona. It has
hitherto been confused with V. pumila Rydb.

VERBENA GENTRYI Moldenke, sp. nov.

Herba perennis ramulosa; ramis graciusculis tetragonis
sparsissime pilosis vel glabris; petiolis indistinctis et
alatis vel obsoletis; laminis chartaceis ellipticis acutis,
ad basin cuneatis, regulariter arguteque serratis utrinque
adpresso-sirigillosis non scabris.

"Low spreading bush, branched from base"; branches rather
slender, tetragonal, often purplish, very sparsely scattered
pilose with rather long weak hairs or glabrous; nodes annul-
ate; principal internodes 1--3.5 om. long; leaves decussate-
opposite; petioles indistinct, to 5 mm. long, and winged, or
absent, ampliate and clasping the stem at base, sparsely
scattered-pilose or glabrescent; blades chartaceous, lighter
beneath, elliptic, 3--7 cm. long, 1--1.7 cm. wide, acute at
apex, cuneate at base and prolonged into the winged petiole,
regularly sharp-serrate from the apex to below the middle,
rather abundantly appressed—strigillose on both surfaces,
more densely so beneath, not scabrous above, not glandular;
venation slightly subimpressed above, prominulous beneath;
inflorescence spicate, compound, the spikea very slender, to
18 or more cm. long, many-flowered, the flowers closely im-
brieate before and during anthesis, rather uniformly separea-
ted in fruit; peduncles (1—3 cm. long) and rachis slender,
glebrate; bractlets lanceolate, very small, about 1--2 m.
long, subglabrate or very minutely ciliolate at the base,
sharply acuminate; calyx narrow-tabular, about 1.5 mm. long
(to 2 mm. long in fruit), glabrous or subglabrate; corolle
very tiny.

The type of this species was collected by Howard Scott
Gentry (no. 5923) -- in whose honor it is named — in a
moist canyon bottom, short-tree forest, altitude 1500 feet,
Quebrado de Platano, Sierra Monterey, Sinaloa, Mexice, on
March 13, 1940, and is deposited in the Britton Herbarium at
the New York Botanical Garden. It was distributed by the
collector as V. urticifolia L., to which it is related.

VERBENA PINETORUM Moldenke, sp. nov.
Herba; caulis gracilibus argute tetragonis crasse hirsut-
is; petiolis late alatis; laminis chartaceis profunde pinn-

28 PHY 20 bOG 22 Vol. 2, no. l

atifido-incisis vel tripeartitis, supremis plerumque oblongis
vel linearibus et integris, utrinque dense hirsutis, suprem
is adpresso-strigosis; inflorescentiis elongatis spicatis.

Herb; stems slender, sharply tetragonal, bristly-hirsute
with whitish heirs about 1 mm. long, much more densely so at
base of plant; leaves decussate~opposite, 2--4 om. long; pe-
tioles broadly winged, not very distinct from the blades;
blades chartaceous, uniformly green on both surfaces, deeply
pinnatifid-incised, the lower ones often 35-parted and each
division again pinnatifid-incised, the uppermost much reduc-
ed and simply 3-parted with entire divisions or even oblong
or linear and entire, densely hirsute on both surfaces, the
larger leaves scabrous with bulbous-based hairs above and
very densely white-hirsute beneath, the upper leaves with
mich more appressed-strigose hairs, especially above; inflo-
rescence spicate, elongates spikes slender, to 21 or more
cme long, loosely many~-flowered (dense in bud and during ar
thesis, the rachis later elongating considerably), not glan-
duler; peduncles slender, sharply tetragonal, 2--}5 cm. long,
hirsute like the stems; rachis also tetragonal and densely
hirsute; bractlets lanceolate, about 4 mm. long, attenuate
at apex, rather densely strigose-pilose, about equaling the
calyx in anthesis and fruit; calyx tubular, 3.5--4 mm. long,
densely strigilloses corolla pale-blue, showy, 10--11 m.
long, its limb large and spreading.

The type of this species was collected by Howard Scott
Gentry (no. 1522) in pine flats, transition habitat, Sierra
Charuco, Rio Fuerte, Chihuahua, Mexico, on July 22, 1935,
and is deposited in the herbarium of Dr. Forrest Shreve at
Tucson, Arizona. It.was originally distributed as V. neomex-
icana (A. Gray) Small

VERBENA PINNATILOBA (Kuntze) Moldenke, comb. nov.
Verbena megapotamica var. tweediana f. pinnatiloba Kun-
tze, Reve Gen. Pl. 3%: 256. 1898.

xVERBENA TEASII Moldenke, hybr. nove

Herba oultorum hybrida; ramis decumbentibus vel adscend-
entibus multoramosis gracilibus obtuse tetragonis sparse vel
dense hirsutulis; foliis pervariabilis dense strigosis vel
supra sparse strigillosis et subtus patento-pubescentibus,
pluemims profunde inciso-pinnatifidis plerumque tripartit-
is; inflorescentiis spicatis, juventute subcapitatis, dein
elongatis, densissime multifloris; floribus arcte imbpicatis

Garden hybrid between Ve. tenuisecta Briq. and V. hybrida
Voss, with intermediate characters; stems decumbent or as-
cending, abundantly branched with ascending branches, slend-
er, obtusely tetragonal, sparsely or densely hirsutulous
with rather stiff whitish hairs or merely spreading-pilose,

1941 Moldenke, Novelties 29

the smaller branches often more acutely tetragonal; leaves
decussate-opposite, numerous, very variable in shape and
size, varying from densely strigose with long white appress-
ed hairs on both surfaces to sparsely strigillose above and
spreading-pubescent along the midrib and larger veins be-
neath, more or less deeply incised in pinnatifid fashion,
often more or less 3-parted, the lowest divisions usually a-
gain pinnatifid-incised, the lobes all sharply acute at ap-
ex, the body of the blade and lowest lobes often relatively
very broad and with recurved secondary lobes; inflorescence
spicate, at first flattened~subcapitate, later elongating to
15 cm. or more, very densely many-flowered, the flowers
closely overlapping before, during, and after anthesis; ped-
uncles (1.5--7 em. long) and rachis slender, acutely tetrag-
onal, densely hirsutulous or spreading-pilose, not glandu-
lar; bractlets relatively very short, lanceolate, about 4 m
long, about 1/3 as long as the calyx, attermate to the apex,
rather densely strigillose with white appressed hairs, den-
sely white-ciliate toward the base; calyx elongate-tubular,
8--13 mm. long, densely short-pubescent with spreading hair
or densely white-strigose with closely appressed hairs; cor-
olla 15--20 mm. long, showy, blue, purple, red, pink, or
white, its tube about 1 1/3 times as long as the calyx, gla-
brous throughout or slightly pubervlent at the apex outside,
its limb 5—9 mm. in diemeter.

The type of this hybrid was collected by G. A. Stevens in
a nursery at Harrisburg, Dauphin County, Pennsylvania, on
June 20, 1953, and is deposited in the herbarium of the Bai-
ley Hortorium at Ithaca. It is named in honor of Edward Teas
who first developed this hybrid in his nurseries at Houston,
Texas, by crossing Ve hybrida and V. tenuisecta. It is the
source of the races of cultivated verbena called Ceres
(derk red), Rowena (pink), Albion (white), Ruth (pink), Bel-
laire, Madge Roberts, and Teas Hybrid.

xVITEX HYBRIDA Moldenke, hybre nov.

Arbor vel frutex hybridus naturalie; foliolis anguste
lanceolatis 5 longe attenuatis utrinque dense canescento-
puberulis; inflorescentiis distincte ramulosis; ramulis gra-
cillimis ubique dense canescento-puberulis.

A natural hybrid between V. Agnus-castus L. and V. Negun-
do L. with intermediate characters. The leaflets are narrow-
lanceolate, 5 in number, the three central ones 5.5--10 om.
long and 7--16 mm. wide, long-attenuate at apex, densely
canescent-puberulent on both surfaces, on petiolulys 3--5
mm. long, the lowest two very much smaller. The inflorescen-
ces aro distinctly branched, the branches very slender, 5--
15 mm. long, with numerous nodes and flowers (in the fashion
of V. ‘agundo), densely canescent-puberulent throughout.

50 PHYTOLOGIA Vol. 2, noe l

The type of this variety was collected at Bhola in Sindh,
India, in July, 1891, and is deposited in the herbarium of
the University of Michigan at Amn Arbor. No collector is
designated on the label.

VITEX REGNELLI ANA Moldenke, Geogr. Distrib. 27, nom. nud.
(1939), sp. nov.

Frutex vel arbor; ramulis gracilibus medullosis obtuse
tetragonis puberulis vel breviter pubescentibus glabrescent-
ibue; sarmentis densissime ferrugineo-velutinis vel villoso-
tomentosis; foliis oppositis 3-foliolatis; petiolis gracili-
bus densissime velutino-villosis vel tomentosis ferrugineis;
foliolis sessilibus vel subsessilibus oblongis vel anguste
ellipticis vel oblanceolatis acutis vel abruptissime brevi-
terque acuminatis integris, ad basin acutis vel obtusis, ut-
rinque velutinoso-villosis vel tomentosis; inflorescentiis
axillaribus cymosis valde bracteatis ubique dense ferrugin-
eo-velutinis vel villoso-tomentosis.

Shrub or tree; branchlets slender, medullose, obtusely
tetragonal, grayish, compressed and rather ampliate at the
nodes, puberulent or short-pubescent when young, becoming
glabrate in age; twigs slender, tetragonal, compressed, very
densely velutinous with ferruginous villous-tomentose pubes-
cence, ampliate-compressed at the nodes; nodes annulate;
principal internodes 1--6 em. long; leaf-scars very large
and corky, greatly elevated; buds densely ferruginous-vill-
ous or -velutinous; leaves decussate~opposite, 3-foliolate;
petioles slender, 4--10.5 cm. long, slightly ampliate at the
base, flattened above, very densely velutinous-villous or
tomentose with ferruginous hairs; leaflets subequal, sessile
or subsessile; leaflet-blades thin-chartaceous, uniformly
dark- or bright-green on both surfaces under the ferruginous
tomentum, the central one oblong, narrow-elliptic, or oblan=
ceolate, 5.5--10 cm. long, 1.5--3.2 cm. wide, acute or very
abruptly short-acuminate at apex, entire, acute or obtuse at
base, very densely velutinous-villous on both surfaces or
somewhat more tomentose beneath, the pubescence golden or
ferrugineous, the lateral leaflets similar in all respects
only often somewhat inequilateral and usually more obtuse at
the base; midrib slender, flat or subprominulent above, pro-
minent beneath; secondaries slender, about 10 per side, most
ly hidden by the long pubescence on both surfaces or promin-
ulous beneath; vein and veinlet reticulation not discernible
above, mostly obscure beneath or sometimes the largest parts
slightly subprominulous beneath; inflorescence axillary, cy-
mose, 3~--8 cm. long, 2--4.5 cm. wide, 1—3 times dichotorm
ous, dense, the branches much abbreviated, densely ferrugin=
ous-velutinous or villous-tomentose throughout, conspicuous-
ly bracteate; peduncles slender, 1.5--5.2 om. long, flatten-

ne , aae7

1941 Moldenke, Novelties 31

ed, densely ferruginous-velutinous or villous-tomentose like
the twigs and petioles; pedicels very slender, 1-2 mm.
long, or obsolete on lateral flowers; bracts numerous, simp-
le, oblong or lanceolate, 1--1.8 com. long, densely velutin-
-ous like the leaflets, sessile, acute; bractlets linear, 3-—
6 mm. long, densely ferruginous=pubescents; prophylla linear,
about 1 mm. long, hidden by the tomentum; corolla violet or
white.

The type of this very handsome species was collected by
Don Bento Pickel (no. 3211) in a thicket at Tapera, Pernam-
buco, Brazil, on Jamary 26, 19353, and is deposited in the
Langlois Herbarium of the Catholic University of America at
Washington. The species is known also from S&o Paulo and is
named in honor of Anders Fredrik Regnell, famous explorer
and botanist, to whom we owe so much of our knowledge of the
Brazilian flora.

VITEX SPONGIOCARPA var. LONGIDENTATA Moldenke, var. nov.

Haec varietas a forma typica speciei recedit lobis caly-
cis 1.5--2 mm. longis et bracteolis prophyllisque persister-
tibus.

This variety differs from the typical form of the species
in ite calyx-teeth being 1.5--2 mm. long (instead of 0.5--1
mm.) and its bractlets and prophylla being persistent.

The type of this variety was collected by Adolfo Ducke
(Herb. Jard. Bot. Rio de Janeiro 23,763] in "catinga" woods
at Igarape Jurupary, on an affluent of the lower Rio Uaupés,
Amazonas, Brazil, on November 2, 1932, and is deposited in
the Britton Herbarium at the New York Botanical Garden. The
collector describes the plant as a small tree with white
flowers.

VITEX TRIFOLIA var. VARIEGATA Moldenke, var. nov.

Haec varietas a forma typica speciei recedit laminis
foliolorum irregulariter albo-variegatis.

This variety differs from the typical form of the species
in its leaflet-blades being variegated, whitish along the
edges in irreguler mottles.

The type of the variety was collected by my good friend,
Walter M. Buswell, from a cultivated specimen at or near
Miami, Dade County, Florida, in 1940, and is deposited in
the herbarium of the Bailey Hortorium at Ithaca.

VITEX WITTROCKLANA Moldenke, Geogr. Distrib. 20 & 27, nom.
nud. (1939), sp. nove
Arbor; ramulis crassiusculis obtuse tetragonis vel subte-
retibus sparsissime minutissimeque puberulis glabrescentib-
us; sarmentis acutiuscule tetragonis sparse puberulis; foli-
is oppositis 5-foliolatis; petiolis gracilibus sparsiuscule

32 PHEYTOLOG ie Vol. 2, no. l

puberulis; foliolis subsessilibus vel brevipetiolulatis ten-
uiter chartaceis vel submembranaceis oblongis vel lanceolat-
is vel oblanceolatis longe acuminatis vel caudatis integris,
ad basin acutis vel subacuminatis, supra glabris nitidis,
subtus glabratis vel obscure puberulis; inflorescentiis ax-
jllaribus capitatis dense multifloris sparse strigilloso-
puberulis. .

Tree, to 7 m. talls branchlets rather stout, brownish,
obtusely tetragonal or subterete, not very pithy, very spar-
sely and minutely puberulent, becoming glabrous and rather
shiny; twige very slender, rather acutely tetragonal or com-
pressed, short, rather sparsely puberulent, less so in age;
nodes not annulate; principal internodes 1--6 em. long; leaf
-scars mostly not very large or corky or prominont; leaves
decussate-opposite, S-foliolate; petioles slender, 2--5 cm.
long, convex or slightly keeled beneath, conspicuously flat-
tened above, rather sparsely puberulent, not noticeably amp-
liate at base nor disciform at apex; leaflets usually unequ-
al, the 2 lowermost mich smaller than the 3 central ones,
all subsessile or the central one short-petiolulate on a pe-
tiolule which is slightly puberulent and margined and to l
mn. long; leaflet-blades thin-chartaceous or submembranous,
dark-green and rather shiny above, lighter beneath, the cen-
tral one oblong, lanceolate, or oblanceolate, 53.5--8.5 cme
long, 2--3 cm. wide, long-acuminate or caudate at apex, en-
tire, acute or subacuminate at base, glabrous and shiny ab-
ove, glabrate beneath or obscurely puberulent on the midrib
and secondaries; midrib slender, flat or slightly impressed
above; secondaries slender, 7--15 per side, ascending, not
much arcuate except at the margins, where they are arcuately
joined, flat or subprominulous above, prominulous beneath;
vein and veinlet reticulation abundant, very fine, subprom
inulous on both surfaces; inflorescence axillary, capitate,
5--8.5 cm. long, 1--2 cm. wide, densely many-flowered, some-
times with a few very short branches arranged in subumbell-
oid form; peduncles slender, compressed, 4--7.3 cm. long,
sparsely strigillose-puberulent; pedicels very slender and
to 1 mm. long or usually obsolete; bracts absent; bractlets
linear, 1--3 mm. long; prophylla setaceous, minute; corolla
violet.

The type of this species was collected by Jofo Geraldo
Kuhlmann (no. 2915) in campo at Caracarahy on the Rio Bran-
co, Amazonas, Brazil, in February, 1913, and is deposited in
the United States National Herbarium at Washington. The spe-
cies is also known from adjacent Venezuela and is named in
honor of. Gustave Ludwig Wittrock, custodian of the herbarium
at the New York Botanical Garden, conscientious worker on
all botanical subjects, and expert on the plants used by the
North American Indians.

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d evolution

Additional notes ont :
shed and Harold N.Moldenke == —™S
The New York Botanical Garden ae A oor ae
ae ae RATS 1 RON eee VE ee gee a Dy Rage ee Bh te
Bronx Park, New York, N. Y : 3
| er volume, $5.00 | |
oiath

ENERGY AND EVOLUTION
James B. pained GARE Es
The object of this paper is to develop the theory that

species formation occurs during periods of increased activi-
ty, that plants which do the hardest (most difficult) work

have evolved to the highest positions; that in this regard

quality of products is mre important than quantity; and
that as morphological structures evolve from simple to com
plex, so plant chemical compounds evolve from simple to com
plex. .

Species Definition from a Chemical Standpoint. - It is
assumed that each species is in a state of mobile equilibri-
um between reversible reactions which fluctuate and are mt-
able under the action of modifying agents (Marcello, 1930).

Individuality has, therefore, a complex chemical basis. The

existence and permanency of a species is controlled and de-
pends upon the existence of constant external and internal
conditions and shows a fixed ability to synthesize charact-
eristic compounds which constitute its physiologico-chemical
characteristics (in part S. L. Ivanov, 1926). Thus, an in-
crease in habitat temperature will stimlate the formation
of more saturated fatty acids in glycerides and vice versa.
Also, at moderately elevated temperatures starch is formed
in evergreen leaves which is converted into oil when the
temperature is gradually lowered and vice versa (Tuttle,
1919). As another instance of change of conuitions affecting
plant physiologico-chemical characteristics we know that an
increase in water in the soil and in the plant promotes oil
formation and vice versa (Sinnott 1917, Ivanov, Lavrova and
Japochko 1931, Geddes 1934, Halden 1934). The concentration
of electrolytes in plants is a factor controlling the amount
of alkaloids and cyanogenetic compounds formed (McNair,

1941). As instances of the effect of internal conditions on

plant chemical products we have the influence of changes in
genetic constitution. Genetic strain affects HON production
in white clover (Williams, 1939) and sorghum (Nowosad and
MacVicar, 1940). Genetic strain affects alkaloid production
in tobacco (Rasmssen, 1915), opium (Annett and coworkers,
1920-1925) ,and aconite (Bonisteel, 1940, 1941), And genetic
strain also affects the amount and distribution of oil in
corn kernels (Pearl and Bartlett 1911, Lindstrom and Ger-
Species Developed During Greater Activity. - It is the
consensus of opinion that although species may originate in
a number of different ways they all originate during periods
35

34 PHY T10-b 6 G teak Vol. 2, now 2

of greater activity. This greater activity may take place
internally in or externally to the plant. According to geo-
logical evidence the splitting off of new species apparently
falls within the times of greater range of variation in ail
characters, therefore of greater plasticity of species
(Brinkmann, 1929). According to the biologists, especially
physiologists, structure varies with function (Tait, 1928)
and functional activity is emphasized as the foundation of
structural differentiation (Leathes 1926, Fox 1932), or in
the words of Pycraft (1950) changes of form are responses to
continuous and persistent needs. The geneticists, as pointed
out by Huxley (1941), have shown that new species may arise
suddenly at a single bound. Instances of such greater activ-
ity are shown in chromosome-doubling (6. g. Osnothera), the
inverting end-to-end of a considerable section of one chrom
osome (e. ge. Datura) or the detachment of a bit of one
chromosome which may become attached to a different kind of
chromosome (e. g. in Drosophila). A chemist, Henderson
(1922), has suggested that apparent instances of orthogenes-
is may sometimes depend upon a single important chemical
change in an organism, followed by slow and progressive mod-
ifications leading up to a definitive morphological result.
Such a process, he says, might be somewhat analogous to the
establishment of a condition of equilibrium.

Climate, Energy and Evolution. - As Parks (1926) points
out from geological evidence, there is an undoubted tendency
to increased complexity in the organic world. Consequently
the greatest complexity in both form and substance may be
expected to be found in such regions and in such plants as
undergo the most rapid changes of external and internal con=
ditions. There are, of course, optimum conditions above
which the foregoing statement would not be true.

From a detailed study of the varietal diversity of culti-
vated plants and their wild relatives Vavilov (1932) found
that the majority have had their origin in comparatively
small territories concentrated mainly in the mountains and
foothills of the subtropics and tropics. The mountain and
foothill regions in the subtropics he found especially fav-
orable for the development of species and varietal diversity.
Mountains provide geographic types of isolation in the nat-
ure of differences between habitats - woodland and open
country, pond and swamp, high ground and low ground, sunny
southern slopes and shady northern slopes, canyons and
ridges. These barriers isolate small populations and then
useless accidental characters automatically accumlate. Much
greater divergence is achieved on small areas (islands) as
compared to large continental areas. Sewell Wright offers
the explanation that if isolated populations are small
enough in numbers, mere chance will step in and largely

See SS ee es er eS

A tiie te St

Meg Ae

Lid

1941 ‘McNair, Ener gy & Evolution 35

override the effects of selection.

Greater differences between habitats are found in the
mountains of the tropics and subtropics than in those nearer
the poles. We have in tropical mountains various life zones
from tropical, lower sonoran, upper sonoran, transition, to
boreal, while in mountains nearer to the poles some of these
zones are absente

In tropical lowland climates where conditions are more
stable one would not expect to find the most highly evolved
plants or the most complex chemical compounds. But rather
the most highly evolved plants and the most complex chemical
compounds would be found more likely in the subtropics and
temperate zones where fluctuations of environment occur. For
a similar reason aquatics with their more equable environ-
ment would be more primitive than land plants. In tnis con-
nection Went (1941) has shown that in tomatoes either a high
uniform temperature or a low uniform temperature did not
promote nearly as mich growth or fruiting as when a fluctua-
ting temperature consisting of a high day temperature and
low night temperature was provided. ’

Alkaloids. - If the alkaloids be first separated accord-
ing to the habitat climates of the plant families producing
them, it becomes apparent that the alkaloids of the highest
molecular weight are produced by temperate plants and that
those with the lowest are obtained from tropical families
(Table I) (McNair, 1934).

The greatest number of plant families and also the great-
est number of plant families from which alkaloids have been
analyzed is found in the tropics. Some 299 alkaloids have
been analyzed. All else being equal a largest number of an-
alyses should lead to the most accurate results. Consequent=
ly tropical alkaloids are used. When this is done it is
found that the higher the tropical plant family is in .evolu-
tionary development, the greater will be its tendency to
RG ape of large average molecular weight (McNair,
1934). |
Inasmich as it generally requires more difficult work to
produce chemical compounds of large molecular weight than
those of small molecular weight, it can be argued that the
higher evolved plants which likewise manufacture alkaloids
of greater molecular weight perform more difficult work than
more primitive plants.

A specific example in which the molecular weignt of al-_
kaloids may serve to indicate the degree of evolution of
species is shown in the members of the genus Aconitum. Acon-
itum is noteworthy in giving a new chemical species of acon-
itine for each new botanical species analyzed, although all
the aconitines are apparently closely related. Perhaps India
is the center of distribution of this gems for here we find

36 P H-Y T’OrhO-G LA Vol. 2, now 2

A. chasmanthum Stapf with indiaconitine CyyHy70;oN (mol. wt.
629), A. demorrhizum Stapf with pseudoaconitine C3,H4q0;2N
(mol. wt. 667) and A. spicatum Stapf which contains bikhac-
onitine C3,Hs00,,N (mol. wt. 672). Japan may be at the outer
boundary of distribution with a more recently evolved spec-
ies for here is found A. japonicum Thynb. which furnishes
jesaconitine CypH5;0;.N (mol. wt. 737) of a higher molecular
weight than the Indian alkaloids (Carr 1912, Schafer and La
Cour 1954). In a comparison of the chromosome numbers with
toxicity Bonisteel has found (1940, 1941) that the diploid
aconites are for the most part non-toxic, while the triploid
and tetraploid aconites contain some of the most powerful
poisons known. There is, therefore, an increase in toxicity
with an increase in chromosome number.

Glycerides. - Analyses of 318 fats (glycerides) are av-
ailable for study. In Table I the fatty oils from temperate
plant families have been separated from those produced by
tropical plant families. It is apparent from this table that
temperate fatty oils have higher average iodine values (and
lower melting points) than the tropical (McNair, 1934).

As in the case of alkaloids, the greatest number of plant
families from which glycerides have been analyzed is found
in the tropics. By use of the more abundant tropical data it
has been found (McNair, 1934) that the higher the plant fam-
ily is in evolutionary development the greater will be its
tendency to produce glycerides of large average iodine num-
bers (i. e. of greater unsaturation).

In the plant economy, saturated fatty acids are first
produced wnich become less saturated later. In this way add-
itional and more difficult work is necessary to form the
less saturated fatty acids and consequently it is evident
that the higher evolved plants which produce them perform
harder, more difficult work. :

The molecular weignts of tropical glycerides (in agree-
ment with the molecular weights of tropical alkaloids) are
lower then those of temperate regions. Hilditch (1928) found
that the tropical families Palmae and Myristicaceae had one
specific fatty acid for each family, respectively lauric
(mol. wt. 200, mep. 48° C.) and myristic (mol. wt. 228, m.p.
58° C.) and that the temperate families Cruciferae and Um-
belliferae had likewise one specific acid for each family, .
respectively erucic (mol. wt. 338, m.p. 33 .5° C.) and petro-
selinic (mol. wt. 282, m.ep. 14° C.). From this data it is
evident that the average molecular weight of the tropical
families, 214, is lower than that of the temperate, 310. As
it requires more energy to compound fatty acids of higher
molecular weight it is evident that these temperate families
which likewise occupy a higher evolutionary rank have more
difficult work to do than these tropical lower evolved fam-

{

ee ee ae

1941 McNair, Energy & Evolution ~ 37

ilies.

In the latest compilation of analyses of seed fats (Hil-
ditch, 1940) data from sixteen natural ordere (Engler and
Prantl classification) are given. When the component acids
of the families of these orders are considered it is found
that seven orders have an increase in the number of acids,
eight have an equal number of acids and one has a decrease
in the number of acids with an advance in evolutionary posi-
tion of their constituent families.

When the number of carbon atoms of these acids is consid-
ered it is found that eight orders have an increase in the
number of C-atoms, six have equal numbers of C-atoms and two
have a decrease in the number of C-atoms with an advance in
evolutionary position. If, however, the terminal families of
those analyzed of the Malvales, Myrtiflorae, Contortae and
Tubifloras (i.e. respectively Sterculiaceae, Myrtaceae, As-
clepiadaceas and Acanthaceae) be removed from consideration,
then three of these four orders show an increase in the nun
ber of acids and all four show an increase in the number of
C-atoms in these acids with an increase in evolution. An in-
crease in the number of C=-atoms indicates in these instances
an increase in molecular weight of the acids which contain
them. It is hardly necessary to add that both an increase in
the number of fatty acids as well as an increase in their
molecular weights require an increased expenditure of energy.

Volatile Oils. - Nilov (1936) shows in a study of the
essential oils in various stages of growth of Coriandrum
sativum, Trachyspermim copticum and other plants that,
parallel with the evolution of the plant, there occurs an
increase in the complexity of the molecules.

In the volatile oils the genus Eucalyptus provides an ex-
cellent demonstration of the progressive increase in the
number and variety of chemical products with the morphologi-
cal advance in evolutionary position in the genus. These
comprise in order of occurrence pinene, cineole, phelland-
rene, aromadendral (cuminal, cryptal, etc.) and piperitone
(Baker and Smith data 1920). The amount of oil in the leaf
also increases with the increase in evolutionary position,
@. g. the most primitive average 1/2 of 1 per cent (e. g.

E. corymbosa Sm.), while the most advanced (e. g. EB. dives
Schad.) have 2 per cent (4 times as mich).

Volatile Oils, Specific Gravities. - Analytical data from
938 volatile oils is available for study. When the average
specific gravities of the volatile oils produced by tropical
and temperate plant families are inspected, it is evident
that the volatile oils of tropical plant families have lower
specific gravities than those produced by temperate plants
(Table I) (McNair, 1932).

Further analysis of the more abundant tropical data shows

38 PHYTOLOGIA Vol. 2, no. 2

that the higher the tropical family is in evolutionary de-
velopment the greater will be its tendency to produce vola-
tile oils of high specific gravity (McNair, 1934).

It can likewise be inferred (McNair, 1932) in accordance
with these differences in specific gravity, that terpenes
and compounds of the fatty (aliphatic) series predominate in
the volatile oils produced lowest in the evolutionary posi-
tion, while volatile oils formed by the families highest in
evolution contain more aromatic, sulphur and nitrogen com
pounds. As more energy is generally required to produce aro-=
matic than aliphatic compounds we can conclude that families
highest in evolution carry on the most difficult work.

Volatile Oils, Refractive Index. ~ The refractive index
is another property which may be used to measure variations
in composition of volatile oils. From Table I it is evident
that tropical volatile oils have higher values than ventas
ate (McNair, 1932).

In addition it has been definitely shown (McNair, 1934)
that the higher the tropical plant family is in evolutionary

development, the smaller will be the average refractive in-.

dex of its volatile oil.

It can likewise be inferred (McNair, 1932) that a small
number or lesser amounts of saturated substances are formed
in the volatile oils produced lowest in the evolutionary
position. A high refractive index may also indicate a large
quantity of compounds of high molecular weight; therefore it
might be that the volatile oils produced lowest in the evo-
lutionary scale have less of these compounds.

However, it has been observed in volatile oils (McNair,
1932) that a low index of refraction carries with it a con-
comitant increase in specific gravity. Consequently, a trend
downward in the case of the refractive index (Fig. 5, McNair
1934) and upward in the case of specific gravity (Fig. 4,
McNair 1934), indicate that the values verify each other in
the case of evolutionary progression as well as in climatic
difference.

It can therefore be concluded that the volatile oils of
the tropical families highest in evolutionary development
have constituents with a large number of double bonds (low
saturation), more aromatic compounds, or more sulphur and
nitrogen compounds with small amounts of substances of low
molecular weight cr small quantities of terpenes or bodies
of the fatty series.

Because it requires more energy to form substances of
high than of low molecular weight, aromatic than fatty (ali-
phatic) compounds, it can be concluded that plant families
which manufacture these substances can be classed in the
same manner. That is, that temperate volatile oil producing
families are more energetic than tropical volatile oil fami-

1941 McNair, Energy & Evolution 39

lies, and that in tropical volatile oil families those high-
est in evolutionary development are more energetic and per-
form more difficult work than those lower in evolutionary
position.

Volatile Oils, Tropical Acids and Alcohols. - It has been
shown from a consideration of both the specific gravity and
refractive index of volatile oils that the higher the devel-
opment of a tropical plant family the greater is the com-
plexity of its chemical constituents. The study can likewise
be continued to the various components of volatile oils, e.
ge their acids and alcohols. When this was done, it was
found that the heats of combustion of the alcohols and acids
of tropical volatile oils increased in harmony with the in-
crease in evolutionary differentiation of the plant families
producing them (McNair, 1934).

The greater the heat combustion the greater the amount of
energy required in the making of the burned compound. Conse-
quently the higher the tropical plant family in evolutionary
position the harder the work it has had to perform.

Plant Form Versus Energy and Evolution. - Aristotle long
ago (384-322 B.C.) and his pupil Theophrastus (372-287 B.C.)
classified plants as trees, shrubs and herbs, and this
simple classification (in the words of A. M. Johnson) is the
one we.all first become aware of in our youth. It is plain
that this classification is based on "life-form" and that
the structure of the flower is ignored.

Bames (1911) brings forward evidence that the earliest
dicotyledons possessed a solid tubular woody cylinder of
considerable thickness which has gradually been reduced and
finally broken up into a circle of separate strands, which
is characteristic of the "typical" herbaceous condition.
Such an hypothesis of reduction from primitive arborescent
_forms has also been worked out under the direction of Prof-
essor Jeffrey by several other members of his laboratory
(Adkinson 1913, Bailey 1911, and Jeffrey 1912). In more re=
cent papers, Sinnott and Bailey (1914, 1922) produced evi-
dence in support of this view from paleobotany, phylogeny,
anatomy. and geographical distribution. It is no wonder that
Bessey (1915) included in his "general principles adopted
for the classification of plants" the postulate that "in
certain groups, trees and shrubs are probably more primi-
tive than herbs."

This hypothesis may be considered from the standpoint of
the chemical products derived from plants. In Table II the
glycerides, alkaloids and volatile oils from tropical plant
families are considered in this respect.

From the final average obtained of the molecular weights
of the alkaloids, there is a clear indication that trees
produce alkaloids of lower molecular weights than shrubs,

40 PHY 1.020 ¢:5a4 Vol. 25 NOe 2

and that shrubs have lower alkaloid averages than herbs.
Corresponding results are obtained from the iodine numbers
of glycerides. The average refractive indices and specific
gravities of volatile oils in respect to the dominant form
of plant growth in the families is also developed in Table
Il. Here again the findings clearly indicate that trees may
be the ancestors of herbs. This is shown in the specific
gravities. It has been observed that volatile oils with a
high specific gravity have a correspondingly low index of
refraction (McNair, 1932). If then the specific gravities of
volatile oils decrease from herbs to trees, the refractive
indices should increase from herbs to trees. This is the
case as shown by the averages (Table II), There is chemical
support, therefore, for the contention of Bessey (1915),
Sinnott and Bailey (1914) and others that in the angiosperms
herbs have been derived from woody plants. c

It has been shown previously in this paper that plants
which manufacture glycerides of the highest iodine numbers,
alkaloids with the largest molecular weights, volatile oils
With the highest specific gravities and lowest refractive
indices perform the most difficult work; therefore, it may
be concluded that herbs which are higher evolved than shrubs
or trees, also perform the most difficult work.

The chemical data used in Table II are condensed and re-
arranged according to plant form from McNair (1934). The
following families used in the calculations are considered
as consisting mainly of trees: Bombacaceae, Caricaceas, Dip=-
terocarpaceae, Lecythidaceae, Moringaceae, Palmae, Rhizoph-
oraceae and Winteranaceae; the families consisting mostly of
shrubs and trees are Anacardiaceae, Anonaceae, Araliaceaé,
Bignoniaceae, Bixaceae, Burseracese, Caryocaraceae, Cochlo~
spermaceae, Combretaceae, Ebenaceae, Erythroxylaceae, Flac-
Monimiaceae, Moraceae, Myristicaceae, Myrtaceae, Ochnaceae,
Olacaceae, Oleaceae, Proteaceae, Salvadoraceae, Sapindacéae,

Sapotaceae, Simarubaceae, Staphyleaceae, Symplocaceae, Tili=

aceae, Vochysiaceae and Zygophyllaceae; mostly shrubs, Apo-
cynaceae, Asclepiadaceas, Humiriaceae, Loranthacese and Vit-

aceae; the families consisting mostly of herbs, shrubs and
trees, Loganiaceae, Menispermaceas, Phytolaccacese, Rubiace-
ae, Sterculiaceae and Verbenaceas.

Intensity of Assimilation. - Although the amounts of mat-
erials such as ligneous matter, sugars and chlorophyll assi-
milated in plant structures may not have a bearing on evolu-
tion, yet the rapidity of assimilation of some substances in
plants apparently does have a bearing on evolution. For in-
stance, by a rapid rate of metabolism in those plants which
produce fruit only once in their lives, the foods and re-
serve materials necessary for fructification and seed prod-

1941 McNair, Energy & Evolution 4)

uction are produced in sufficient quantity more rapidly and
earlier reproduction and death are thereby made possible. It
is these plants of rapid metabolism which generally occupy
the most highly evolved positions on the plant family tree.

Length of Plant Maturation Period. - In relation to the
length of plant maturation period plants may be divided into
two categories, those plants which bear fruit only once dur-
ing their lives (monocarpic), and others which do so several
times or frequently (polycarpic).

Plants which bear fruit only once generally tend to have
the shortest longevity consistent with a normal reproductive
period (Molisch, 1938). These germinate, develop and, as
soon as they are fully grown, store reserve materials and
then proceed to fructification, seed production and death.

This principle of the greatest possible abbreviation of a
natural life does not apply to polycarpic plants, however,
for we know that many such forms, particularly trees and
shrubs, continue to live a long while, sometimes for many
centuries after attaining maturity.

As monocarpic annual herbs have shorter plant maturation
periods than polycarpic perennial shrubs and trees, and as
annual herbs also occupy, in general, more recent evolution-
ary positions, therefore it can be concluded that these
plants with shorter plant maturation periods work harder and
are higher evolved than the longer maturing shrubs and trees

PLANT PARTS. -- Length of Fruit Maturation Period. - The
time consumed between the moment that the ovule becomes fer-
tilized and the moment that the seed becomes viable varies
greatly among plants. The length of this maturation period
may require from a few weeks (Tradescantia virginica) to
from two to three years as in the Pinaceae and Myrtaceae
(Cheel, 1931). The shortest periods are found in monocarpic
species and among the monocarpic species the ephemeral or
annual plants generally require less time than the peremni-
als. As the ephemeral or anmal plants are herbs we have a-
nother instance where the greatest intensity of work is
shown by plants highest in evolutionary position.

Flowers, Leaves and Stems. - The rate of metabolism ap-
pears to have a definite relation to the evolutionary posi-
tion not only of the plant forms themselves (as shown above)
but also of plant parts. Intensity of respiration can indeed
be regarded, to a certain degree, as a measure of intensity
of metabolism, since we know, for example, that flowers ex-
hibit an unusually high rate of respiration, leaves less so
and stems stiil less, and that the longevity of these organs
parallels these rates, i. e., flowers live only a short per-
iod, leaves for a longer period and stems still longer.

The systematic position of a plant in evolution is deter-

mined mainly through flower differences. In the flower

4a PHYTOLOGIA _ Nols 2, Sane

structure and function many more changes and more rapid
changes have taken place than in either the leaves or stems.

The period of longevity of an angiosperm flower is here
considered as extending from the first opening of the blos—_
som to the final withering or shedding of its important
parts (calyx, stamens).

So considered, the duration of flowers among various
plants lasts from three hours to three months. If the plant
puts forth only one flower (which is considered by some to
be a more primitive condition than an inflorescence) annual-
ly, as is true of Galanthus, Moneses uniflora, Paris quadri-
folia and the different species of Trillium, or when the
flowers are only two or three in number, as in Cypripedium
calceolus and the tropical orchids of the genera Oncidiun,
Stanhopea and Cattleya, these single flowers remain fresh
and open a long time.

Evolution, we know, does not necessarily Licoiv all
parts of the flower at one time or in the same direction.
One flower’ part may be advancing while another is stationary
or retrograding. Because of this, all short-lived flowers
are not all evolved to the same uniform degree of advance=
ment, but nearly all of them are found on short-lived herbs
(although staminate aments are on trees), and herbs consti-
tute the most recent evolved plant form. Consequently it can
be concluded that in general the flowers of shortest dura-
tion which also exhibit the most intense metabolism occupy
the most advanced phylogenetic positions. ;

Leaves. - As compared with flowers, leaves are of greater
duration; compared with the entire plant, however, they are
rather short-lived, sometimes conspicuously so, except in
those cases where their death is approximately simultaneous
with that of the entire plant.

Under the most favorable circumstances the leaves of ann-
uals attain the age of the plants which bear them, usually
that of only one vegetative period, namely, several months.

The leaves of gymmosperms vary greatly from one hundred
years (those of Welwitschia) to one year (Cupressus, Ephedra,
etc.). The longevity of the leaves of the monocotyledons al-
so varies from a number of years, as in the palms, to less
than one year (Amaryllis, etc.). Although some of the leaves
of the dicotyledons live as long as five years, many are in
the one-year class.

There is, therefore, apparent decrease in the length of
life (with an accompanying increase in intensity of work) of
leaves as the plants on which they occur advance in evolut-
ionary position.

Stems. - In general, it may be said, aside from except-
ions, that the life of stems of herbs is relatively shorter
than that of shrubs and trees as a whole. In the case of

1941 - MeNair, Energy & Evolution 43

gymnosperms, woody etems and great longevity attain the ult-
imate dominace. And among monocotyledons long life prevails
in the palms and shrubby forms, while short-lived annuals

‘are relatively rare. In the dicotyledons short-lived anme.-

are more common.

It seems highly probable, therefore, that the position in
evolution of some spermatophytes is indicated by the kinetic
energy of their life cycles and the potential energy of some
of their chemical compounds.

Summary

The object of this paper is to develop the theory that
species formation occurs during periods of increased activ-
ity, that plants which do the hardest (most difficult) work
have evolved to the highest positions; that in this regard
quality of products is more important than quantity; and
that as morphological structures evolve from simple to com-
plex, so plant chemical compounds evolve from simple to com-
plex.

Chemically each species is in a state of mobile equilib-
rium between reversible reactions.

The existence and permanency of a species depends upon
the existence of constant external and internal conditions
and shows a fixed ability to synthesize characteristic
chemical compounds.

The splitting off of new species falls within the times
of greater activity.

There is a tendency to increased complexity both in mor-
phology and chemical compounds with evolutionary progress.

The stable conditions in the tropics are not as liable
to produce these changes as the fluctuating conditions in
the temperate zones.

Alkaloids are of greater molecular weight in temperate
regions and likewise in the higher evolved tropical plants.
Therefore higher evolved plants carry on more difficult
worke

Glycerides produced in temperate zones and in the higher
evolved tropical families have greater unsaturation (higher
iodine values), and their fatty acids have higher molecular
weights than the average tropical products. Thus more dif-
ficult work is performed by the higher evolved plants.

Volatile oils of temperate families and tropical famil-
ies highest in evolutionary placement have constituents of
low saturation, more aromatic compounds, or more sulphur
and nitrogen compounds with small amounts of substances of
low molecular weight or small quantities of terpenes or
bodies of the aliphatic (fatty) series. Because it requires
more energy to form substances of high than of low molecu-

et PHYTO. 0'S "th Vol. 2, noe 2

lar weight, aromatic than aliphatic compounds, it can be
concluded that plant families which manufacture these sub-
stances can be said to have reached an advanced place in ev-
olution.

Volatile oil tropical acids and alcohols likewise show
that the highest evolved tropical families form the acids
and alcohols of greatest molecular weight and therefore pro-
mote more difficult work. é

Trees are shown to do less difficult work than shrubs or
herbs through a study of their alkaloid, glyceride and vola-
tile oil production.

Plants that produce fruit only once in their lives (annu-
al herbs) have a more rapid rate of metabolism than the
polycarpic shrubs and trees. This rapid rate of metabolism
is therefore indicative of more difficult work of -the more
highly evolved plant forms.

The length of the fruit maturation period is shorter in
annual herbs than in shrubs and trees and therefore is an
indication of the more difficult work carried on by the
highly evolved herbs in apposition to that of the more prim
itive shrubs and trees.

Among plant parts longevity is a measure of metabolism.
In this regard flowers greatly exceed leaves, and leaves ex-
ceed stems. The shortest lived flowers are produced by annu-
al herbs and consequently exhibit the most intense metabol-
ism, the most difficult work and occupy in general the most
advanced phylogenetic positions.

It seems highly probably, therefore, that the position in
evolution of some spermatophytes is indicated by the kinetic

energy of their life cycles and the potential energy of some

of their chemical compounds.
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a

1941 McNair, Energy & Evolution . 45

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Table I

Peete i ee ee Ni Sas a ee
Substances Properties Producing Climates

Tropical Temperate

(iodine number ) 85.56.) 124.00

(molecular weight) 293 .00

Volatile oils (specific gravity) 0.9188 -

Volatile oils (refractive index) 1.4932

47

Substances

lycerides

McNair, Energy & Evolution

1941

Properties

(iodine numbers )
(molecular weight)
(specific gravity )

(refractive index)

Table II

Families consisting essentially of (average va lues )

shrubs and herbs and herbs
trees shrubs

66 94 96 102 117

191 348 236 231 307
£8878 9175 9010 9340 887

1.4650 1.4938 1.4295 1.4918 1.490

48 PHYTOLOGIA Vol. 2, noe 2
Table III

Camparison of Some Characteristics of Primitive
and Higher Evolved Plants

More Primitive Plants Higher Evolved Plants

More stable environments More fluctuating environ-
@.g. tropics and water ments @.g. temperate
zone and land

TEnvironment

Alkaloids Alkaloids
Lower. average molecular Higher average molecular
weights weights |
Glycerides Glycerides
Lower average melting Higher average melting
points points
Lower average iodine num- Higher average iodine
bers numbers .
2 Lower average molecular Higher average molecular
§ weights weights
rs}
2. Volatile Oils Volatile Oils
S Lower average specific Higher average specific
= gravity grevity
s More aliphatic compounds Fewer aliphatic com
| pounds
© Higher average refrac- Lower average refractive
6 “tive index index
Fewer compounds of high More compounds of high

molecular weight molecular weignt

Ener gy Energy

less energy required to More energy required to
make compounds of lower make compounds of high-
molecular weight cited er molecular weight
& above cited above
© Volatile oil alcohols and Volatile oil alcohols
ta volatile oil acids with and volatile oil acids
lower heat combustion with higher heat’ com

bustion

McNair, Energy & Evolution 49

Table III - (continued)

More Primitive Plants Higher Evolved Plants

4
es
A f

Longevity of plant parts

Energy (continued )
Therefore less energy
required to make

Trees more primitive
Alkaloids
- Lower average molecular
weights
Glycerides
Lower average melting
points
Lower average iodine
numbers
Lower average molecular
weights
Volatile Oils
Fewer compounds of high
molecular weight

Plant form

Intensity of assimilation
Polyearpic plants have
less

3 &
o3
PS
aa
Ss 8
® -
» @
Ss a
Ho

Length of plant matura-

tion period
Polycarpic plants have
greater
Perennial shrubs & trees
Length of fruit matura-
tion period
Polycarpic perennials
(longer )

Longevity of flowers
Longer

Energy (continued)
Therefore more energy
required to make

Herbs more advanced
Alkaloids
Higher average molecu-
lar weights
Glycerides
Higher average melting
points
Higher average iodine
numbers
Higher average molecu-
lar weights
Volatile Oils
More compounds of high
moleculer weight

Intensity of assimilatio
Monocarpic plants have
greater

Length of plant matura-

tion period

Monocarpic plants have
less
Annual herbs

Length of fruit matura-

tion period
Monocarpic ephemeral or

annual (shorter)

Longevity of flowers

Shorter

50 PHYTOLOGIA Vol. 2, noe 2

Table III - (continued)

More Primitive Plants Higher Evolved Plants

Longevity of leaves - Longevity of leaves
Longer Shorter

Longevity of stems
Shorter

Longevity of stems
_ Longer

PLANT NOVELTIES

- Harold N. a

ALOYSIA FONCKI (R. A. Phil.) Moldenke, Suppl. List Invalid
Names 5, hyponym (1941), comb. nov.
Lippia Foncki R. A. Phil., Anal. Univ. Chile 90: 620.1896

BAILLONIA AMABILIS var. PUBESCENS Moldenke, vare nove

Haec varietas a forma typica speciei recedit ubique dense
breviterque pubescentibus vel puberulis.

This variety differs from the typical form of the species
in having ite twigs, branchlets, petioles, and both leaf=
surfaces densely short~pubescent, the racis and bracts den-
sely puberulent or short=pubescent, and the calyx more or
less puberulent.

The type of this variety was collected by my good friend
and respected colleague, Dr. Frederico Carlos Hoehne [Com.
Rondon 4739] at Triumpho, Rio S. Lourengo, Mattogrosso,
Brazil, in February, 1911, and is deposited in the herbarium
of the Departamento do Botanica do Estado, So Paulo.

CALENDULA OFFICINALIS f. PROLIFERATA Moldenks, f. nov.

Heec forma a forma typica speciei recedit ramlis 4--15
em. longis in axillis bracteolorum involucri ornatis.

This form differs from the typical form of the species in
bearing one or more short branchlets which issue from the
axils of involucral bractlets beneath the main head of flow-

al a

1941 ; Moldenke, Plant Novelties ! 51

ers and which each bear one or more miniature leaves and a
small terminal head of orange flowers.

The type of this form was collected by H. Ne Moldenke
(no. 11,830) in cultivation at Mount Vernon, Westchester Co,,
New York, on September 20, 1941, and is deposited in the
herbarium of the Bailey Hortorium at Ithaca, New York.

CITHAREXYLUM KUNTHIANUM Moldenke, nom. nov.

Citharexylum tomentosum H.BeK., Nov. Gen. & Sp. Pl. 2:
258. 1818 aoe C. tomentosum Poir. in Lam., Encycl. Méthe
Bot. Suppl. 2: 368. 1811].

JUNELLIA ILLAPELINA (Re Ae Phil.) Moldenke, Suppl. List In-
valid Names 9, hyponym (1941), comb. nov.
Verbena illapelina R. A. Phil., Anal. Univ. Chile 90:
612. 1896.

JUNELLIA THYMIFOLIA (Lag.) Moldenke, Suppl. List Invalid
Names 10, hyponym (1941), comb. nov.
Ver bena thymifolia Lage, Gen. & Sp. 18--19. 1816.

LAMPAYA CASTELLANI Moldenke, sp. nove

Frutex; caulis prostratis abbreviatis; ramis numerosis
brevibus erectis vel adscendentibus dense viscidulo-puberu-
lis; nodis valde annulatis; foliis numerosis confertis de-
cussato-oppositis; petiolis abbreviatis viscidulo-puberulis
vel glabratis, ad basin subamplexicaulis; laminis coriaceis
brunnescentibus ellipticis acutis integris, glanduloso-
punctatis utrinque glabratis aromaticis, ad basin attenuat-
is; floribus fasciculatis; bractéolis parce pilosulis.

Dwarf shrub; stem prostrate, woody, gnarled, abbreviat-
ed; branches numerous, short, erect or ascending, quadrang-
ular, densely viscidulous-puberulent, becoming scabrous
through attachment of sand particles; nodes very conspicu-
ously annulate; principal internodes mich abbreviated, 1--
12 mm. long; leaves numerous, crowded, decussate-opposite;

' petioles abbreviated, 1.5--3 mm. long, sub-clasping at

base, viscidulous<puberulent and eventually scabrous, or
glabrate; blades coriaceous, uniformly gray-green on both

surfaces, brunnescent in drying, elliptic, 6--14 mm. long,

3—-7 mm. wide, acute at apex, entire, somewhat attenuate
into the margined petiole at base, glandular—punctate,
glabrate on both surfaces, aromatic when crushed; midrib
flat above, sharply prominent beneath; secondaries and
veinlet reticulation not visible on either surface; flowers
borne in fascicles of 1 or more near the tips of the
branches; bractlets deltoid, 1.5--2 mm. long, sparsely pil-
osulous; calyx tubular, about 5 mm. long, sparsely pilosul-
ous outside, its rim 2-lipped, S-lobed, the lobes involute

52 PHYTOLOGIA Vol. 2; now 2

at the tip; corolla zygomorphic, hypocrateriform, its tube
curvate, 10--12 mm. long, pilose within, glabrous outside,
its limb 4—parted; stamens 4, didynamous; filaments abbrevi-
ated; stigma obliquely 2-lobed; ovary glabrous, 2-celled;
ovules 1 per cell, apparently basally attached.

The type of this species was collected by my good friend,
Alberto Castellanos -- in whose honor it is named -- between
Cienaga Grande and Cerillos, Jujuy, Argentina, on February.
5, 1937 [Herb. Mus. Argent. Cisnc. Nat. 20,161] and is dep-
osited in the Britton Herbarium at the New York Botanical
Garden.

LAMPAYA HIERONYMIE Schum. & Moldenke ex Moldenke, Suppl. List
Invalid Names 4, hyponym (1941), sp. nov.

Frutex; ramis crassis tetragonis glabris; nodis valde an=
nulatis argute ampliatis; foliis numerosis decussato-opposi-
tis; petiolis obsoletis vel usque ad 3 mm. longis crassis
glabris non-amplexicaulibus; laminis coriaceis oblongo-
ellipticis obtusis vel subacutis integris, ad basin acutis,
utrinque glabris.

Erect shrub; stems heavy, tetragonal; branches numerous,
heavy, tetragonal, glabrous; nodes very conspicuously annu-
late, sharply ampliate on the branches; principal internodes
1--2.5 cm. long; leaves decussate-opposite, numerous, borne
on prominent corky sterigmata; petioles obsolete or to sev-
eral mm. long, heavy, glabrous, not clasping at base; blades
thick-coriaceous, 2.5--4 cm. long, 7--12 mm. wide, oblong-
elliptic, obtuse or subacute at apex, entire, acute at base,
glabrous on both surfaces, the epidermis wrinkling in dry-
ing; midrib flat or subimpressed above, rounded=prominent
beneath; secondaries and veinlet reticulation hidden; in-
florescence not séen.

The type of this species was collected by G. Hieronyms
and G. Niederlein (no. 191) at Las Cortaderas, between El
Penou and El Jaguel, Rioja, Argentina, on February 22, 1879,
and is deposited in the herbarium of the Botanisches Museum
at Berlin. The collectors report the vernacular name
"lampaya" and say that the plant is used to was wounds and
in the treatment of gonorrhoea.

LANTANA BRITTONI Moldenke, spe nove

Frutex scandens; ramis acutiuscule tetragonis obscure
aculeatis plusminus pilosis glabrescentibus; internodiis 6-
longatis; nodis distincte piloso-annulatis; foliis opposit-
is; petiolis gracilibus sparse obscureque pilosulis; laminis
tenuiter membranaceis fragilibus nigrescentibus ovatis acut-
is vel breviter acuminatis regulariter serratis, ad basin
subcuneatis, utrinque glabris vel subglabris vel supra ob=
scure scabridis; inflorescentiis axillaribus capitatis.

1941 Moldenke, Plant Novelties 53

Trailing shrub. or climbing to a height of over 7 me; main
stem about 1.5 cm. in diameter at base; branches rather ac-
utely tetragonal, obscurely prickly, more or less scattered-
pilose toward the apex, becoming glabrous in age; principal
internodes elongate, 8--10 cm. long; nodes distinctly annu-
late,. with a band of pilose hairs on the youngest parts;
leaves decussate-opposite; petioles slender, about 2.5 cm.
long, sparsely and obscurely pilosulous; leaf-blades thin=-
membranous, very fragile in drying, uniformly green on both
surfaces, nigrescent in drying, or slightly lighter beneath,
ovate, about 15 cm. long and 6.5 cm. wide, acute or short~
acuminate at apex, subcuneate at base, uniformly serrate a-
long the margins with broad more or less rounded (and minu-
tely apiculate) teeth, glabrous or subglabrous on both sur-
faces or very slightly and obscurely roughened above; midrib
slender, flat above, prominent beneath; secondaries slender,
6 or 7 per side, arcuate-ascending, flat above, slightly
prominulous beneath; veinlet reticulation obscure or indis-
cernible above, abundant and plane beneath; inflorescence
axillary, 2.5--6 cm. long, capitate, densely many-flowered;
peduncles slender, 1.5--4.5 cm. long, glabrate or very ob-
scurely puberulent, sometimes incrassate or ampliate at a-
pex; bractlets small, linear, not very conspicuous, 2--4 mm.
long, minutely strigillose; calyx campanulate, about 2 mm.
long; corolla yellow, its tube about 6 mm. long, its limb a-
bout 5 mm. in diameter, the three upper lobes smaller than
the lower lobe.

The type of this species was collected by William Harris
and Nathaniel Lord Britton (no. 10,541) at Tweedside, below
Moody's Gap, Jamaica, on September 10, 1908, and is deposit-
ed in the Britton Herbarium at the New York Botanical Gard-
en. It is named, in grateful appreciation, in honor of Dr.
Britton, who assigned to it the cheironymous name, Lantana
scandens Britton [not L. scandens Moldenke, Phytologia 2:
18--19. 1941].

LANTANA INVOLUCRATA var. ODORATA (L.) Moldenke, comb. nov.
Lantana odorata L., Syst. Veg., ed. 12, 418. 1774.

LIPPIA SCHLECHTENDALII Moldenke, nom. nov.
Dipterocalyx scaberrima Schlecht., Linnaea 26: 647. a
(not Lippia scaberrima Sond., Linnaea 23: 87. 1850].

NASHIA ARMATA (Urb.) Moldenke, comb. nov.
Lippia armata Urb. in Fedde, Repert. 16: 194. 1922.

NASHIA MYRTIFOLIA (Griseb.) Moldenke, comb. nov.
Lippia myrtifolia Griseb., Cat. Pl. Oub. 215. 1866.

54 PHY TOLOGLA Vol. 25 noe 2

NASHIA NIPENSIS (Urb.) Moldenke, comb. nov.
Lippia nipensis Urb. in Fedde, Repert. 20: 344. 1924,

NASHIA SPINIFERA (Urb.) Moldenkse, comb. nov.
Lippia spinifera Urb. in Fedde, Repert. 18: 195. 1922.

NASHIA VARIIFOLIA (Urb.) Moldenke, comb. nov.
Lippia variifolia Urb. in Fedde, Repert. 20: 344. 1924.

PYGMAEOPREMNA HERBACEA (Roxb.) Moldenke, comb. nov.
Prema herbacea Roxb., Hort. Beng. 46, hyponym (1814),
Fl. Ind. 3: 80, 1852.

PYGMAEOPREMNA SUBACAULLS (F. Muell.) Moldenks, comb. nov.

Tatea subacaulis F. Muell., Trans. Roy. Soc. S. Austral.
63 34. 1883.

STACHYTARPHETA CEARENSIS Moldenke, sp. nove

Herba perennis; ramis subtetragonis adpresso-puberulis
vel pubescentibus; foliis oppositis brevipetiolatis; petiol-
is gracilibus adpresso-strigilloso-pubescentibus; laminis
firme chartaceis obovatis, ad apicem rotundatis, supra medi-
am uniforme crasseque dentatis, ad basin cuneatis, marginib-
us subrevolutis, supra subscabridis et obscure strigillosis
glabrescentibus, subtus leviter adpresso-puberulis.

Perennial herb; stems appressed—pubescent. or puberulent,
less so in age, subtetragonal; leaves decussate-opposite,
short-petiolate; petioles slender, about 4 mm. long, appres-
sed-strigillose-pubescent; blades firmly chartaceous, obov-
ate, 3--3.3 cm. long, 1.6--1.8 cm. wide, rounded at apex,
rather uniformly coarsely dentate above the middle with
broad and rounded or subacute teeth, cuneate at base, sub-
revolute along the margins, subscabridous and obscurely
strigillose with very minute caducous hairs above, glabres-
cent in age, lightly appressed-puberulent beneath; midrib
slender, impressed above, prominulous beneath; secondaries
slender, 4 or 5 per side, arcuate-ascending, somewhat im-
pressed above, prominulous beneath; peduncles short, tetrag-
onal, about 1.5 cm. long, rather densely puberulent with
cinereous or sordid hairs; floriferous portion of spike 5.5
cme long (in immature specimen?); rachis stout, densely cin-
ereous~puberulent, sharp-angled; bracts oblong, 8--9 mm.
long, about 3 mm. wide from base to near the apex, then ab=
ruptly long-acuminate (the acumination about 2 m. long), a-
bundantly puberulent on the back, not at all scarious; flow-
ers closely imbricate; calyx tubular, about 10 mm. long and
5 mm. wide, prominently S-ribbed or -plaited, its rim short-
ly S-toothed; corolla-tube slightly exceeding the calyx,
glabrous outside, its limb spreading, about 8 mm. wide.

1941 - Moldenke, Plant Novelties 55

The type of this species is a poor fragment collected by
Freire Allem&o (no. 1152) in Ceard, Brazil, and is no. 32247
in the herbarium of the Museu Nacional at Rio de Janeiro.

VERBENA GALAPAGOSENSIS Moldenke, sp. nov.

Herba, in siccitate ubique nigrescens; ramis ramulisque
gracilibus acute tetragonis saepe sulcatis striatisque spar-
sissime pilosis; nodis annulatis; foliis oppositis sessilib-
us linearibus vel angustissime lanceolatis utrinque adpres-
so—pilosis, ad basin subamplexicaulibus; inflorescentiis
terminalibus spicatis laxe multifloris.

Herb, nigrescent throughout in drying; stems and branches
slender, acutely tetragonal, often sulcate and striate be-
tween the angles, very sparsely pilose with short whitish
widely scattered antrorse hairs; nodes annulate; principal
internodes 1--5.8 cm. long; leaves decussate-opposite, sess—-
ile, sub-clasping at base, linear or very narrowly lanceol-
ate, 5--15 mm. long, rather abundantly pilose with appressed
antrorse hairs on both surfaces; midrib and veinlet reticu-
lation indiscernible; inflorescence terminal, spicate, 4--15
cme long, loosely many-flowered; peduncles slender, similar
to the branches in all respects, nigrescent in drying, acut-
ely tetragonal, longitudinally striate, very sparsely scatt-
ered-pilose, 1~-3 cm. long; bractlets ovate-lanceolate,
about 2.5 mm. long, ciliolate-margined, acuminate at apex,
usually glabrate or very obscurely pilosulous except for the
margins; mature flowers and fruit not seen.

The type of this remarkable species was collected by Alb=
an Stewart (no. 3318) at Cowley Bay, Albemarle Island, Gala-
pagos Islands, on August 10, 1905, and is deposited in the
Britton Herbarium at the New York Botanical Garden. The col-
lector stated that the species is common at 2000 feet eleva-
tion. [t has hitherto been confused with V. litoralis H.BK.

VERBENA RUSSELLII Moldenke, sp. nov.

Herba; ramis gracilibus argute tetragonis ubique minutis-
sime obscureque puberulis; nodis annulatis; foliis opposit-
is, supremis sessilibus; petiolis alatis hirsuto-pubescenti-
bus; laminis inferioribus ellipticis vel ovato-ellipticis a-
cutis irregulariter inciso-dentatisa utrinque densiuscule
glanduloso-hirsutulis, ad basin longo-acuminatis, marginibus
subrevolutis; laminis supremis lineari-lanceolatis irregu-
lariter inciso-dentatis vel integris utrinque dense glandu-
losomhirsutulis; inflorescentibus terminalibus spicatis.

Herb, not at all nigrescent in drying; stems slender,
sharply tetragonal, very minutely and obscurely puberulent
throughout; nodes annulate; principal internodes 2--6.5 cm.
long; leaves decussate-opposite, the uppermost ones sessile,
the lower and older ones petiolate; petioles to 1 cm. long,

56 PY EOL OF 42.4 Vol. 25 noe 2

winged, hirsute-pubescent; lower leaf-blades elliptic or ov=
ate-elliptic in outline, 1--4 cm. long, 8--15 mm. wide, ac-
ute at apex, irregularly incised-dentate along the margins,
long-attenuate into the petiole at base,rather densely hirs-
utulous and glandular on both surfaces, especially beneath,
with stiff whitish hairs, the margins slightly revolute in
age; midrib and secondaries flat or subimpressed above,
prominent beneath} upper leaves linear-lanceolate, sessile,
5--15 mm. long, irregularly incised-dentate or entire, dens
ely glandular—hirsutulous on both surfaces, with a very
prominent midrib beneath; inflorescence terminal, spicate,
peaniculately branched, the branches to 20 cm. long, densely
many~flowered; peduncles slender, sharply tetragonal, minu-
tely puberulent, often elongate; rachis densely glandular-
puberulent; bractlets lanceolate, 2--2.5 mm. long, attenuate
to the sharply acute apex, densely glandular-pubescent and
ciliate-margined; calyx tubular, about 2 mm. long, densely
glandular=—puberulent, its rim S-apiculate; corolla small,
its tube 3--3.5 mm. long, very slender, its limb 2--3 mm.
wide in anthesis.

The type of this species was collected by Joseph Nelson
Rose, Paul Carpenter Standley, and Paul George Russell (noe
14,850) in a moist field in the vicinity of Culiacan, Sinal-
oa, Mexico, on April 21, 1910, and is deposited in the Brit-
ton Herbarium at the New York Botanical Garden. It is named
in honor of Paul Yeorge Russell and was annotated by Lily M.
Perry in 1922 as "aff. V. officinalis L."

VERBENA STEWARTII Moldenke, sp. nov.

Herba plusminus nigrescens; ramis ramulisque gracillimis
subfiliformibus argute tetragonis ubique glabris nitidisque;
nodis annulatis; foliis oppositis sessilibus, supremis lin-
earibus, inferioribus 2--3-lineari-lobatis, obtusis revolut-
is utrinque plusminus sparse adpresso=pilosis; inflorescent-
iis terminalibus spicatis paucifloris.

Herb, more or less nigrescent in drying; stems and
branches very slender, the latter almost filiform, sharply
tetragonal, glabrous and shiny throughout; nodes annulate;
principal internodes mostly elongated, 2--6 cm. long; leaves
decussate-opposite, sessile, the upper ones linear, the low-
6r ones with 2 or 3 linear widely divergent lobes, revolute-
margined, blunt=pointed, more or less sparsely scattered=-
pilose with appressed whitish antrorse hairs on both surf-
aces; midrib prominent beneath; iniloréscence terminal,
spicate, rather few-flowered, dense t ward the apex and dur-
ing anthesis, the lower flowers often scattered after an-
thesis; peduncles slender, elongated, 6.5--7.5 cm. long,
glabrous and shiny; rachis filiform, glabrous and shiny or
very obscurely scattered-pulverulent; bractlets lanceolate,

1941 Moldenke, Plant Novelties 7

1.5--2 mm. long, acuminate at apex, glabrous except for the
Ciliolate margins; calyx tubular, about 2 mm. long, minutely
appressed=-puberulent; corolla barely exceeding the calyx,
its tube usually only about 2 mm. long, its limb about 1.5
mm. wide.

The type of this remarkable species was collected by Alb-
an Stewart (no. 3320) -- in whose honor it is named — at
Tagus Cove, Albemarle Island, Galapagos Islands, on March
27, 1906, and is deposited in the Britton Herbarium at the
New York Botanical Garden. The collector states that the
species is common in lava beds at 300 fest altitude. It has
hitherto been confused with V. litoralis H.B,K.

ADDITIONAL NOTES ON THE GENUS ARGIPHILA -- VII

Harold N. Moldenke

The following notes constitute a continuation of those
published in Phytologia 1: 182--208, 222--240, and 248--272
(19357), 289--504 (1938), 364--368 (1959), and 372-400
(1940). An additional herbarium abbreviation herein employ-
ed and not previously explained is "Ra", which stands for
the herbarium of the Museo Nacional de Historia de Buenos
Aires.

Bentham in Benth. & Hook. f., Gen. Pl. 2%: 1151 (1876)
recognized ahout 50 species in the genus Aegiphilea. Today we
recognize 158 species and varieties. An additional misspell-
ing of the generic name is Aegyphylla Jacq. ex Moldenke,
Suppl. List Invalid Names 1, in syn. (1941). Junell in Symb.
Bot. Upsal. 4: 83, fig. 133 (1934) shows the structure of
the gynoecium and gives important notes on the floral morph-
ology of the group.

49a. AEGI PHILA ACULEIFERA Moldenke.

Austin Smith has furnished us with detailed notes about
this species, including some characters not before recorded.
He states that it is a tree 4--12 m. tall, of open growth,
inhabiting the cloud forests of the Caribbean watershed and
thickets on the edges of woodlands, where it is “quite a
common tree", at altitudes of 4200 to 7000 feet, often in
clay-loam and open shade. The bark is neutral-brown, gray,
or gray~brown, a little glandularly roughened. The leaves
are "barely stiffened", nearly glabrous, dark dull-green a-
bove, the venation prominent on the under surface. The under
surface of the leaves is dark-green, not shiny, with a faint

58 FP HY TO. FOG: E' a Vol. 2, no. 2

tomentum, and has the midrib there cream-colored. The buds
are roundish, pale- or grayish-green, more or less woolly-
tomentose. The flowers, stems, pedicels, and petioles, acc-
ording to this fine collector, are more or less woolly
tomentose. The expanded flowers are 20 mm. long and 10--13
mne Wide. The calyx is campanulate, pale-green, and glandu-
lar, the sepals 4 in number. The corolla is pure-white,
creamy-white, or cream-colored, glandular, cruciform, and
fleshy, the tube about 12 mm. long, the lobes recurved. The
4 stamens are "as long as the petals" and are placed alter-
nately with them. The filaments are slender and white, the
anthers large, brown, compressed, bilobed or "2-segmented".
The style is either much longer than or mich shorter than
the stamens depending on whether the plant is male- or
female-predominant. The stigma is mich shorter than the
style, 2-parted. The fruits ripen pale-yellow and semi-
pellucid. The tree is in full inflorescence in June and
Auguste |

Additional citations: COSTA RICA: Alajuela: A. Smith
4210 (F, F).» A-252 (F, N), Ho1104 (F, N, N).

6. AEGIPHILA ANOMALA Pittier.

Austin Smith has furnished detailed notes about this rare
species, stating that it is a tree 55--50 feet tall, of open
spreading growth, often spreading to 50 feet, the base 1 1/2
to 5 feet in diameter, of open forests on wooded hillsides
in light shade and in clay~loam soil on the Continental Div-
ide and within the zone of the Pacific cloud-forest, often
in semi=shade on clay ridges, at altitudes of from 4500 to
5800 feet. On one label he states that it is "common" and on
another "scarce so high, more common westward". The sap is
not milky. The bark is brown, thick, of cork-like structure,
well-sutured, much roughened and shaggy by raised and
slightly curled granulated narrow-oblong strips. The cambium
is cream-yellow. The buds are round and gray-green, the un-
opened clusters grayish-yellow, the open ones showing brown-
ish; the flower-buds are buffy-yellow in color. The leaves
are rather lax and soft or slightly stiffened, mich disfig-
ured by the wind, clustered, bright—green or the newer ones
light-green, velvety to touch. The calyx-cup is light-green.
The flowers resemble those of Capparidaceae, 14--20 mm. wide
when expanded, with a faint odor. The corolla is pure-white,
"resembling Cleome", "springing independently of the base
cluster from stem", the tube very small and urceolate, "can-
aliculated in throat". The corolla-limb opens flat, 2 of the
S petals sometimes recurving. The 5 stamens are long-exsert-

ed and curved, the anthers brown, and the style short. "Seed

capsules suggesting Capparidaceae." It has been collected in
flower and fruit in July and August and has been confused

fae ae
eer

1941 | Moldenke, Notes on Aegiphile 59

with A. Valerii.

Additional citations: COSTA RICA: Alajuela: A. Smith 138
(F), A.242 (F, N, N--photo, Z--photo), A379 (Fy Fs F).
Limon: H. Pittier s.n. [Herb. Instit. Physico-geogr. Nat.
Costaric. 16,711] (F=-photo of type).

70. AEBGIPHILA BARBADENSIS Moldenke.
Additional citations: BARBADOS: Warming 101 (F--photo of

type).

3. AEGIPHILA BOGOTENSIS (Spreng.) Moldenke.

This species has been collected in anthesis in September,
December, and January, inhabiting low woods on slopes. It is
said by Miss Mexia to be a small tree to 6 m. tall, with
yellow-white and waxy flowers.

Additional citations: COLOMBIA: Cundinamarca: Bonpland s.

T2] Wel.

n. (F=-fragment); Triana ae [1] (Je), 3743/4 Jel.

Tolima: Purdie s.n. (F=-photo). ECUADOR: Pichincha: Mexia
1683 (N).

80. ABGIPHILA BOLIVIANA Moldenke.

Additional citations: BOLIVIA: Santa Cruz: Steinbach 6437
(F--photo of cotype), 7071 [Herb. Mus. Nac. Hist. Nat.
Buenos Aires 30/2720] (Ra), 7289 (Z--photo).

25. ABGIPHILA BRACHIATA Vell.

Jorgensen states that the species grows to be a tree 4 m
tall, with sulphur-yellow flowers, blooming in September. He
states that it is very common in Pseraguay. |

Additional citations: BRAZIL: State undetermined: Sellow
1269 [Macbride photos 17,590] (F--photo, Kr--photo). PARAGU-
AY: Jérgensen 3662 (F, F, N).

120. AEGIPHILA BRACTEOLOSA Moldenke.

The species is said by Ducke and by kKrukoff to inhabit
secondary not~inundated forests or terre firma. It is des-
cribed as a shrub to 12 feet tall and has been confused with
A. arborescens (Aubl.) Gmel. [=A. integrifolia (Jacq.)
Jacks.].

Additional citations: BRAZIL: Amazonas: Ducke 444 (F);
Krukoff 5060 (F, Mi).

102. AEGIPHILA BUCHTIENII Moldenke.
The species has been collected at an altitude of 800 m.,

in anthesis in Jamary. .
Additional citations: BOLIVIA: La Paz: Buchtien 1716 (N).

99. AEGI PHILA CANDELABRUM Brig.
The specific name of this species is sometimes lower-

60 PHYTOLOGIA Vol. 2, no. 2

cased, but was written with a capital initial letter in the
original publication, and being a substantive, not agreeing
with the generic name in gender, it probably would be class=
ed among those which the International Rules of Nomenclature
(more properly called "International Exceptions in Nomencla-
ture"$) in Recommendation 43 allows to be capitalized. It is
gratifying to the present writer to note how many modern
authors are disregarding this unfortunate recommendation.

Additional citations: PARAGUAY: Hassler 8120 [Macbride
photos 24,621] (F--photo of type, Kr--photo of type).

90. AEGIPHILA CEPHALOPHORA Standl.
Additional citations: PANAMA: Canal Zone: Kenoyer 607 (F
--fragment of type, F==photo of type).

98. AEGIPHILA CHRYSANTHA Hayek. ~

An additional synonym is Aegiphila chrysantha Poepp.e ex
Moldenke, Suppl. List Invalid Names 1, in syn. (1941). The
Klug 2104 and 2204 from Loreto, Peru, cited by me in Britt-
onia 1: 423 & 474 (1934) and Phytologia 1: 297 (1938) as A.
vitelliniflora Klotzsch are actually A. chrysantha and the
citations are repeated in their correct place hereinafter.
Klug 2204 has the disks very conspicuous along the midrib on
the lower leaf=surface and was identified as A. Smithii Mol-
denke by Standley. It has been collected in anthesis from
March to July and at altitudes of 180--200 m. The common
name "fetord-ey" is recorded by Klug and applies to this
species instead of to A. vitelliniflora as erroneously stat-
ed by me in Brittonia 1: 423 (1934) and Alph. List Common
Names 12 (1939).

Additional citations: ECUADOR: Guayas: Eggers 14,348
(Macbride photos 20,349] (F--photo, F--photo, Kr--photo).
PERU: Loreto: Klug 2027 (F), 2104 (A, B, E, F, G, K, Mi, N;
S, W), 2204 (B, Cb, E, F, G, K, S, W)3 Poeppig 2314 [Mac-
bride photos 34,313] (F--fragment of isotype, F--photo of
logotype).

98a. AEGIPHILA CHRYSANTHA var. GLABRA Moldenke, var. nov.

Haec verietas a forma typica speciei recedit calyce ubi-
que glabro.

This variety differs from the typical form of the species
in its calyx being entirely glabrous. Tne corolla is descri-~
bed by the collector as being cream-colored. Ths type was
collected by Guillermo Klug (no. 2894) at Juan Jui, Alto Rio
Huallaga, at an altitude of about me, San Martin, Peru,
in October, 1934, and is deposited in the Britton Herbarium
at the New York Botanical Garden. The collector state that
it is a liana, and Standley identified it as A. Smithii.

Citations: PERU: San Martin: Klug 3894 (F--isotype, N--

a ee ee ee ee ee oS ee

eee Tee

1941 Moldenke, Notes on Aegiphila 61

type).

44b. AEGIPHILA CONTURBATA Moldenke.
Additional citations: BRAZIL: Maranhao: Newman sen. [Mac=-
bride photos 28,377] (Kr--photo of type).

112. AEGIPHILA CORDATA Poepp.

An additional synonym, due to misaccrediting, is Aegiph-
ila cordata P. & E. ex Moldenke, Suppl. List Invalid Names
I, in syn. (1941).

Additional citations: PERU: Loreto: Poeppig 2158 [Mac-
bride photos 34,312] (F--photo of type, F--fragment of iso-
type, F--photo of type).

1l2a. AEGIPHILA CORDATA var. COLOMBIANA Moldenke.
Additional citations: COLOMBIA: Santander Sur: Haught
1885 (F--isotype).

113. AEGIPHILA CORDIFOLIA (Ruiz & Pav.) Moldenke.
Additional citations: PERU: Department undetermined:

Ruiz & Pavon 12/68 (F), sen. [Mifla, Panatehua] (Kr-~photo of

isotype).

10. AEGIPHILA COSTARICENSIS Moldenke.

Austin Smith has furnished us with copious field notes a-
bout this species. He states that it is an erect bushy shrub
4m. tall or a tree to & m. tall, the trunk 20 cm. in diame-
ter at breast-height, with a broad flat crown, inhabiting
the shade of Caribbean rain-forests, in mould and loamy soil
at altitudes of 600--1100 m. The bark is brown or pale=
brown, slightly roughened by raised striations. The leaves
are "thin-chartaceous" or thin-membranous, "faintly stiffen-
ed", dark opaque~green above, glabrous, with a faint sheen.
The "cupules" [calyx?] are green; “dried peduncles and cup-
ules brownish to black". The drupes are verious shades of
green, pale-green to greenish-yellow, ripening to Prussian-
green. It has been collected in anthesis in December and in
fruit in March and April and has been confused with the gen-
us Vitex.

Additional citatione: MEXICO: Chiapas: Matuda 572 (F--
fragment), 2101 (F). GUATEMALA: Quezaltenango: Skutch 2012
(F). COSTA RICA: Alajuela: A. Smith F.1816 (F, N), Fel907
(F, N). Cartago: Pittier & Toute & gen. [Herb. Instit. Physi-
co-geogr. Nat. Costaric. 9167] (F—-fragment of isotype).
Guanacaste: Standley & Valerio 45 22828 F==-photo ).

35. AEGIPHILA ORENATA Moldenke.
Additional citations: BRAZIL: Parand: Dusén 10,541 [Mac-
bride photos 30,182] (F--isotype, F=-photo of | Laotype, kr-=

62 PHYTOLOGIA Vole. 25 noe 2

photo of isotype), 16,238 (F).

17a. “eats CUATRECASASI Moldenke, Phytologia 2: 7-8.
1941.
Citations: COLOMBIA: Caquetdé: Cuatrecasas 8566 (N—frag-
ment of type, N--photo of type, W--type, W--isotype, Z=-—
photo of type).

9. AEGIPHILA CUNEATA Moldenke.

The species is said by Ule to be a shrub 2--9 m. tall,
With white flowers, blooming in July.

Additional citations: BRAZIL: Acre Territory: Ule 9859
("14,684") (F).

117. AEGIPHILA DEPPEANA Steud.

In Phytologia 1: 291 (1938) I cited a "Dugand & Mina 950"
from "Department undetermined", Colombia, and in Phytologia
1: 383 (1940) I stated that this collection was actually
made in the Panama Canal Zone. Dr. Armando Dugand, in a let=-
ter to me, dated March 18, 1941, has kindly pointed out to
me that I was in error in the above references. The collect-
ion was made by Dr. Armando Dugand G. at Juan Mina, a small
place some 10 miles southwest of Barranquilla, in the Dep-
artment of Atlantico, Colombia. The collector describes the
plant as having "small tubular reddish flowers" and records
the vernacular name "sauco monte". Elias calls it a rather
abundant woody vine around Barranquilla, with light bark
and no economic uses. Williams and Martinez-Calderdn des-
cribe it from Mexico as an uncommon straggly scandent shrub
on the edge of paths in thickets or a vine in llanos, found
at altitudes of 20--50 m., collected in fruit in March. It
has been confused with A. elata Sw. by some recent workers.
An additional synonym, due to misaccrediting, is Aegiphila
Deppeana Moldenke, Suppl. List Invalid Names 1, in syn.
(lou). In Brittonia 1: 452 (1934) I stated that the type of
A. Berteriana Schau. was collected by Bertero. This is an
error. It was collected by Balbis, as is shown by Macbride
photo 33,932, cited below.

Additional citations: MEXICO: Tamaulipas: Schiede 1165
(F--photo of type). Veracruz: Ll. Williams 8872 (F). Oaxaca:
Martinez-Calderdn 418 (W). State undetermined: Sessé, Moc-
iflo, Castillo, & Maldonado 6035 (F), 1074 (F). COSTA RICA:
Department undetermined: C. W. Dodge 6189 (F). COLOMBIA: At-
ldntico: Dugand G. 950 [Mius. Yale School of Forestry 32,382]
(F); Elias 1621 (F, N). Magdalena: Balbis s.n. [Macbride
photos 33,932] (F--photo); H. H. Smith 661 (Ca).

~ 101. AEGIPHILA ELONGATA Moldenke.
Additecit.: BOLIVIA: La Paz: Buchtien 1645 (F=ph.of type)

7
7
j
7
;

1941 Moldenke, Notes on Aegiphila 63

124. AEGIPHILA ELATA Sw.

Williams describes the species as a "slender shrub, at
times scandent, in secondary growth", a "vine on shrubs in
clearings or rough pastures", and as a "scandent herb". As a
shrub it is said to attain a height of 4 to 5 feet. Williams -
reports the flowers as sometimes white and the common name
“bejuco de peine mico" in Oaxaca. It has been collected in
fruit in February and has been confused with the genus
Cestrum of the Solanaceae. .

Additional citations: MEXICO: Oaxaca: Ll. Williams 9143
(F), 9271 (F), 9281 (F), 9354 (F), 9566 (F). Tabasco: Matuda

031 oe 081 (F), 3406 (F). BRITISH HONDURAS: Gentle 2653
F, Mi), 2684 (Mi), 2843 (F, Mi, Mi), 3047 (F, Mi), 3350
(N). HONDURAS: Atldntida: Yuncker, Koepper, & Wagner 8
(F). COSTA RICA: Alajuela: Brenes 20,535 ["30"] (F). COLOMB-
IA: Cundinamarca: Triana 3135 Us 77"] (Jc). Tolima: Pérez
Arbeldez & Quatrecasas 6529 (W). VENEZUELA: Bolivar: Ll.
Williams 12,828 (Ve, W).

121. AEGIPHILA ELEGANS Moldenke.

Krukoff describes the species as a vine, growing on terra
firma in high forests.

Additional citations: BRAZIL: Amazonas: Krukoff 8701 (F).

61. AEGIPHILA FALCATA Donn. Sm.

Standley reports that the species is a shrub 2-—-4 m.
tall, inhabiting dry or damp thickets, at an altitude of ab=
out 240 m., that the flowers are greenish-yellow or pale
greenish-yellow, and that a vernacular name in Guatemala is
"chiploque"; collected in flower and fruit in September.

Additional citations: MEXICO: Chiapas: Matuda 666 (Mi,
Mi). GUATEMALA: Escuintla: J. D. Smith 2111 (F==photo of
type). Retalhuleu: J. D. Smith 1479 (F--photo); P. C. Stand-
ley 88,699 (N), 88,767 (N). COSTA RICA: Department undeterm-
ined: Calvert & Calvert son. [Surubres, Oct. 1909] (Up).

15. AEGIPHILA FASCICULATA Donn. Sm.

An additional synonym, due to mis-accrediting, is Aegiph-
ila fasciculata H.BeK. ex Moldenke, Suppl. List Invalid
Names 1, in syn. 1941.

Additional citations: GUATEMALA: Alta Verapaz: Turckheim
4013 (F--photo of type). .

114. AEGIPHILA FENDLERI Moldenke.

The species grows at altitides of 1450--1650 m., and has
been collected in anthesis in Octobe.

Additional citations: VENEZUELA: Aragua: Chardon 189 (N--
fragment, Ve, W).

64 PHYTOLOGIA Vol. 2, no.2

31. AEGIPHILA FERRUGINBA Hayek & Spruce.

Rimbach has furnished some additional information about
this species. He states that the cortex of the trunk is gray
and slightly fissured, soft in texture, and the wood is
white, with distinct growth-rings; also that it is a medium
sized tree of the forest. An additional synonym, due to mis-
accrediting, is Aegiphila ferruginea Hayek ex Moldenke,
Suppl. List Invalid Names 1, in syn. 1941.

Additional rinks ECUADOR: Chimborazo: Rimbach 616
(F). Imbabura: Lehmann 4700 [Macbride Bagot 17,504] (F--

photo, Kr--photo). Pichincha: Firmin 632 (F--photo); Penland

& Summers 939 (F); Spruce 5473 (F—fragment of isotype, F--
photo of isotype). |

63. AEGIPHILA FILIPES Mart. & Schau.

The species is described by collectors as a tall shrub or
small tree, 10—-18 feet tall, with a stem 1/2 to 1 inch in
diameter, sometimes to 22 feet tall, inhabiting pastures,
forests, and jungles on varzea land, the flowers white or
yellowish. The calyx is green, remarkably and quite charact-
eristically wide during anthesis. The fruit is yellow or or=-
ange. It has been collected in anthesis in March and from ~
July to September, and in fruit in August. It ascends to
1900 me. altitude in Colombia.

Additional te COLOMBLA: Huila: Pérez Arbeldez &
Cuatrecasas 8360 (W). PERU: Loreto: Ll. Williams 5335 (F),
2165 (F), 2469 (F), (F), 2778 (F), 28352 (F), 2850 (F),

2856 (F), 3115 (F), 3146 (F). BRAZIL: Amazonas: Krukoff 5125
coon Dept. Bot. Est. 8S. Paulo 35,047] (Ca, F, Mi, Sp),
8041 (F), 8042 (F); Riedel 1418 (F). Pard: Martius s.n.
[Herb. Mohac. 1020 & 1689; Macbride photos 20,550 50] (F=-photo
of cotype, Kr--photo of cotype).

66..AEGIPHILA FLORIBUNDA Moritz & Moldenke.
The label. on the photogrpah cited below says "Moritz &
Moldenke 1765" for the collection number, in errore
Additional citations: VENEZUELA: Aragua: Moritz 1765
[Macbride photos 34,310] PN

57. AEGIPHILA FOETIDA Sw. ;
Additional citations: JAMAICA: Re C. Alexander sen. (F--
photo); Swartz sen. [Jamaica] (F--photo of type). 7

62a. AERGIPHILA GLANDULIFERA var. PARAENSIS Moldenke.

The variety is described as a shrun 12 feet tall, bloom
ing in August; the fruit red when ripe. It has been confused
With A. filipes.

Additional citations: BRAZIL: Pard: Ginzberger 908 (F);
Krukoff 59235 (F, Mi).

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ae LIBRARY

Beis Baye: NEW YORK
Sa - BOTANICA!
hose PHYTOLOGIA GARDEA
dys 3 aa ae
ey: Designed to expedite botanical publication
ahaa December, 1944 _ No.3
CONTENTS
ig N. MoLDENKE: Additional common and vernacular
names recorded for members of the Verbenaceae
GM AVIGENNIACEGE, \ncdiecsc ins -scvsdessbactad Ackccebsgeceveigcincsdccenske 65

LN. MoLpDENKE: The recorded common and vernacular
names of Verbenaceae and Avicenniaceae ar-
ranged according to genera And SPeCIES..eescccccceseeeseeeeeseeeee 89

— laceae, Avicenniaceae, and Verbenaceae of
_ Texas. Ti: eeeeeeseserveee COPS eee ee eeteeeeeeree THOS eee eseeeseeeee the tes

ne Published by H. A. Gleason and Harold N. Moldenke
ae The New York Botanical Garden
ate Bronx Park, New York 58, N. Y.

Price of this number, 75 cents; per volume, $5.00 in advance

Vol. 2, No. 2 was issued December 8, 1941

AW EY loo

=a

=

SADNAK Y

NEW YORI

BOTANICA
GARDE

ADDITIOCNAL COMMON AND VERNACULAR NAMES RECORDED FOR MEMBERS
| OF THE VERBENACEAE AND AVICENNIACEAE

Harold N. Moldenke

Since the publication of my original alphabetic list of
2202 common and vernacular appellations for members of the
Verbenacese and Avicenniaceae (1) and the supplementary list
of 1387 additional names (2) numerous names have come to my
attention on the labels of herbarium specimens and in vari-
ous manuals, horticultural catalogues, floral lists, and de-
scriptive accounts of regions in which these plants grow.
These 1012 additional names are given herewith together with
some corrections and emendations of previous listings. As in
the previous works, all variations in orthography or accent-
uation are listed separately in exactly the manner as given.
by the original recorder. Entries which are merely correc
tions or emendations of entries in previous installments of
this list are here indicated by an asterisk (*).

abisoa = Vitex Doniana Sweet, V. grandifolia Gtirke
abontennua = Stachytarpheta jamaicensis (L.) Vahl
ada = Vitex Doniana Sweet, V. grandifolia Gtirke
adabi = Clerodendrum splendens G. Don .
adaga = Vitex Doniana Sweet, V. grandifolia Giirke
ade = Vitex Doniana Sweet
_adefia = Vitex grandifolia Gtirke
adelamanyi = Lantana Camara L.
*adgdu = Premna corymbosa var. obtusifolia (R. Br.) Fletcher
aegiphilas = Aegiphila Jacq.
afetewa = Vitex Doniana Sweet, V. grandifolia Gtirke
afia-nunung = Avicennia africana P. Beauv.
afifia omya = Clerodendrum splendens G. Don
afurati = Lippia adoénsis Hochst.
*agdu = Premna corymbosa var. obtusifolia (R. Br.) Fletcher
agba = Stachytarpheta jamaicensis (L.) Vahl
agbul u uwagh = Clerodendrum capitatum (Willd.) Schum. &
Thonn.
*apdau = Prema corymbosa var. obtusifolia (R- Br.) Fletcher
*agetha = Premna corymbosa (Burm. f.) Rottl. & Willd.
*aonimantha = Premna corymbosa (Burm. f.) Rottl. & Willd.
agnocasto = Vitex Agnus-castus L.
&gdgo igin = Stachytarpheta jamaicensis (L.) Vahl
agug = Teijsmanniodendron Ahernianum (Merr.) Bakh.
*aguydb&t = Premna corymbosa (Burm. f.) Rottl. & Willd.
ahgao = Premna Gaudichaudii Schau.
*aker ketu-kétu = Sphenodesme pentandre Jack
5

66 PHY TOL ‘O'Git<A Vol. 2, now 3

*“akear lintang ruas = Sphenodesme pentandra Jack :

akojoe méla kali = Stachytarpheta cayennensis (L. C. Rich.)
Vahl

akotongmé = Lantana Camara L.

akwakora-gyahini = Vitex Fosteri C. H. Wright, V. rivularis
Gtirke a:

*alagdu = Premna corymbosa var. obtusifolia (R. Br.)Fleteher

*alagéu-blanko = Premna corymbosa var. obtusifolia (R. Br.)

Fletcher

*alagdu-ddgat = Premna corymbosa var. obtusifolia (R. Br.)
Fletcher

*alalgéu = Premnma corymbosa var. obtusifolia (R. Br.) Flet-
cher

alasaobo =.Vitex Stahelii Moldenke

Albion Verbena = Verbena Teasii Moldenke

albocar = Callicarpa acuminata H.B.K.

alfornbrilla = Verbena ciliata Benth.

algarrobo = Avicennia nitida Jacq.

aloalo = Prema taitensis var. rimatarensis F. H. Br., Pe
corymbosa (Burm. f.) Rottl. & Willd.

aloch = Vitex Agnus-castus L.

ama-kosikati = Vitex Wilmsii var. reflexa (H. H. W. Pearson)
Pieper :

*amamah{t = Teijsmanniodendron Ahernianum (Merr.) Bakh.

*amambol{gan = Clerodendrum minahassae Teijsm. & Binn.

*ambuligan = Clerodendrum minahassae Teijsm. & Binn.

American callicarpa = Callicarpa americana L.

amor de hombre = Verbena temiisecta Briq.

amér de hombre = Verbena tenuisecta Brig.

amu-ati = Avicennia africana P. Beauv.

amu-tsi = Avicennia africana P. Beauv.

ananse ddkono = Lantana Camara L.

ananse dua = Lantana Camara L.

ananse kono = Lantana Mearnsii Moldenke

ananu kOmi = Lantana Camara L., L. Mearnsii Moldenke

ananu kén-tsho = Lantana Camara .., L- Mearnsii Moldenke

*andarése = Premna corymbosa (Burm. f.) Rottl. & Willd.

andofiti = Vitex micrantha Giirke

angalem = Vitex Doniana Sweet

éngel = Aloysia ligustrina (Lag.) Small

angkasa~angkasa = Stachytarpheta jamaicensis (L.) Vahl
angma-tsho = Avicennia africana P. Beauv.

RET nena tee neces

ankasa = Stachytarpheta jamaicensis (L.) Vahl
*anobrang = Premna corymbosa (Burm. f.) Rottl. & Willd.

antelope's garden egg = Vitex rivularis Gtirke

*aoepaloelan hahoela = Clerodendrum Rumphianum De Vriese

api-api = Avicennia alba Blume, *A. marina var. Rumphiana
(H. Hallier) Bakh.

api-api putik = Avicennia marina (Forsk.) Vierh.

;

1944 Moldenke, Common Names 67

apokotja = Vitex compressa Turcz.

apord = Clerodendrum polycephalum J. G. Baker

*appel = Premna corymbosa (Burm. f.) Rottl. & Willd.
*eragéu = Premna corymbosa (Burm. f.) Rottl. & Willd.
aranga = Vitex Doniana Sweet, V. grandifolia Gtirke

*arbre & la migraine = Premna corymbosa (Burm. f.) Rottl. &

Willd.
*arbre de la migraine = Premna corymbosa (Burm. f.) Rottl. &
Willd.

*argéu = Premna corymbosa (Burm. f.) Rottl. & Willd.
arisgo = Duranta repens L.

ariya = Clerodendrum inerme (L.) Gaertn.

*arni = Premna corymbosa (Burm. f.) Rottl. & Willd.
*arnrai = Vitex quinata (Lour.) F. N. Will.

asokoro = Avicennia africana P. Beauv.

asokpolo = Avicennia africana P. Beauv.

asopro = Avicennia africana P. Beauv.

asukuru = Avicennia africana P. Beauv.

ata-nunung = Avicennia africana P. Beauv.

atiaci = Lippia Hoehmei Moldenke

*ayam~ayam = Clerodendrum minahassae phigh & Binn. ©
ayeti = Clerodendrum capitatum (Willd.) Schum. & Thonn.

*babon = Premna corymbosa (Burm. f.) Rottl. & Willd.

bacatén = Lippia Pringlei Brigq.

badi = Clerodendrum barba-felis H. Hallier

baéh zitang = Clerodendrum paniculatum L.

*bagalbak = Clerodendrum minahassae Teijsem. & Binn.
*bagduak = Clerodendrum minahassae Teijem. & Binn.
*bagduak-itim = Clerodendrum minahassae Teijsm. & Binn.
*bagduak-ne-puti = Clerodendrum minahassae Teijsm. & Binn.
Neahema tea =-Lantana Camara L.

bahé = Lippia adoénsis Hochst.

bahé-bahé = Lippia adoénsis Hochst. |

*bakarcha = Premna corymbosa (Burm. f.) Rottl. & Willd.
*bakébak = Clerodendrum minahassae Teijsm. & Binn.
bakoréné = Clerodendrum Buchholzii Gtirke

ba-kudu-ne = Vitex barbata Planch., V. chrysocarpa Planch.
*balabi = Prenna corymbosa (Burm. f.) Rottl. & Willd.
balamagnian kan = Vitex chrysocarpa Planch.

balsamo = Citharexylum fruticosum L.

balunakuta = Stachytarpheta mutabilis (Jacq.) Vahl
*bangana = Gmelina elliptica J. ©. Sm.

bartanucha = Verbena pumila Rydb.

*basal = Vitex quinata (Lour.) F. N. Will.

bastard vervain = Stachytarpheta jamaicensis (L.) Vahl
bataydqui = Lippia Pringlei Brigq.

beauty-berry = Callicarpa americana L-

*bSbuas = Premna corymbosa (Burm. f.) Rottl. & Willd.
*bSbuat = Prema corymbosa (Burm. f.) Rottl. & Willd.

-

68 PHYTOLOGIA Vol. 2, no. 3

bee blossom = Aloysia ligustrina var. Schulzii (Standl.)
Moldenke

bee brush = Aloysia ligustrina (Lag.) Small

*beech = Gmelina Leichhardtii (F. Mueli.) F. Muell.

bejuco de peine mico = Aegiphila elata Sw.

Bellaire Verbena = Verbena Teasii Moldenke

*bélongeh = Gmelina elliptica J. 3. Sm.

benturosa morada = Lantana trifolia L.

bercul = Verbena menthaefolia Benth.

Bermudian mulberry = Callicarpa americana L.

*bhuijam = Pygmaeoprema herbacea (Roxb.) Moldenke

*bhumijambu = Pygmaeopremna herbacea (Roxb.) Moldenke

*bhumi-jambiika = Pygmaeopremna herbacea (Roxb.) Moldenke

*bhit-bhirari = Prema corymbosa (Burm. f.) Rottl. & Willd.

bichicho = Verbena crithmifolia Gill. & Hooks

bilankuru fida = Premna hispida Benth.

blacktree.= Avicennia nitida Jacq.

blackwood = Avicennia nitida Jacq.

bleeding heart = Clerodendrum Thomsonae Balf. f.

blue plumeria = Duranta repens L.

blue rats tail = Stachytarpheta urticaefolia (Salisb.) Sims

Blue Sentinel Verbena = Verbena hybrida Voss

blue spirea = Caryopteris incana (Thunb.) Miq.

blue verbena = Verbena hastata L., V. stricta Vent.

blue vervain = Verbena Blanchardi Moldenke, V. Engelmannii
Moldenke, V. Halei Small, V. stricta Vent.

boandjo = Avicennia africana P. Beauv.

*Bocksblatt = Prema corymbosa (Burm. f.) Rottl. & Willd.

*boelangan = Gmelina asiatica var. villosa Bakh.

*boenato = Clerodendrum minahassae Teijsm. & Binn.

boenga-in tah = Lantana Camara var. aculeata (L.) Moldenke

*boenga panggil = Clerodendrum Rumphianum De Vriese

*boenga pluim = Clerodendrum Rumphianum De Vriese

*boenga poean = Clerodendrum Rumphianum De Vriese

*boengis = Vitex quinata (Lour.) F. N. Will.

boerta-boerta = Clerodendrum adenophysum H. Hallier

boesie droifi = Clerodendrum aculeatum (L.) Schlecht.

*boewah kerandjang = Gmelina asiatica var. villosa Bakh.

bofuluk = Vitex grandifolia Girke

*bohol = Gmelina elliptica J. @. Sm.

*bois & cotelettes = Citharexylum B. Juss.

*bois cotelet = Citharexylum B. Juss.

*bois de bouc = Premna corymbosa (Burm. f.) Rottl. & Willd.

*bois de guitare = Citharexylum B. Juss.

*bois sureau sauvage = Premna corymbosa (Burm. f.) Rottl. &
-Willd.

*bokkeblad = Premna corymbosa (Burm. f.) Rottl. & Willd.

bok wat tan = Callicarpa longissima (Hemsl.) Merr.

bollo limpio = Aegiphila puberulenta Moldenke

1944 Moldenke, Comnon Names 69

*bongogon = Viticipremna philippinensis (Turez.) H. J. Lam
borom-borom = Lippia adoénsis Hochst.

boschkalebas = Vitex compressa Turcz.

bracted vervain = Verbena bracteata Lag. & Rodr.
Brazilian lantana = Lantana fucata Lindl.

Brazilian tea = Stachytarpheta jamaicensis (L.) Vahl
broedae nahatti = Clerodendrum Thomsonae Balf. f.
bué = Avicennia africana P. Beauv.

bué-dinté = Avicennia africana P. Beauv.

buji = Vitex simplicifolia Oliv.

*bulang = Gmelina ellivtica J. %. Sm.

*bulang gajah = Gmelina elliptica J. EH. Sm.

*bulang hutan = Gmelina elliptica J. &. Sm.

*bulang k&chil = Gmelina elliptica J. %. Sm.
*bulbuol = Gmelina elliptica J. %. Sm.

bummehi = Vitex simplicifolia Oliv.

bummeji = Vitex simplicifolia Oliv.
bummere = Vitex simplicifolia Oliv.

*bunalun-babay = Avicennia marina var. Rumphiana (H. Halli-
er) Bakh.

bunch-berry = Lantana horrida H.B.K.

*bunga k¥rtas = Sphenodesme pentandra Jack

*bunglas = Tectona philippinensis Benth.

Burnett's Scarlet Verbena = Verbena hybrida Voss

burzun = Vitex Doniana Sweet

buttonweed = Phyla nodiflora var. reptans (H.B.K.) Moldenke

buwe = Avicennia africana P. Beauv.

cabaraé-caa& = Lantana Camara L., L. montevidensis (Spreng.)
Briq.

cabradora simarona = Aloysia macrostachya (Torr.) Moldenke

café cimarrén = Aeginhila monstrosa Moldenke

calico bush = Lantana horrida H.B.K.

camaré = Lantana montevidensis (Spreng.) Brig.

camaradinha = Verbena phlogiflora Cham.

camara faux thé = Lippia Psevdo-thea (A. St. Hil.) Schau.

camara roseo = Lantana fucata Lindl.

cambaré = Lantana Chamissonis (D. Dietr.) Benth., L. tiliae-
folia Cham.

canahuite = Githsarexylum hexangulare Creenm.

capa-blanca = Petitia domingensis Jacq.

capa rosa = Callicarpa ampla Schau.

- capa savannah = Petitia domingensis Jacq.

a

capitao do mato = Lipviea Pseudo-thea (A. St. Hil.) Schau.
*caragra = Liopia oxyphyllaria (Donn. Sm.) Standl.
cariaquito = Lantana “oritziana Otto & Dietr.

cariaquito blanco = Lantana achyranthifolia Desf.
carrioquito = Lantana Camara L.

Carter's Dwarf Coerulea Verbena = Verbena hybrida Voss
Carter's Dwarf Compact Verbena = Verbena hybrida Voss

70 PA POM OG Lak Vol. 2, now 5

Carter's Holborn.Mammaoth Verbena = Verbena hybrida Voss

carvoeiro = Citharexylum myrianthum Cham.

cateicillo = Citharexylum caudatum L.

cawuira = Aegiphila racemosa Vell.

cedron = Aloysia ligustrina (Lag.) Small, A. triphylla
(iter) Britton, A. virgata (Rufz & Pav.) A. L. Juss.

Ceres Verbena = Verbena Teasii Moldenke

chaek tsai shue = Callicarpa rubella Lindl.

cha de pedreste = Lippia Pseudo-thea (A. St. Hil.) Schau.

*chah leud = Premna corymbosa (Burm. f.) Rottl. & Willd.

*chéméri = Premna corymbosa (Burm. f.) Rottl. & Willd.

chaste-trees = Vitex Tourn.

chau pin tung = Clerodendrum fragrans var. plenitlorum
Schau.

chau shi mut li = Clerodendrum Bungei Steud.

chéne calebassic = Petitia domingensis Jacq.

chicharra caopi = Aloysia virgata (Ruiz & Pav.) A. Le Juss.

chile pdjaro = Citharexylum brechyanthum (A. Gray) A. Gray

chiligua = Lippia cardiostegia Benth.

chilillo = Stachytarpheta angustifolia (“ill.) Vahl

Chinese beardwort = Caryopteris incana (Thunb.) Miq.

Chinese-hats = Holmskioldia sanguinea Retz.

chingari = Clerodendrum indicum (L.) kuntze

chinkuro = Lantana hispida H.B.K.

chinquillo = Neosparton ephedroides Griseb.

chiploque = Aegiphila falcata Donn. Sm.

chisnan = Duranta triacantha A. L. Juss.

chuul = Citharexylum Donnell-Smithii Greenm.

cidrera = Lippia alba (Mill.) N. %. Br.

cinzeiro = Aegiphila Sellowiana Cham.

*clérodendron = Clerodendrum Burm.

comasi = Stachytarpheta urticaefolia (Salisb.) Sims

commode mulberry = Callicarpa americana L.

common deep orange lantana = Lantana Camara L.

common lantana = Lantana Camara var. aculeata (L.) Moldenke,
L. Camara var. mista (L.) Le H. Bailey

common lilac lantana = Lantane tiliaefolia Cham.

common verbena = Verbena bipinnatifida Nutt., V. hybrida

Voss
common vervain = Verbena Abramsi Moldenke, V. lasiostachys
Link

comnon white vervain = Verbena urticifolia L.

confite = Lantana velutina Mart. & Gal.

confite: blanca = Lantana velutina Mart. & Gal.
confituria amarilla = Lantana glandulosissima Hayek
confiturilla = Lantana horrida H.B.K.

confiturilla amarilla = Lantana glandulosissima Hayek
confiturilla blanca = Lantana velutina Mart. & Gal.
coralillo rosado = Duranta repens L.

1944 Moldenke, Common Names 71

corocillo = Stachytarpheta cayennensis (L. 0. Rich.) Vahl
*cotelet = Citherexylum B. Juss. ~

*cotelets = Citharexylum B. Juss.

*cotelet tomenteux = Citharexylum Kunthianum Moldenke

cuul = Citharexylum Donnell-Smithii Greenm.

*dabtan = Vitex trifolia var. bicolor (‘lilld.) Moldenke

dadiangas = Gmelina elliptica J. #. Sm.

dagba = Clerodendrum volubile P. Beauv.

Dakota verbena = Verbena bipinnatifida Nutt.

*delipapa = Teijsmanniodendron Ahernianum (Merr.) Bakh.

dame cubre galanos = Lantana Camara L.

*danasi = Geunsia Cumingiana (Schau.) Rolfe

*denata = Clerodendrum minahassae Teijsm. & Binn.

dancundi = Vitex trifolia var. simplicifolia Cham.

*dangl& = Vitex trifolia var. bicolor (Willd.) Moldenke

*dafigila = Teijesmanniodendron Ahernianum ('err.) Bakh.

*danhanges = Gmelina elliptica J. H. Sm.

danna = Citharexylum macradenium Greenm.

*daoen kambina = Premna corymbosa (Burm. f.) Rottl. & Willd.

¥dauhon lagondie = Vitex trifolia var. bicolor (Willd.) Mol-
denke \

d'dap mira = Hosea Lobbii (C. B. Clarke) Ridl.

dengd = Prema quadrifolia Schum. & Thonn.

*der surinamsche Thé = Lantana Camara L-

devil's coach whip = Stachytarpheta jamaicensis (L.) Vahl

*didigkalin = Teijsmanniodendron Ahernianum (Merr.) Bakh.

*didip&pak = Teijsmanniodendron Ahernianum (Merr.) Bakh.

dinchi = Vitex Doniana Sweet

'dinya = Vitex Doniana Sweet

'dinyar biri = Vitex simplicifolia Oliv.

diohuli = Lippia adoénsis Hochst.

djin-akwa = Vitex micrantha Giurke

drap d'or = Lantana urticaefolia Mill.

droceria = Lippia umbellata Cav.

'dumniya = Vitex Doniana Sweet

*dufigila = Teijsmanniodendron Ahernianum (‘err.) Bakh.

‘dunya = Vitex Doniana Sweet

'dunyar biri = Vitex simplicifolia Oliv.

durancia = Duranta repens L.

duranta = Duranta repens L.

duranta de Plumier = Duranta repens L.

Dwarf Coerulea Verbena = Verbena hybrida Voss

Dwarf Compact Verbena = Verbena hybrida Voss

dwarf lantana = Lantana Camara var. hybrida (Neubert) Mol-
denke

dyob = Vitex Doniana Sweet

*ear-stud climber = Sphenodesme pentandra Jack

Sbenote = Clerodendrum volubile P. Beauv.

Sbisaa = Vitex Doniana Sweet, V. grandifolia Gurke

72 PHY @' OL. 0G Ls Vol. 2, now 5

8-bure = Avicennia africana P. Beauv.

ede = Avicennia africana P. Beauv.

edin = Vitex Doniana Sweet

efinrin-gogara = Lippia adoénsis Hochst.
egwa = Clerodendrum Thomsonae Balf. f.
ehrodo = Avicennia africana P. Beauv.

&ji = Vitex Doniana Sweet

ek@nyieya = Clerodendrum splendens G. Don
ele-ele = Vitex Doniana Sweet

eleku = Lantana Mearnsii Moldenke

Blis's duranta = Duranta repens L.
English sage bush = Lantana Camara var. mista (L.) Le H.

Bailey
espina de pescado = Junellia s seriphioides (Gill. & Hook. Es
Moldenke

espino = Clerodendrum Pittieri Moldenke

Btn adé1é.= Lantana Camara L.

dwin agogo = Lantana Camara L-

fafa-hinei = Clerodendrum Buchholzii Gtirke

fafe = Clerodendrum Buchholzii Gtirke

false vervain = Verbena Blanchardi Moldenke

fasau = Lippia adoénsis Hochst.

*faux thé = Lippia Pseudo-thea (A. St. Hil.) Schau.

ferem&mi = Clerodendrum capitatum (Willd.) Schum. & Thonn.

fetfetti = Lippia adoénsis Hochst.

fetoréd-ey = Aegiphila chrysantha Hayek

feve = Vitex micrantha Gtirke

fevei = Vitex micrantha Glrke, V. Ripe ee J. G. Baker

fiddlewood = | Citharexyl un B. Juss.

*fiddle-wood = Citharexylun B. Juss.

*fiddlewood tree = Citharexylum B. Juss.

*fidelle-wood = Citharexylum B. Juss.

filigrana = Lantana Camara L., L. montevidensis (Spreng.)
Briq.

fiolintraee = Citharexylum B. Juss.

fioltraed = Citharexylum B. Juss.

firi-fore = Clerodendrum capitatum (Willd.) Schum. & Thonn.,
C. umbellatum Poir.

flor de chichalaque = Callicarpa acuminata H.B.K.

flor de la rosa merte = Clerodendrum Bungei Steud. .

flor de sangre = Lantana Moritziana Otto & Dietr.

flowering verbena = Verbena canadensis (L.-) Britton

£5 = Vitex Doniana Sweet, V. ~grandifolia Gtirke

fog fruit = Phyla | lanceolata (Michx.) Greene, P. nodiflora

(L.) Greene
fog-fruit = Phyla incisa Small
for chai tsai = Oallicarpa formosana Rolfe
forget-me-not = Duranta repens var. alba (Masters) L. H.
Bailey

4
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z
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1944 Moldenke, Common Names 73

Fordhook Famous Verbena = Verbena hybrida Voss |
f5-ti = Vitex Doniana Sweet, V. grandifolia Gtirke
fox-fruit = PI Phyla lanceolata (fichx.) Greene
£5 yi = Vitex Doniana Sweet, V. grandifolia Gtirke
£8 yi-ti = Vitex Doniana Sweet, V. grandifolia Gtirke
f%yi-tsho = Vitex Domiana Sweet, V. prandifolia Glirke
fragrant clerodendron = Clerodendrum Bungei Steud.
French mulberry = Callicarpa americana var. lactea F. J.
Muller
frog fruit = Phyla incisa Small, P. nodiflora (L. ) Greene
frog-fruit = Phyla Lour.
fruta de iguana = Duranta repens L.
fruta de macaco Oitharexylum myrianthum Cham.
fruta de paloma = Duranta repens var. canescens Moldenke
frutilla = Lantana scorta Moldenke
frutilla blanca = Lantana achyranthifolia Desf.
frutillo = Lantana achyranthifolia Desf.
fuemémi = Clerodendrum capitatum (Willd.) Schum. & Thonn.
fumaria = Verbena tenuisecta Briq.
furu-fure = Clerodendrum capitatum (Willd.) Schum. & Thonn.,
C. umbellatum Poir.
*papayug = Geunsia Cumingiana (Schau.) Rolfe
galbihi = Vitex Dor Doniana Sweet
galbije = Vitex Doniana Sweet
*pealipépa = = Teijsmanniodendron Ahernianum (Merr.) Bakh.
EPDS ba = Lippia adoénsis Hochst.
*ganidri = Prema corymbosa (Burm. f.) Rottl. & Willd.
hated “= Premna corymbosa (Burm. f.) Rottl. & Willd.
*canniari = Premna corymbosa (Burm. f.) Rottl. & Willd.
garigari = Avicennia africana P. Beauv.
gb%1léti = Avicennia africana P. Beauv.
geakoi = Clerodendrum m splendens Ge Don
*Geigenholz = Citharexylum B. Juss.
*Geigenholzbaum = Citharexylum B. Juss.
gerbéo = Stachytarpheta cayennensis (L. C. Rich.) Vahl
gervao = Stachytarpheta australis Moldenke, S. Maximiliani
Schau., S. polyura Schau.
*cervad = Stachytarpheta cayennensis (L. C. Rich.) Vahl
*chebu-nelli = Premna corymbosa (Burm. f.) Rottl. & Willd.
Giant Pink Verbena = Verbena hybrida Voss
gidjiko = Vitex Doniana Sweet
gigatraed = Citharexylum B. Juss.
gigetraee = Citharexylum B. Juss.
*pineri = Premna corymbosa (Burm. f.) Rottl. & Willd.
globito = Priva lappulacea (L.) Pers.
*slorybower = Clerodendrum Burm.
*clory tree = Clerodendrum Burm.
godon kada = Phyla nodiflora (L.) Greene
God's coconut = Vitex prandifolia Gurke

Wo OP EE TOROE TA Vole 2, no. 3

*oemira = Premna corymbosa (Burm. f.) Rottl. & Willd.
_ *gofasa = Vitex quinata (Lour.) F. N. Will.
golden-dewdrop = Duranta Le -
golden dewdrops = Duranta repens L.
goo yis hai = Clerodendrum inerme (L.) Gaertn.
grigri = Avicennia africana P. Beauv.
guearataro = Vitex capitata Vahl
guilel guéri = Lippia adoénsis Hochst.
*cuitar wood = Citharexylum B. Juss. _
gulinda = Clerodendrum inerme (L.) Gaertn-
gumhar = Gmelina arborea Roxb. .
gyengya aforowa = Premna quadrifolia Schum. & Thonn.
hai ngan = Callicarpa cana L. |
hairy lantana = Lantana Camara var. mista (L.) L. H. Bailey
hairy verbena = Verbena pumila Rydb. 3
*hamurduon-asu = Viticipremna philippinensis (Turez.) H. J.
Lam
harlequin.glorybower = Clerodendrum trichotomum Thunb.
harlequin lantana = Lantana Camara var. varia (Kuntze) “ol-
denke
hayariballi = Petrea bracteata Steud. —
a

*headache tree = Premna corymbosa (Burm. f.) Rottl. & Willd.

hemptree = Vitex Tourn.

herba Luisa = Aloysia triphylla (L'Hér.) Britton

herb o' grace = Verbena officinalis L.

herimena-kola = Phyla nodiflora (L.) Greene

herva cidreira = Aloysia triphylla (L'Hér.) Britton

herva de picapdéo = Amasonia campestris (Aubl.) Moldenke

heul = Vitex Doniana Sweet ;

hierba de Christo = Lantana horrida H.B.K.

hierba de hormiga = Phyla nodiflora var. canescens (H.B.K.)
Moldenke, P, nodiflora var. reptans (H.B.K.) Moldenke

*hierba del incordio = Verbena tenuisecta Briq.

hierba del negro = Lippia alba (Mill.) N. 3. Br. ;

hierba dulce = Lippie praveolens H.BeK.

hierba negra = Lippia alba (Mill.) N. E. Br.

hoar vervain = Verbena stricta Vent.

hoary verbena = Verbena moechine Moldenke, V. stricta Vent.

hoary vervain = Verbena stricta f. albiflora Wadmond

hoi = Verbena bonariensis L.

Holborn Mammoth Verbena = Verbena hybrida Voss

honawai = Clerodendrum umbellatum Poir.

honey~mangrove = Avicennia nitida Jacq.

huhwwali = Vitex mollis H.B.K.

*hukre-mara = Clerodendrum viscosum var. nilagiricum H.

Hallier
huniyan = Pygmaeoprema humilis Merr., *B. herbacea (Roxb)
Moldenke

hunter-does-not=eat-it = Lantana Camara L.

rs Pe ee aD

1944 Moldenke, Common Names 75

hunters' scent = Lantana Mearnsii Moldenke
hunters' spice. = Lantana Mearnsii Moldenke
hwana wulie = Clerodendrum umbellatum Poir.
ibang = Vitex Fosteri CO. H. Wright
idj8li = Vitex simplicifolia Oliv.
*ipang = Teijsmanniodendron Ahernianunm (Merr.) Bakh.
iguanero = Avicennia nitida Jacq.
ildn-ildn = Aloysia Looseri Moldenke
ilang-ilang = Aloysia Looseri Moldenke
illiri = Clerodendrum capitatum (Willd.) Schum. & Thonn.
*incdic = Vitex rapinoides Guillaum.
*indjaro = Premna corymbosa (Burm. f.) Rottl. & Willd.
ingari = Vitex Doniana Sweet, V. grandifolia Gtirke
ink tree = Vitex Doniana Sweet, V. grandifolia Gtirke
*inrelo = Prema corymbosa (Burm. f.) Rottl. & Willd.
insuo-koto = Vitex chrysocarpa Planch.
irk aldngba = Stachytarpheta jamaicensis (L.) Vahl
irt amure = Stachytarpheta jamaicensis (L.) Vahl
ishé-din = Clerodendrum violaceum Gitirke
_ Italian verbena = Verbena tenera var. Maonetti Regel
-iye = Clerodendrum capitatum (Willd.) Schum. & Thonn.
jaia-guli = Avicennia africana P. Beauv.
_jampang laki = Vitex flava Ridl.
jaqueca = Verbena ephedroides Cham.
jaua verbena = Stachytarpheta cayennensis (Lo. heen)
Vahl ;
*ka-aungeyl = Clerodendrum infortunatum L.
ka-bure = Avicennia africana P. Beauv.
kadamanakku = Vitex altissima L. f.
*kada met = Pygmaeopremna herbacea (Roxb.) Moldenke
kaddunochchi = Vitex leucoxylon L. f.
kaf8i = Prema hispida Benth.
kafi = Premna hispida Benth.
kaikoa = Premna corymbosa var. sambucina (Wall.) Moldenke
kajie = Clerodendrum umbellatum Poir.
kajoe boerta-boerta = Clerodendrum adenophysum H. Hallier
*kajoe sémoet = Vitex quinata (Lour.) F. N. Will.
kaju titi = Gmelina macrophylla Wall.
kaju titie = Gmelina macrophylla Wall.
kaju tittie = Gmelina macrophylla Wall.
kaka kairkau = Stachytarpheta jamaicensis (L.) Vahl
kakoli = Clerodendrum inerme (L.) Gaertn.
*kalimantau = Viticipremna philippinensis (Turcz.) H. J. Lam
*kalipa&pa = Teijsmanniodendron Ahernianum (Merr.) Bakh.
ka liu tsoi = Vitex quinata (Lour.) F. N. Will.
*kaluingun = Gmelina elliptica J. ". Sm.
*kamalan = Viticipremna philippinensis (Turez.) H. J. Lam
kamiyo = Verbena delticola Small, V. Gooddingii var. nepeti-
folia Tidestr.

a

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76 PHYTOLOGIA Vol. 2, no. 8

*kenanga woeba = Gmelina asiatica var. villosa Bakh.
*keng mao = Gmelina elliptica J. E. Sm.
kani ba = Lippia adoénsis Hochst.

kapni = Holmskioldia sanguinea Retz.
*karnika = Premna corymbosa (Burm. f.) Rottl. & Willd.

karuana = Premna foetida Reinw.

kasaroballi = Citharexylum macrophyllum Poir.
*kasopaéngil-gibat = Clerodendrum minahassae Teijsm. & Binn.
kataboawin = Vitex rivularis Gtirke

katu-hinguru = Lantana sp., L. Camara var. aculeata (L.) Mok

denke
kawiyo = Verbena pumila Rydb.
*kemandiang = Gmelina asiatica var. villosa Bakh.
kena-qele-yago = Stachytarpheta urticaefolia (Salisb.) Sims
kenhenda = Clerodendrum serratum (L.-) Moon
*k8tiléng = Vitex quinata (Lour.) F.N. Will.
*ki bangbara = Vitex quinata (Lour.) F. N. Will-
kimbar mahalba = Lantana Mearnsii Moldenke
kimbo = Lippia adoénsis Hochst.
kingkilli ba = Lippia adoénsis Hochst.
*ki pahang = Premna corymbosa (Burm. f.) Rottl. & Willd.
*ki seungit = Premna corymbosa (Burm. f.) Rottl. & Willd.
*koefo-koefo = Vitex quinata (Lour.) F.N. Will.
*koetiléng = Vitex quinata (Lour.) F.N. Wille
*kojoe semoet = Vitex quinata (Lour.) F. N. Will.
koli = Clerodendrum inerme (L.) Gaertn.
ko. ling ngio = Vitex Negundo L.
*kalipapa = Teijsmanniodendron Ahernianum (Merr.) Bakh.
koorsoe wiwierie = Lantena Camara L.
korlejiga = Clerodendrum capitatum (Willd.) Schum. & Thonn.
koro koronta = Vitex Fosteri OC. H. Wright
koto = Vitex Doniana Sweet 4
kpar-seh = Vitex oxycuspis J. G. Baker, V. rufa A. Chev.
kua = Premna foetida Reinw.
kuabalon = Premna corymbosa var. sambucina (Wall.) Moldenke
kudu = Vitex Doniana Sweet
kukpweli = Vitex Doniana Sweet, V. grandifolia Gtirke
*lu-ku = Clerodendrum minahassae Teijsm. & Binn.
kukui = Vitex Doniana Sweet, V. grandifolia Glirke
*kulipdpa = Teijsmanniodendron Ahernianum (“ferr.) Bakh.
kama-tsuzura = Verbena officinalis L.
kumbil = Gmelina philippensis Cham.
kuru = Vitex barbata Planch., V. chrysocarpa Planch., Ve
Simplicifolia Oliv.
kurugh = Vitex Doniana Sweet
kuru kudulé = Vitex madiensis Oliv.
kutu-fingo = Vitex barbata Planch.
kwai tim foh = Clerodendrum canescens Wall.
kyet yo = Vitex pinnata L.

a a i a a i

1944 Moldenke, Common Names 77

*laban = Vitex guinata (Lour.) F. N. Will.

la ché rat = Stachyterpheta cayennensis (L. ©. Rich.) Vahl
lagoon tree = Avicennia africana P. Beauv.

lagrima de Cristo = Clerodendrum Thomsonae var. delectum Hort.
*lagund{ = Vitex trifolia var. bicolor (Willd.) Moldenke
lagunding dagat = Vitex trifolia var. simplicifolia Cham.

lagunding gapang = Vitex trifolia var. simplicifolia Cham.

lala tea = Vitex trifolia var. bicolor (Willd. ) Moldenke

lampaya = Lampaya medicinalis R. A. Phil.

lampayo = Lampaya medicinalis R. A. Phil.

lantan = Lantana Camara var. aculeata (L.) Moldenke

Lantana = Lantana Camara var. aculeata (L.) Moldenke, L. Cam-
ara var. mista (L3) bs Hs Balley, L. horrida H.B.K., Le.
insularis Moldenke, L. montevidensis (Spreng.) Brigq.,
L. scorta Moldenke, L. trifolia L.

lantanna = Lantana Camara var. flava (Medic.) Moldenke, Le
Camara var. hybrida (Neubert) Moldenke, L. Camara var.
mutabilis (Hook.) L. H. Bailey, L. Camara var. nivea
(Vent.) L. H. Bailey, L. Camara var. sanguinea (Medic.)
L. H. Bailey, L. Camara var. varia (Kuntze) Moldenke

large-bracted vervain = "Verbena bipinnatifida Nutt."
{error for V. bracteata Lag. & Rodr.]

large-bracted vervaine = Verbena bracteata Lag. & Rodr.

large-bracted vervane = Verbena bracteata Lag. & Rodr.

large flowered verbena = Verbena canadensis (L.) Britton

large flower verbena = Verbena canadensis (L.) Britton

large-leaved vervain = Verbena bracteata Lag. & Rodr.

lavender ground-flower = Verbena bracteata Lag. & Rodr.

*layaupan = Geunsia flavida (Eim.) He J. Lam, G. pentandra
(Roxb.) Merr.

lazo de amor = Verbena tenuisecta Briq.

*1Sban boenga = Vitex quinata (Lour.) F.N. Will.

le bois cotelet = Citharexylum B. Juss.

le bois de guitard = Citharexylum B. Juss.

le cotelet = Citharexylum B. Juss.

*Leierholz = Citharexylum B. Juss.

*18i18m in asoe = Clerodendrum minahassae Teijsm. & Binn.

*]i18m in taloen = Clerodendrum minahassae Teijsm. & Binn.

2

leja gado = Citharexylum macrophyllum Poir.

*lentang ruas = Sphenodesme pentandra Jack

liane rude = Petrea Kohautiana Presl

lilac lantana = Lantana Camara var. aculeata (L.) Moldenke,
L. Camara var. mutabilis (Hook.) L. H. Bailey

*iffigei = Vitex trifolia var. bicolor (Willd.) Moldenke

*lifigo-lingo = Viticipremna philippinensis (Turcz.) H. J.
Lam

- *linolfno = Viticiprema philippinensis (Turcz.) He J. Lam

*liffo-lffio = Viticipremna philippinensis (Turez.) H. J. Lam

lizard's tail = Stachytarpheta jamaicensis (L.) Vahl

78 ~ PHYTOLOGIA Vol. 2, no. 3

*loewarang = Gmelina asiatica var. villosa Bakh.

lo hai ngan = Callicarpa cana L.

1o kop ngan = Callicarpa longifolia Lam.

*loloet = Clerodendrum Rumphianum De Vriese

long-fruited duranta = Durante Mutisii L. f.

long-spiked fiddle-wood = Citharexylum caudatum L.
*Loosbaum = Clerodendrum Burm.

*Losbaum = Clerodendrum Burm.

*lotboom = Clerodendrum Burm.

lubei = Vitex Doniana Sweet, V. grandifolia Giirke

lugbei = Vitex Doniana Sweet, V. grandifolia Giirke

lung nga ts'o = Verbena officinalis L.

luwu-wului = Vitex Doniana Sweet, V. grandifolia Gtirke

lyre = Lantana Camara var. mista (L.) L. H. Bailey

madan polan = Clerodendrum fragrans var. pleniflorum Schau.
Madge Roberts Verbena = Verbena Teasii Moldenke

*madolau = Geunsia flavida.(@lm.) H. J. Lam *
*magilak'= Geunsia Cumingiana (Schau.) Rolfe

*magomo = Viticipremna philippinensis (Turez.) H. J. Lam
makwaiwa = Vitex Doniana Sweet :

*malabulaon = Symphorema luzonicum (Blanco) Fern.-Will. 3

*malafgang = Teijsmanniodendron Ahernianum (Merr.) Bakh. ‘

*mala-moldve = Viticiprema philippinensis (Turcz.) H. J. q
Lam - 4

*malamulduin = Viticipremna philippinensis (Turcz.) H. d. ;
Lam

*malapangit = Tectona philippinensis Benth.

*malasiad = Symphorema luzonicum Blanco) Fern.-Will.

*malasiag = Symphorema luzonicum (Blanco) Fern.-Will.

*malaskog = Symphorema luzonicum (Blanco) Fern.-Will.

*malatabaéko = Geunsia Cumingiana (Schau.) Rolfe 4

*malduing-aso = Viticipremna philippinensis (Turez.) H. J.
Lam

*mala-usd = Viticipremna philippinensis (Turcz.) H. J. Lam

male = Clerodendrum Buchholzii Gurke

*malet = Caryopteris odorata (Hamilton) B. L. Robinson

malmequer do mato = Lippia alba (Mill.) N. &. Br

*malvena = Lippia Recolletae Morong .

*mamahit = Teijsmanniodendron Ahernianum (Merr.) Bakh.

*mamali = Vitex quinata (Lour.) F.N. Will.

mamath = Verbena hybrida Voss

mameira = Vitex flavens H.B.K.

Mammoth Rose Queen Verbena = Verbena hybrida Voss

Mammoth Scarlet Queen Verbena = Verbena hybrida Voss

Mammoth Snow Queen Verbena = Verbena hybrida Voss

*manabéko = Geunsia Cumingiana (Schau.) Rolfe

mandarin's-hat = Holmskioldia sanguinea Retz.

man kaka kakkan = Stachytarpheta cayennensis (L. C. Rich.)
Vahl

1944 | Moldenke, Common Names 79

manpraseara = Aegiphila laeta H»B.K., A. laevis (Aubl.) Gmel.

manzanillo = Lippia integrifolia (Griseb.) Hieron.

ma pin ts'o = Verbena officinalis L.

margarita morada = Verbena dissecta Willd., *V. laciniata
(L.) Brig.

margarita punzé = Verbena incisa Hook.

*masarwet = Vitex quinata (Lour.) F. N.. Will.

‘mashayi = Clerodendrum capitatum (Willd.) Schum. & Thonn.

mata negro = Junellia tridens (Lag.) Moldenke

matorro moro = Junellia Lorentzii (Niederlein) Moldenke

maukakarawa = Stachytarpheta urticaefolia (Salisb.) Sims

mbalhat = Lippia adoénsis Hochst.

mbormbor = Lippia adoénsis Hochst.

mbougand = Avicennia africana P. Beauv.

mejorana = Lantana macropoda Torr.

*middf-gass = Premna corymbosa (Burm. f.) Rottl. & Willd.

misiwahchil = Holmskioldia sanguinea Retz.

Miss Willmott Verbena = Verbena hybrida Voss

misteriosa olorosa = Clerodendrum fragrans var. pleniflorum
Schau «

mofalu = Stachytarpheta urticaefolia (Salisb.) Sims

*mohéni = Oaryopteris odorata (Hamilton) B. L. Robinson

mokaukarau kedra = Stachytarpheta urticaefolia (Salisb.)
Sims

molauin = Vitex parviflora A. L. Juss.

molave = Vitex parviflora A. L. Juss.

*nongpong = Teijsmanniodendron Ahernianum (Merr.) Bakh.

*moni = Caryopteris odorata (Hamilton) B. Le Robinson

monks-pepper-tree = Vitex Tourn.

moradia = Verbena delticola Small

moradilla = Verbena ciliata Benth., V. elegans var. asperata
Perry .

mosongo-songo = Olerodendrum Buchholzii Gtirke

*moss verbena = Verbena laciniata (L.) Briq.

‘motofu = Stachytarpheta urticaefolia (Salisb.) Sims

(ar oS oy

mouse's bowstring = Stachytarpheta jamaicensis (L.) Vahl
*maldéuin = Viticipremna philippinensis (Turcz.) H. J. Lam
*mul duin g-baging = Symphorema luzonicum (Blanco) Ferb.-Will.
mullenm-leafed vervain = Verbena stricta Vent.

_“munnay = Prema corymbosa (Burm. f.) Rottl. & Willd.

“miney kiray = Premna corymbosa (Burm. f.) Rottl. & Willd.

‘*munni-vayr = Premna corymbosa (Burm. f.) Rottl. & Willd.

Mutis's duranta = Durante Mutisii L. f.

mu-tewani = Lippia scabra Hochst.

mutuku-tsho = Avicennia africana P. Beauv.

muyuyu de) monté = itharexylum quitense Spreng.

myrrh tree = Vitex Agnus-castus L.-

‘nage = Premna corymbosa var. obtusifolia (R. Br.) Fletcher

“nago = Geunsia Oumingiana (Schau.) Rolfe

80 PHeYot Op oO. Gtk Vol. 2, no. 3

nahuire = Aloysia nahuire Gentry & Moldenke

nembalerri = Vitex s simplicifolia Oliv.

narenga = Vitex Dor Doniana Sweet, V. grandifolia Gtirke

*narvel = Prema corymb: corymbosa (Burm. a) Rottl. & Willd.

Nassau-rose = Olerodendrum fragrans var. pleniflorum Schau.

*nay-mof-si = Vitex rapinoides Sai Tlsum.

nebedda = Vitex leucoxylon L. f.

negrito s Vitex pyramidata B. L. Robinson

*néla nfredu = Pygmaeopremna herbacea (Roxb.) Moldenke

_ nettle leaved vervain = Waviead urticifolia L.

nettle-leaved vervain = Verbena Zngelmannii Moldenke, V. ur-
ticifolia var. leiocarpa Perry & Fernald

nettle leaved Virginian vervain = Verbena urticifolia L.

ngakawa = Faradaya ovalifolia (A. ee Seem.

ngalbihi = Vitex Doniana Sweet

ngasu = Lappis pia adoénsis Hochst.

ng&suru = Lippia adoénsis Hochst.

ng chi fung = Vitex Negun Negundo L.

ngoh sat na = Verbena officinalis L.

ngurunguru = Prema Gaudichaudii Schau.

nici = Prema taitensis Schau.

nigua = Cornutia obovata Urb.

nika = Vitex Negundo L.

nifia rupé = Aloysia ligustrina (Lag.) Small

niue = Vitex "quate cLour.) F. N. Will.

nja-wului = af 2 dat canis africana P. Beauv.

nomeoluides = Duranta repens var. alba (Masters) L. H. Bailey

no=me=oluides = Duranta repens var. + alba (Masters) L. H.
Bailey

*ném méo = Gmelina elliptica J. 3. Sm.

nsunsu = Stachytarpheta jamaicensis (L.) Vahl

nuenu-pichada = Stachytarpheta cayennensis (Le G. Riche
Vahl

nuna del monte = Aloysia ligustrina (Lag.)- Small

nya = Vitex Doniana Sweet, V. grandifolia Gtirke

nyam@le-kukwe = Vitex grandifolia Giirke

nyarine = Vitex Doniana Sweet, V. grandifolia Gtirke

ny &kpe = Clerodendrum capitatum um (Willd.) Schum. & Thonn.

nyo6na = Lippia adoénsis Hochst.

obranmotuam = Clerodendrum c capitatum (Willd.) Schum. & Thonn.

obuban = Vitex Fosteri C. H. Wright

Scha koro = Vitex Doniana Sweet .

odonumon = Avicennia africana P. Beauv.

oema koorsoe wiwirie = Lantana Camara var. mista (L.) L. He
Bailey

oeroejatoe = Citharexylum macrophyllum Poir.

-ogboso-tsho = Premna quadrifolia Schum. & Thonn.

ogbosu = Prema guadrifolia Schum. & Thonn.

Ogbun = Avicennia africana P. Beauv.

a =)
7 ery

1944 Moldenke, Common Names 81

Ogi = Vitex Fosteri CO. H. Wright

ogikhimi = Vitex grandifolia Gtirke

oi = Stachytarpheta jamaicensis (L.) Vahl

ojédiballi = Stachytarpheta cayennensis (L. C. Rich.) Vahl
okurutu = Vitex grandifolia Gtirke

old man's shin-bone = Vitex rivularis Giirke

Spd para = Stachytarpheta jamaicensis (L.) Vahl

opé-éshi = Clerodendrum splendens G. Don

orabia = Vitex grandifolia Gtirke

orcujuela = Citharexylum Berlandieri B. L. Robinson

oreganillo = Aloysia ligustrina (Lag.) Small

oregano = Lippia alba (Mill.) N. H. Br.
orégano = Lippia Berterii Spreng., L. micromera Schau.
orégano de burro = Lippia Berterii Spreng.

orégano di burro = Lippia affinis Schau.

Ori = Vitex Doniana Sweet, V. grandifolia Gtirke

6ri-%ta = Vitex Fosteri C. H. Wright

origanum = Lippia micromera Schau.

Sri-nla = Vitex Doniana Sweet

Sri-odan = Vitex Doniana Sweet

oriri = Vitex grandifolia Gtirke

orozuz de latierra = Phyla scaberrima (A. L. Juss.) Moldenke
Stwe-ntdrowa = Vitex rivularis Gtirke

oviakuku = Clerodendrum Thomsonae Balf. f.

ovuruburu = Vitex grandifolia Gtirke

Swenkundigbon = Vitex grandifolia Gtirke

Oxford Pink Verbena = Verbena hybrida Voss

paak pui ip = Vitex trifolia var. simplicifolia Cham.
pagil = Vitex pinnata L.

pagode-flower = Clerodendrum Burm., C. paniculatum L.
pak yat hung = Clerodendrum Kaempferi (Jacq.) Sieb.
pak yat pak = Clerodendrum fragrans (Vent.) Re Br.

palo amarillo = Aloysia ligustrina (Lag.) Small

*palo blanco = Citharexylum Kunthianum Moldenke
*pamagsen = Teijsmanniodendron Ahernianum (Merr.) Bakh.
pampa oregano = Lippia alba (Mill.) N. %. Br.
*pamulaklakin = Symphorema luzonicum (Blanco) Fern.-Will.
*pananagok = Geunsia flavida (Elm.) H. J. Lam

panda = Vitex Stahelii Moldenke

pan poregano = Lippia alba (Mill.) N. %. Br.

panyerd = Vitex Doniana Sweet

panyerd buda = Vitex simplicifolia Oliv.

papagaio = Aegiphila Sellowiana Cham.

*paper flower = Sphenodesme pentandra Jack

paraguita de China = Holmskioldia sanguinea Retz.
parasol-flower = Holmskioldia sanguinea Retz.

parwa = Avicennis nitida Jacq.

pau de tamanco = Aegiphila Sellowiana Cham.

pau de viola = Citharexylum myrianthum Cham.

82 PHYTOLOGIA Vol. 2, no. 3

pechiche = Vitex gigantea H.B.K.
pedrésy = Stachytarpheta Mexiae Moldenke
pendola de sierra = Citharexylum caudatum L.
pendiila = Citharexylum fruticosum L. i
pengua = Priva aspera H.B.K.
*péragu = Clerodendrum Burm.
*péragut = Clerodendrum Burm.
perajil = Verbena tenuisecta Brig.
perennial verbena = Verbena canadensis (L.) Britton
pfufulla = Clerodendrum capitatum (Willd.) Schum. & Thonn.
piedrero = Vitex capitata Vahl
pigeonberry = Duranta L.
pimenteira = Citharexylum myrianthum Cham.
pingdang = Clerodendrum paniculatum L. | |
pink verbena = Verbena hybrida Voss, V. pumila Rydb. |
pink vervain = Verbena pumila Rydb.
*pinna-nelli = Prema corymbosa (Burm. f.) Rottl. & Willd.
pipe-tree = Clerodendrum capitatum (Willd.) Schum. & Thonn.
Plumier's duranta = Duranta repens L.
*pokok agak paya = Teijsmanniodendron pteropodum (Miq.)Bakh.
poko kwat tdn = Callicarpa macrophylla Vahl
poleo = Aloysia barbata (T. S. Brandeg.) Moldenke, Lippia
Berterii Schau., L. turbinata f. angustifolia Osten
poleo de burro = Aloysia polystachya (Griseb.) Moldenke
poleo de Castilla = Aloysia polystachya (Griseb.) Moldenke
poleo de Castillo = Aloysia polystachya (Griseb.) Moldenke
polinalina = Vitex trifolia var. simplicifolia Cham.
*ponranga = Gmelina asiatica var. villosa Bakh.
prickly lantana = Lantana Camara var. aculeata (L.) Moldenke
*puhting = Gmelina elliptica J. @. Sm.
*pukang mata hari = Gmelina elliptica J. %. Sm.
punyo-tsho = Vitex Doniana Sweet, V. grandifolia Gtirke
Purple Garnet Verbena = Verbena hybrida Voss
purple verbena = Verbena bipinnatifida Nutt., V. canadensis
(L.) Britton, V. hastata L.
purple wreath = Petrea racemosa Nees
puta de noche = Clerodendrum ternifolium H.B.K.
qarovo = Premna corymbosa var. sambucina (Wall.) Moldenke
-*Queensland beech = Gmelina Leichhardtii (F. Muell.) F.
Muell.
querendereniqua = Vitex pyramidata B. L. Robinson
queue de rat = Stachytarpheta jamaicensis (L.) Vahl
quilau = Clerodendrum disparifolium Blume
ra-bina = Clerodendrum splendens G. Don
rat's tail = Stachytarpheta jamaicensis (L.) Vahl
rauvula = Premna taitensis var. rimatarensis F. H. Br.
red lantana = Lantana Camara var. sanguinea (Medic.) Le He
Bailey
red verbena = Verbena hybrida Voss

ete i a
he Apo
we :
aeons
se +
tee
,

1944 Moldenke, Common Names 83

remako = Premma corymbosa var» sambucina (Wall.) Moldenke,
P. foetida Reinw.

rica-rica = Acantholippia deserticola (R. A. Phil.) Molden-
ke, A. hastulata CGriseb.

*rimoewas = Vitex quinata (Lour.) F. N. Will.

rithoul = Holmskioldia sanguinea Retz.

rosa blanca = Lantana velutina Mart. & Gal.

rose glorybower = Clerodendrum Bungei Steud.

rose verbena = Verbena canadensis as Britton

rouen = Verbena tenuisecta Briq.

Rowena Verbena = Verbena Teasii Moldenke

Ruth Verbena = Verbena Teasii Moldenke

saa-nunum = Lippia adoénsis Hochst.

sacha-poleo = Aloysia sp. (Salta)

sagarai = Vitex trifolia L., *V. trifolia var. bicolor
(Willd.) Moldenke

sah-sah = Vitex micrantha Giirke

sai fa min = Callicarpa cana L.

saigun = Tectona grandis L. f.

sai hong hun = Callicarpa formosana Rolfe

sai ip lo hai ngan = Callicerpa dichotoma (Lour.) K. Koch

sai ko din nuang = Prema acuminatissima Merr., P. octoner-
via Merr. & Metc.

sai tsio tau = Vitex quinata (Lour.) F.N. Will.

saivonta = Vitex quinata (Lour.) F. N. Will.

*sajor kambing = Prema corymbosa (Burm. f.) Rottl. & Willd.

saladillo = Avicennia nitida Jacq.

salvia = Aloysia salvisefolia (Hook. & Arn.) Moldenke, Lip-
pia alba (Mill.) N. &. Br.

salvia alta = Lantana velutina Mart. & Gal.

*salvia blanca = Aloysia salviaefolia (Hook. & Arn.) Mold-
enke

salvia morada = Lantana montevidensis (Spreng. ) Briq., Lip-
pia alba (Mill.) N. &. Br., L. Grisebachiana Moldenke

salvia santa = Lippia substrigosa Turcz.

samanibir = Vitex Doniana Sweet, V. grandifolia_ Giurke

*sambuyut = Geunsia Cumingiana (Schau.) Rolfe

sana = Avicennia africana P. Beauv.

sanango sacha = Petrea peruviana Moldenke

sandia lahuen = Verbena laciniata (L.) Briq., V. tenuisecta
Briq.

*sandialahuén = Verbena tenuisecta Brig.

San Judn de la verdad = Aegiphila puberulenta Moldenke

sanzgatillo = Vitex Agnus-castus L.

*saoce masarawet = Vitex quinata (Lour.) F. N. Will.

*sace poeti = Vitex quinata (Lour.) F.N. Will.

*sace réndai = Vitex quinata (Lour.) F. N. Will.

inata (

*sace sSla = Vitex qui ours ) Pathe Weil s
*“saonad = Gmelina elliptica J. @. Sm.

84 FP HY of O' 4--O7:G4i2 Vol. 2, now 3

sapu-milla = Vitex altissima L. f.

*sarogang salaki = Gmelina asiatica var. villosa Bakh.
*sasalit = Tei jsmanniodendron Ahernianum (Merr.) Bakh.
sa sha ping = Clerodendrum Bungei Steud.

*sasilit = Tei jsmanniodendron Ahernianum (Merr.) Bakh.
*sasulit = Teijsmanniodendron Ahernianum (Merr.) Bakh.
sauco monte = Aegiphila Deppeana Steud.

scarlet verbena = Verbena hybrida Voss

shah's favourite = Verbena L., V. hybrida Voss

shan pak tang = Sphenodesme pentandra Jack
sha-passan = Verbena L., V. hybrida Voss
*shechin = Oaryopteris odorata (Hamilton ) B. Le Robinson

shek tzi shu = Gmelina racemosa (Lour.) Merr.

short—hair: white vervain = Verbena urticifolia var. leiocar-
pa Perry & Fernald

siba = Prema corymbosa var. sambucina (Wall.) Moldenke

sibo = Prema foetida Reinw.

sidraera = Lippia alba (Mill.) N. &. Br.

silvery duranta = Duranta argentea Lodd.

simplers' joy = Verbena hastata L.

*singkil = Prema corymbosa (Burm. f.) Rottl. & Willd.

*singkil alas = as = Frenne corymbosa (Burm. f.) Rottl. & Willd.

sisiling hyamo = Lippia adoénsis Hochst.

skyflower = Duranta L.

*sky-flower = Duranta L.

slender vervain = Verbena Halei Small

small-flowered ES ae = "Verbena canadensis (L.) Britton"
(error for V. bipinnatifida } Nutt.

snake-rattle = Stachy tarpheta cayennensis (L. C. Rich.) Vahl

*sobsoganbogo = Geunsia’ Cuminpiana (Schau. ) Rolfe

solande = Lantana Camara var. mista (L.) L. H. Bailey

Soldaten Thee = Lantana Camera L. |

song-sho = Vitex Doniana Sweet, V. grandifolia Giirke

song tsio gun = Gmelina racemosa (Lour .) Merr.

so pa = Vitex trifolia var. simplicifolia Cham.

s6-tsho = = Vitex Doniana Sweet, V. grandifolia Gtrke

spatulate-lear - fog-fruit = " Aloysia macrostachya (Torr.)
Moldenke" [error for Phyla nodiflora (L.) Greene]

spatulate-leaved fog-fruit = Phyla incisa Small, FP. lanceo-
lata (Michx.) Greene

spider's kenki = Lantana Camara ie

*stilbé = Stilbe Berg.

*stidamerikanisches Tisenkraut = Verbena DOueT 4 Ouse Le

sumpin = Syhenodesme borneénsis Merr.

*susanna = Citharexylum B. Juss.

sweet sage = Lantana trifolia L.

sweet William = Verbena bipin bipinnatifida Nutt.

sylvania = Aegiphila martinicensis Jacq.

taasen dua = Clerodendrun capitatum (Willd.) Schum. & Thonn.,

! aed, + "
PO ee ee ee ee ae ae eS SS

pra Ce oa

1944 Moldenke, Common Names 85

C. formicarum Gtirke, C. polycephalum J. G. Baker
tabaquillo = Aegiphila intermedia Moldenke
tab&Sto = Clerodendrum capitatum (Willd.) Schum. & Thonn.
tabonsu = Stachytarpheta jamaicensis (L.) Vahl
*tabugok = Clerodendrum minahassae Teijsm..& Binn.
tai chung lo kop muk = Callicarpa nudiflora Hook. & Arn.
tai ip shan po = Prema octonervia Merr. & Metc.
t&l = Cornutia odorata (Poepp. & "ndl.) Poepp.
talabao = Lippia Pringlei Briq.
tala blanco = Duranta serratifolia (Griseb.) Kuntze
talalachi = Aegiphila monstrosa Moldenke
*talauan = Gmelina elliptica iptica J. &. Sm.
talitue = Prema corymbosa var. sambucina (Wail. ) Moldenke
tall ssegiphila = Aegiphila elata Sw.
*talunigud | = Gmelina elliptica J. =. Sm.
*taliingun = Gmelina elliptica J. E. Sm.
tamanqueira = = Aegiphila Sellowiana Cham.

tanagyea = Stachytarpheta _ lamaicensis (L.) Vahl
*tanliungun = Gmelina elliptica J. 2. Sm.

tanodza = Seacaytarshts jamaicensis (L.) Vahl

*tarbay = raveolens H. eee

tarete = Seer cap ake Mill.) N. &. Br.

tarrafe = aoe aie nia africana P. Beauv.

taruma = Vitex cymosa Bert.

tarumé = Vitex’ calothyrsea Sandw., V. flavens H.B-K.
taruma = Vitex montevidensis Cham., > V. polyg polygama Cham.
taruma de mata = Vitex Froesii Moldenke

tarumé de terrafirma = Vitex triflora Vahl

tarumé de varzea = Vitex cymosa Bert.

_ taruman = Vitex Schaueriana Moldenke

tarumén = Citharexylum montevidense (Spreng.-) Moldenke
tarumansinho = Vitex Schaveriana Moldenke

tasajo = Vitex orinocensis H.B.K.

tataba = Olerodendrum capitatum (Willd.) Schum. & Thonn.
tatumo = Aegiphile tri truncata Moldenke

tavotavo = Premna taitensis Schau.

tavu = Prema taitensis var. rimatarensis F. H. Br.
*tayupuk = Teijsmanniodendron Ahernianum (Merr.) Bakh.
tcaucui = Clerodendrum barba-felis H. Hallier

*tea plant = Lantana Camara L.

Teas Hybrid separ = Verbena Teasii Moldenke

té del pais = Lippia turbinata Griseb.

*te del pais = Lippia " graveolens H.BeK.

te negro = la eee (L.) Small

*thé de piéton = Lippia Pseudo-thea (A. St. Hil.) Schau.

-tialu = Verbena officinalis L.

ti dah = Clerodendrum umbellatum Poir.

tinho = Stachytarpheta australis Moldenke
tiogbi = Vitex ferruginea Schum. & Thonn.

86 PHYTOLOGIA Vol. 2, no. 3

tokalau = Lantana Camara var. aculeata (L.) Moldenke

tolochocho = Lantana velutina Mart. & Gal.

tomillo = Acantholippia seriphioides (A. Gray) Moldenke,
Junellia asparagoides (Gill. & Hook.) Moldenke

tomillo macho = Junellia seriphioides (Gill. & Hook.) Mold-
enke

toronjil = Lantana Langlassei Moldenke

tosatido = Petrea arborea H.Bek.

“tostadito = Petrea aspera Turcz.

totumillo blanco = Vitex divaricata Sw.

*toung-than-gyoe = Premma corymbosa (Burm. f.) Rottl. &
Willd.

tra-tsho = Avicennia africana P. Beauv.

tres colores = Lantana glandulosissima Hayek
trdmen = Clerodendrum capitatum (Willd.) Schum. & Thonn. 1
tearkiyar kusu = Stachytarpheta jemaicensis (L.) Vahl
tachangbaio = Vitex Doniana Sweet
tschangmaro = Vitex Doniana Sweet

tschingmera = Vitex Doniana Sweet

tuetu = Stachytarpheta jamaicensis (L.) Vahl
*tugas = Viticipremma philippinensis (Turez.) H. J. Lam
*tugas-buhgogon = Viticipremna philippinensis (Turcz.) H. J.

Lam
*tulufigun = Gmelina elliptica J. %. Sm.
*tufgdlnol = Gmelina elliptica J. &. Sm.
turkey tangle = Phyla nodiflora (L.) Greene .
turu-Llevu = Stachytarpheta urticaefolia (Salisb.) Sims
tzou tsing tsoi = Clerodendrum cyrtophyllum Turcz.
ububan = Vitex rivularis Gtrke
ucha koro = Vitex Doniana Sweet

ucullucui-sacha = Stachytarpheta cayennensis (Lb. C. Riche):

Vahl
ucullucuy sacha = Stachytarpheta cayennensis (L. C. Rich.)
Vahl

ufiri = Avicennia africaria P. Beauv.

ufuchi = Clerodendrum splendens G. Don

um-diguilgul = Vitex Doniana Sweet

um-dugulgun = Vitex Doniana Sweet

unarmed duranta = Duranta repens L.

u8li = Vitex Doniana Sweet

urdi loho'be = Lantana Mearnsii Moldenke

uruahu = Vitex grandifolia Girke

usillo = Aloysia ligustrina (Lag.) Small °
-*ustabunda = Prema corymbosa (Burm. f.) Rottl. & Willd.

' uvalema’= Vitex pyramidata B. L. Robinson
uvulama = Vitex mollis H.B.K., V. pyramidata B. L. Robinson
vanilla-= Duranta repens L.

vara de San José = Castelia cuneato-ovata Cav.

vara dulce = Aloysia macrostachya (Torr.) Moldenke

4

‘ *

Y
rm
a

1944 . Moldenke, Common Names 87

varo = Premma foetida Reinw.

vasari = Vitex cofassus Reinw.

*vasung = Viticiprema philippinensis (Turcz.) H. J. Lam

*vedelhoutboom = Citharexylum B. Juss.

vedel houthboom = Citharexylum B. Juss.

venturosa = Lantana trifolia L.

verbean vervain = Verbena bracteata Lag. & Rodr.

verbena = Stachytarpheta trimitensis Moldenke, Verbena cana-
densis G.) Britton, V. carolina L., V. domingensis
Urb., V. ephedroides Cham., V. hispida Ruiz & Pave, V.
integrifolia Sessé & Moc., V. laciniate (L.) Briq., V.
tenuisecta Briq.

verbena ancha = Stachytarpheta cayennensis (L. C. Rich.)
Vahl

verbena aztil = Stachytarpheta jamaicensis (L.) Vahl

verbena blanca = Stachytarpheta cayennensis (Le C. Rich.)
Vahl, Verbena platensis Spreng.

verbena blanca serrana = Verbena litoralis H.B.K.

verbena cim = Stachytarpheta jamaicensis (L.). Vahl

verbena falsa = Stachytarpheta cayennensis (L. C. Rich.)Vahl

verbena negra = Stachytarpheta cayennensis (L. 0. Rich.) _
Vahl, S. straminea Moldenke

verbena oil — from Aloysia triphylla (L'Hér.) Britton

verbena shrub = Caryopteris incana (Thunb.) Miq.

verbena-shrub = Caryopteris Bunge

verevere = Clerodendrum inerme tL.) Gaertn.

vervain = Verbena bipinnatifida Nutt., V. canadensis (L.)
Britton, V. canescens var. Roemeriana (Scheele) Perry;
Ve Halei Small, V. simplex Lehm., V. xutha Lehn.

*verveine 4 fleurs rouges = Stachytarpheta mutabilis (Jacq.)

_ Vahl
verveine de Drummond = Verbena canadensis (L.) Britton
verveine de Miquelon naine magenta = Verbena canadensis (L.)

Britton
verveine de Miquelon naine rose = Verbena canadensis (53
Britton

*verveine élégante = Verbena tenuisecta Brigq.
vervena = Verbena litoralis H.B.K.

villatermin = Duranta repens L.

Violet King Lantana = Lantana tiliaefolia Cham.
violette = Verbena tenuisecta Briq.

. Virginia sage = Vitex Agnus-castus L.

voekoe voekoe tolman = Priva lappulacea (L.) Pers.
vulcana = Clerodendrum fragrans var. pleniflorum Schau.
wilokaka = Vitex trifolia var. simplicifolia Cham.
waiwai = Lantana Camara var. aculeata (L.) Moldenke
wakarovungi = Faradaya ovalifolia (A. Gray) Seem.
wakorovundi = Faradaya ovalifolia (A. Gray) Seem.
wal-gurenda = Clerodendrum inerme (L.) Gaertn.

88 PH YT 0-6 O3Gcr se Vol. 25 noe 3

wan hon na wan njari = Clerodendrum japonicum (Thunb.) Sweet

*waréng = Gmelina elliptica J. 3. Sm.

*waréng kétan = Gmelina asiatica var. villosa Bakh., G. ell-
iptica J. &. Sm.

waro = Prema Gaudichaudii Schau.

waro ndamu = Prema Gaudichaudii Schau.

Warowaro = Prema taitensis Schau.

waterside tree = Avicennia africana P. Beauv.

wa vatu = Faradaya ovalifolia (A. Gray) Seem., F. vitiensis

(A. Gray) Seem.

wedgeleaf frog-fruit = Phyla cuneifolia (Torr.) Greene

wedgeleaf frog fruit = Phyla incisa Small

wedge-leaved fog-fruit = Phyla nodiflora var. reptans

(H.BeK.) Moldenke

weighty fog-fruit = Phyla incisa Small

westindisches Eisenholz = Aegiphila martinicensis Jacq.

wesussi = Stachytarpheta jamaicensis (L-) Vahl -

*wSwéngenga = Gmelina asiatica var. villosa Bakh.

weyhooli = Verbena menthaefolia Benth..

*white beech = Gmelina Leichhardtii (F. Muell.) F. Muell.

white Brazil mangrove = Avicennia nitida Jacq.

white brush = Aloysia ligustrina var. Schulzii (Standl.)
-Moldenke

white bush = Aloysia ligustrina (Lag.-) Small
-white-flowered lantana = Lantana Camara var. nivea (Vent.)

L. H. Bailey
white-flowered verbena = Verbena stricta f. albiflora Wad-
‘mond

white-fruited callicarpa = Callicarpa americana var. lactea
F. J. Muller

white mangrove = Avicennia africana P. Beauv.

white-verbena = Verbena hybrida Voss. V. urticifolia L.

widow of last year = Clerodendrum splendens G. Don

wild coffee = Clerodendrum aculeatum (L.) Schlecht.

wild hyssop = Verbena stricta Vent.

wild sage = Lantana Camara var. mista (L.) L. H. Bailey

wild verbena = Verbena canadensis (L.) Britton

*woelas watoe = Vitex quinata (Lour.) F. N. Will.

work is sweet = Clerodendrum violaceum Gtirke

watsiyar 'bera = Stachytarpheta jamaicensis (L.) Vahl

wutsiyar kadangare = Stachytarpheta jamaicensis (L.) Vahl

wutsiyar kusu = Stachytarpheta jamaicensis (Lie) Vahl. 2)

*xakilche = Lippia graveolens H.B.K.

Xalapa duranta = Duranta repens L. |

yabu-murasaki = Callicarpa mollis Sieb. & Zucc

yapau = Verbena L. . |

yaro = Prema taitensis Schau., P. taitensis var. rimatar-
ensis F. H. Br.

yellow lantana = Lantana Camara var. flava (Medic.) Molden-

—- Tie So. se

Pea

1944 Moldenke, Common Names 8&9

ke, L. tiliaefolia Cham.
yellow sage = Lantana trifolia L.
yemo sigba = Lantana Camara L.
yerba de Inca = Lippia integrifolia (Griseb.) Hieron.
yerba de la muestranza = Lantana Camara L. :
yerba de la princesa = Aloysia triphylla (L'Hér.) Britton
yerba de la Virgen = Phyla nodiflora var. rosea (D. Don)
Moldenke
yerba del Cristo = Lantana horrida H.B.K.
yerba del incordio = Yerbena tenuisecta Briq.
yerba dulce = Phyla scaberrima (A. L. Juss.) Moldenke
yerba mora = Lantana Camara L.
yeung ue yi fa = Lantana montevidensis (Spreng.) Brig.
yoch opp tzimin = Petrea volubilis L.
yoimte = Clerodendrum ligustrinum (Jacq.) R. Br.
*Zitronenlippe = Lippia Houst.

ayvos 3 Vitex Agnus-castus L.
Avyapra = Vitex Agnus-castus L.
Avy os = Vitex Agnus-castus L.
Avyos = Vitex Agnus-castus L.

TOOS = Vitex Agus-cestus L.
oligos = Vitex Agnus-castus L.

(1) Moldenke, H. N., An alphabetic list of common and ver-
neacular names recorded for members of the Verbenaceae
and Avicenniaceae. 34 pp. New York Botanical Garden,
August 31, 1939.

(2) Moldenke, H. N., A supplementary list of common and ver-
nacular names recorded for members of the Verbenaceae
and Avicenniacese. 24 pp. New York Botanical Garden,
February 25, 1940.

THE RECORDED COMMON AND VERNACULAR NAMES OF VERBENACEAE AND
AVICENNIACEAE ARRANGED ACCORDING TO GENERA AND SPECIES

Harold N. Moldenke

The following is a list of the common and vernacular
names of Verbenaceae and Avicenniaceae which were recorded
by me in alphabetic sequence in my previous publications on
this subject (1). In all, four thousand six hundred and one
names are here recorded.

90 PHYTOLOGIA Vol. 2, no. 3

Acantholippia deserticola (R. A. Phil.) Moldenke -- rica-
rica

Acantholippia hastulata Griseb. -- rica-rica

Acantholippia seriphioides (A. Gray) Moldenke -- tomillo

Aegiphila Jacq. -- aegiphilas, aegiphile, bois cabril,
Geissenbaéumchen, geitenboompje, goatwood, Ziegenbaumch-
en, Ziegenstrauch

Aegiphila aculeifera Moldenke -- tabaquilla

Aegiphila alba Moldenke -- koit tree, lulu, tutumbo

Aegiphila anomala Pittier. -- tabaquillo

Aegiphila chrysantha Hayek -- fetord-ey

Aegiphila Deppeana Steud. -- saiico de monte, sauco monte

Aegiphila elata Sw. -- bejuco de peine mico, guairo santo,
guaro, guauro, hoher Ziegenstrauch, Kenia weed, tall
aegiphila

Aegiphila falcata Donn. Sm. -- ah pl nai zorrillo

Aegiphila ferruginea Hayek & Spruce -=- valso

Aegiphila foetida Sw. -- stinkender Ziegenstrauch

Aegiphila glandulifera Moldenke -- chirapa sacha

Aegiphila glandulifera var. pyramidata L. 0. Rich. & Molden-
ke -- tabaquero

Aegiphila integrifolia (Jacq.) Jacks. -- baumartiger Ziegen-

. strauch, bois de golette, bois sendu, carindiba, lard-
wood, tocaneiro

Aegiphila intermedia Moldenke -- tabaquillo

Aegiphilea laeta H.B.K. -- manprasara, San Juan de la Verdad

Aegiphila laevis (AublL.) Gmel. -- gelber Ziegenstrauch,
manabo, man manprasara

Aegiphila laxicupulis Moldenke -- palo de zope
Aegiphila martinicensis Jacq. -- bastard white-root, bois

cabril, bois cabrit, bois de bouc, bois de cabril, bois

de fer, capaillo, lengua de vaca, martiniquische
Geissenb&umchen, martiniquischer Ziegenstrauch, sureau
gros, sylvania, westindisches ®isenholz, wild-jasmine

Aegiphila martinicensis var. gh poneure (Urb. ) Moldenke -=
bois cabrite

Aegiphila mollis H.B.K. -- contra-culebra, heilkréftiger
Ziegenstrauch, totumillo

Aegiphila monstrosa Moldenke -- café cimarron, hulub,
talalachi, vara blanca

-Aegiphila multiflora Rufz & Pav. -- utcus

Aegiphila obovata Andr. -- cutlet-wood, timber fiddlewood

Aegiphila panamensis Moldenke -- hombre grande

Aegiphila perplexa Moldenke -- goat-meat

Aegiphila peruviana Turcz. -- Sci sacha, huaca,
ucullucuy sacha

Aegiphila puberulenta Moldenke -- bollo limpio, San Juan de

la verdad

Aegiphila racemosa Vell. -- cawuira, wanini

ST ee eee

1944, Moldenke, Common Names 91

Aegiphila Riedeliana Schau. -- cajuja

Aegiphila Sellowiana Cham. -- cajugd, carindiba, caujuja,
cinzeiro, habiara, papagaio, pau de tamanco,
tamanqueira

Aegiphila splendens Schau. -- serra dos Christae

Aegiphila truncata Moldenke -- tatumo _

Aegiphila Valerii Standl. -- tabaquillo

Aegiphila verrucosa Schau. -- chicharra

Aegiphila villosa (Aubl.) Gmel. -- bois de tabac, bois tabac,

moracooballi, wolliger Ziegenstrauch

Aegiphila vitelliniflora Klotzsch -- caferana, fetord-ey

Aloysia barbata (T. 8. Brandeg.) Moldenke -- poleo

Aloysia ligustrina (Lag.) Small -- dngel, azahar del campo,
bee brush, cedron, Mexican heliotrope, nina rupd, nuna
del monte, oreganillo, palo amarillo, romerillo,
usillo, white bush

Aloysia ligustrina var. Schulzii (Standl.) Moldenke -- bee
blossom, white brush

Aloysia Looseri Moldenke -- ilang-ilang, ildn-ildn

Aloysia macrostachya (Torr.) Moldenke -- cabradora simarona,
vara dulce

Aloysia nahuire Gentry & Moldenke -- nahuire

Aloysia polystachya (Griseb.) Moldenke -- poleo de burro,
poleo de Castilla, poleo de Castillo

Aloysia salviaefolia (Hook. & Arn.) Moldenke -- salvia,
salvia blanca

Aloysia triphylla (L'Hér.) Britton -- Aloisekraut, cedron,
cedron, citroenboompje, citroenkruid, citroenverbena,
citronelle, citronenduftende Lippie, citronenduftige
Lippie, Citronen-kraut, Citronensstrauch, citron-
scented lippia, dreibladige lippia, herba Luisa, herb
Louisa, herva cidreira, Lemonekraut, lemon plant,
lemon-scented verbena, lemon~scented vervain, lemon
tree, lemon verbena, limouneto, lippie & odeur de
citron, pigeon's herb, Punschkraut, scented verbena,
swest-scented verbena, thé arabe, verbena oil, vervain,
verveine & trois feuilles, verveine citronnelle,.
verveine du Pérou, verveine odorante, yerba de la
princesa, yerba luisa

Aloysia virgata (Rufz & Pav.) A. L. Juss. -- cedron,
chicharra caopi, nino-urupé

Aloysia sp. -- sacha-poleo

Amasonia L. f. -- Amasonie, taligalées

Amasonia campestris (Aubl.) Moldenke -- aufrechte Amasonie,
herva de picapdéo, rothe Amasonie

Amasonia lasiocaulos Mart. & Schau. -- bandiera do Espirito
Santo, pau vermelho

Avicennia L. -- black mangrove, Lebendiggebarend, mangrove,

- Salzbaum

ers. PoE Ya Ob 0: Gsb ok Vol. 2, now 35

Avicennia africana P. Beauv. -- afia-nunung, amu-ati, amu-
tsi, angma-tsho, asokoro, asokpolo, asopro, asukuru,
ata-nunung, black mangrove, boandjo, boanjo, bue, bu®,
bue-dinte, bu%dint&, buwe, conmon white mangrove, 6-
bure, ede, ehrodo, garigari, gbeleti,. gb81l8ti, grigri,
jaia-guli, jaia-guwi, ka-bure, lagoon tree, mbougand,
mofuri, mutuku-tsho, nja-wului, odonumon, ogbun,
roanjo, saanar, samar, sana, tarrafe, tra-tsho, ufiri,
waterside tree, white mangrove

Avicennia alba Blume -- api-api, api-api hitam, black api-
api, elava, gundu mada, lamet, marne, samair dam, ton
samair, unte unte

Avicennia bicolor Standl. -- mangle negro

Avicennia -lanata Ridl. -- api-api bérbulu, api-api puteh,
hairy api-api, white api-api

Avicennia marina (Forsk.) Vierh. -- api-api, api-épi, api-
api merah, api-api puteh, api~api putik, bina, boak,
bufigdlan, burigél on , bungélu, _bufigdlun, cherid, fika-
fika, harav, kalapini, kalapini mafigitit, kalapini-
maput{, kausia, koak, kuldsi, kuydpi, lame apyus
lingig, lingog, mabaran , mada-chettu, mchu, migpi,
mtschu, nalla-mada, pidpi, piksik, pipisig, pipisik;
red api-api, renggou, sagarai, samair kao, showarab,
timmer, tioes 1éwo, upputti, white api-api, white
man grove

Avicenni& marina var. resinifera (Forst.) Bakh. -- manawa,
mangrove, native mangrove, New Zealand mangrove

Avicennia marina var. Rumphiana (H. Hallier) Bakh. -- api-
api, bunalun-babay

Avicennia nitida Jacq. -- algarrobo, black mangrove, black-
mangrove, black tree, blacktree, black wood, blackwood,
bois de méche, button mangrove, , carnoé, cativo mangle,
ciriuba, conrida, courida, cowrida, culumate,
glanzender Salzbaum, green turtle bough, honey-
mangrove, iguanero, istatén, mangel, manggel blanko,
mangle, mangle blanc, mangle blanco, mangle bobo,
mangle chéne, mangle negro, mangle prieto, mangle
Salado, manglier noir, manglo salado, mangrove, mangue
amarello, man gue branco, olive mangrove , paléluvier,
palétuvier, palétuvier blanc, pal étuvier gris,
palstuvier rouge, palo de sal, palo do sal, pariva,
parwa, parwaboom, péré, puyeque, saltbushes, salt pond,
white Brazil mangrove, white mangrove, witte mangrove

Avicennia officinalis L. -- api-dpi, api-api brajoe, api-

api daun lebar, api-api katjang, api-api ludat, api-api
puteh, baen, bani, bara baen, bien, bina, black mada,
bogém, bunalun-lalaque, bufgdlon, kajoe k&ndéka, kajoe
ting, ki balanak, lame, lameb, lamet, ludat, mada,
metbin, midpi, nala-mada, nalla, nalla mada, oepata,

:
1% 2 Pt el as
BEES Toe ON

~~

1944 Moldenke, Common Names oF)

orei, palétuvier, pé-apé, pidpi, saladillo, tamelh&é,
thamé, tiabaen, tivar, udat, white api-api, white
mangrove, zoutboom

Avicennia Schaueriana Stapf & Leechman -- caju, ciriba
preta, fromarina, magae Siriba, mangue, mangue brafico,
mangue seriva, siriuwa, white mangrove

Avicennia Tonduzii Moldenke -- palo de sal

Bouchea Cham. -- gervao

Bouchea fluminensis (Vell.) Moldenke -- gervad de folha
grande, gervao de folha grande, gervao de folha larga,
verveine faux-gervad

Bouchea prismatica (L.) Kuntze -- germander-leaved bastard
vervain, narrow-fruited vervain, prismatischer
BWisenhart, verbena, verbena cimarrona

Bouchea prismatica var. longirostra Grenz. -- arrocillo,
verbena, verbena manza

Bouchea pseudochascanum (Walp.) Grenz. -- gervad, gervao

Bouchea Rusbyi Moldenke -- verbena de flos grande

Burroughsia fastigiata (T. S. Brandeg.) Moldenke -- damiana

Callicarpa L. -- beauty-berries, beautyberry, beauty-berry,
callicarpe, French-mulberry, murasaki, Schonbeere,
Schonfrucht, Spanish-mulberry, Wirbelbeere,
Wirtelbeere

Callicarpa acuminata H.B.eK. -- albocar, blackberry,
ceniciento, flor de chichalaque, fructa de chacha,
patzahumacachil, pukil, pukin, sac pukim, uvilla, vara
de alcalde, vara del alcalde, x puc yim, zacpukim

Callicarra americana L. -- American beautyberry, American
callicarpa, amerikanische Schonbeere, amerikanische
Wirbelbeere, beauty-berry, beauty-fruit, Bermuda
mulberry, Bermudian mulberry, bunchberry, commode
mulberry, filigrana, filigrana de mazorca, filigrana
fructo morado, filigrana morada, French mulberry,
French-mulberry, Mexican-mulberry, sourbush, Spanish-
mulberry, turkeyberry, turkey-berry

Callicarps americana var. lactea F. J. Muller -- French
mulberry, white-fruited callicarpa

Callicarpa ampla Schau. -- capa rosa, capa rosa

Callicarpa angusta Schau. -- dirik-dirik

Callicarpa apoénsis Elm. -- layauvan

Callicarpa arborea Roxb.. -- bogodi, bormala, boropatri,
bundun, daung-sat-pya, dera, doika, doung-sap-pya, dum
kotokoi, ghivala, ghiwala, goehlo, gogdi, khoja, kodo,
kozo, makanchi, sakrela, stinga, turmong

Callicarpa basilanensis Merr. -- linagop

Callicarpa cana L. -- adokk, alalui, alayo-ti-manok,
anobrang, | anuyup, apoe~apoe, apu-apu, arusha, damar
b8si, dynamite-grass, goro-goro oetan, graue
Wirbelbeere, hai ngan, hati-hati ketan, katoempang

94 PHYTOLOGIA Vol. 2, no. 3

badak, katoempang kajoe, katumpang badak, katumpang
kayu, kuping bési, lo hai ngan, m@niran bésar, m¥niran
kasar, m@niran k&bo, mSniran oetan, mSniran utan,
palis, papalsis, red-fruited tampang b¥si, sai fa min,
Besepo, s&tampo bési, songka oetan, songka utan,
tambaldsi, tambul-basi, tampa bési, tampah b¥si, tampal
b&si, tampang bési, tampang b’si merah, tampong bSsi
puteh, tapoeng-tapoeng, tigau, tigau-na-itin, toembar
b¥si, tubang-daldg
Callicarpa caudata Maxim. -- anayop, anigup, arayop,
‘harayhai, kabatiti, mama, suba
Callicarpa cubensis Urb. -- filigrana de mazorquilla
Callicarpa denticulata Merr. -- anaif, mayop
Callicarpa. dichotoma (Lour.) K. Koch -- Chinese beautyberry,
French-mulberry , Japanese-mulberry, ko-murasaki, meka-
sogi, murasaki-sikiboo, purple-mulberry, purple urn-
fruit tree, sai ip lo hai ngan, tsu-kda-udn
Callicarpa’ elegans Hayek -- tambalabdsi
Callicarpa erioclona Schau. -- ka gongs palis, salingérau,
tambalabasi, tigau, tiibang-daldg
Callicarpa ferruginea Sw. -- filigrana, rostfarbene
Schonbeere, turkey berry ~
Callicarpe formosana Rolfe -- anadhiu, anoyop, anoyot,
atolba, for chai tsai, horai-murasaki , palis, sai hong
hun , shan-puchiang, tambalabdsi, tfagau, tigau, tigeu-
tigau, tigbabasi, timbabési, ts'u-k'ang, tubang-dalag,
tubang-daldg, tubaybdsi
Callicarpa Hitchcockii Millsp. -- boar-hog bush
Callicarpa japonica japonica Thunb. -- guiou-sao-si, Jama-murasaki ,
Japanese beautyberry , ko-mourassaki, méka-sogui, mi-
mura-saki, mourassaki-skibou, murasaki, murasaki- .
shikibu, tama-mourassaki, tama-murasaki, yabu-murasaki
Callicarpa longifolia Lam. -- bSb&tih kinana, b&8ning-b@ning,
chapal, chapal kechil, dama b&soi, gambiran, kajoe
séran, karat b&si, katoempang, katumpang, keling-kahan,
kém@niran, khow tok, lo kop ngan, m@niran oetan,
mSniran sapi, nasi-nasi, papalsin, sétampo,
simadgimbadjon, songka, songka kampong, sulap, tama,
tampah b&si, tampal b¥si, tampang b¥si, tampang b&si
puteh, tampoh (bési, tampong b¥si, tapah b¥si, tibabdsi,
tigau, tobaybdsi, tulang besi, white-fruited tampang
besi :
Callicarpa longissima (Hemsl.) Merr. -- bok wat tan
Callicarpa Loureiri Hook. & Arn. -- birodonmurasaki
Callicarpa macrophylla Vahl -- bd&-pattra, bauna, budhi-
ghasit, daya, den, druss, mathara, mattranja,
pattharman, poko kwat tan, shiwali, sumali, thar,
tondi-teregam, urn-fruit tree
Callicarpa magne Schau. -- atimla, eres

1944 Moldenke, Common Names 95

Callicarpa magnifolia Merr. -- agnai

Callicarpa Maingayi King & Gamble -- balek angin laut,
chulak, m&ndapor, tampang b&si, tulo, tutok puteh,
tutor .

Callicarpa megalantha Merr. -- palayan

Callicarpa Merrillii Moldenke -- katonal, palis, tigau

Callicarpa mollis Sieb. & Zucc. -- jabumurasaki, namainoki,
yabu-murasaki, yama~murasaki

Callicarpa nudiflora Hook. & Arn. -- tai chung lo kop muk

Gallicarpa obtusifolia Merr. -- anoyop

Callicarpa pedunculata R. Br. -- béning-b&ning rih.
mSm@niran, m&niran, ringan-ringan, wild heliotrope

Callicarpa reticulata Sw. -- netzblattige Schonbeere

Callicarpa Roigii Britton -- filigrana de pinar, filigrana
fruto blanco :

Callicarpa rubella Lindl. -- chaak tsai shue, sugriimik

Callicarpa stenophylia Merr. -- karangiti, lLayop

Callicarpa subintegra var. parva Merr. -- marataringau

Callicarpa suripsénsis Merr. -- alingtutungau, buyaken

Callicarpa tomentosa L.) Murr. -- aisar, ambong-ambong
bukit, ambong-ambong puteh, bastra coat comul, hu kwai,
kata kéran, k®payang, massandari, sitapoeeng,
sitapueng, tamah kérbau, t&pong-t&pong, teregam,
tindjaoe, tinjau, tondi

Caryopteris Bunge -- Bartblume, bluebeard, caryopteére,
verbena~shrub

Caryopteris incana (Thunb.) Miq. -- blue spiraea, blue

_ spirea, Chinese beardwort, verbena shrub

Caryopteris odorata (Hamilton) B. L. Robinson -- malet,
mohani, moni, shechin

Castelia cuneato-ovata Cav. -- cuchipapa, papilla, vara de
San Jose

Chascanum marrubiifolium Fenzl -- danabén, erg el bugr,
tchingaraguen

Citharexylum B. Juss. -- bois & cotelettes, bois cotelet,
bois de guitare, bois fidéle, citarexilon, cotelet,
cotelets, fiddle wood, fiddle-wood, fiddlewood,
fiddlewood tree, fidelle-wood, fiolintraee, fioltraed,
Geigenholz, Geigenholzbaum, gigatraed, gigetraee,
guitar wood, le bois cotelet, le bois de guitard, le
cotelet, Leierholz, susanna, vedelhoutboom, vedel
houthboom, zither-wood

Citharexylum affine D. Don -- alacate, cacachila, canutillo,
chachalaca, jalacate

Citharexylum Berlandieri B. L. Robinson -~- negrito,orcujuela

Citharexylum brachyanthum (A. Gray) A..Gray -- chile pdjaro

Citharexylum caudatum L. -- bird-seed, cateicillo, cigua,
coffe marron, collarette, dama, fiddle-wood, fidelle-
wood, guairo sando de costa, guairo santo, higuerillo,

96 PHY DOL OG Vol. 2, now 3

juniper-berry, long-spiked fiddle-wood, manglillo, moco
de pavo, palo de dama, penda, pendola de sierra, perda,
pigeon-feed, sauge doncella, white fiddlewood, white
fiddle-wood, wild-cherry

Citharexylum Cooperi Standl. -- corrimiente, wild-lime

Citharexylum Dawei Moldenke -- agracejo

Citharexylum decorum Moldenke -- totumillo

Citharexylum discolor Turcez. -- guayo

Citharexylum Donnell-Smithii Greenm. -- buela noche, buena
noche, chuul, cola de pava, coralillo, coral negro,
cordoncillo, cuul, dama, damas, moca de pava, paraiso,
sorguillo

Citharexylum ellipticum Sessé & Moc. -- anacahueta

Citharexylum Endlichii Moldenke -- manzano del cerro

Citharexylum flexuosum (Rufz & Pav.) D. Don -- turucasa

Citharexylum fruticosum L. -- agracejo, balsamo, balsamo,
black fiddlewood, bois de guitard, bois de guitare,
bois guitarin, canilla de vendado, catcycillo,
cateycillo, cutlet, fairytree, falo blanco, fiddlewood,
fiddle-wood, fiddlewood-tree, gallito, grenad marron,
guairo sando de costa, guairo santo, guayo, guayo
blanco, guayo roble, higuerillo, long Tom, mangle de
sabana, old woman's bitter, old-woman's bitter, palo de
guitarra, palo guitana, palo guitarra, penda, penda
blanca, pender, péndola, pendu, pendula, pendila,
péndulo colorado, pfndula [error for "pendula"],
pindoula, pindula, racemose fiddlewood, roble amarillo,
roble de olor, roble dulce, roble guayo, sangre de |
doncella, savanna-wattle, spicate fiddlewood, susanna
tree, white fiddlewood

Citharexylum fruticosum var. Brittonii Moldenke -- bois
cotelette, bois cutlet, cotelette, cutlet, fiddlewood,
hairy cutlet, white fiddlewood

Citharexylum fruticosum var. subserratum (Sw.) Moldenke --
cotelet denticulé, grenardo, palo santo |

Citharexylum fruticosum var. subvillosum Moldenke:--

cateycillo, gallito, penda, pendula blanca

Citharexylum fruticosum var. villosum (Jacq.) 0. 8. Schulz
-- bois cotelette, cotelet velu, cutlet, fiddlewood,
grenard, pende, white fiddlewood

Citharexylum Herrerae Mansf. -- huairuru

Citharexylum hexangulare Greenm. -- cajjalaco, canahuite,
palomillo

Citharexylum Hintoni Moldenke -- chichalaco

Citharexylum hirtellum Standl. -- sac-xitch-che

Citharexylum Kerberi Greenm. -- aceitumillo, aceitunillo

Citharexylum Kunthianum Moldenke -- cotelet tomenteux, palo
blanco

Citharexylum laetum Hieron. -- caffecillo, coffee chocolate,

)

1944 Moldenke, Common Names 7

jacende, semina, tarumd branco
Citharexylum lucidum Schlecht. & Cham. -- naranjillo, tepesi
Citharexylum macradenium macradenium Greenm. -- damas, danna
Citharexylum macrophyllum Poir. -- cotelet & grandes

feuilles, kasaroballi, leja gado, oeroejatoe

Citharexylum microphyllum (P. DC.) 0. ©. Schulz -- gatigal,
mala-muger

Citharexylum montevidense (Spreng.) Moldenke -- aguay~guazu,

espina de banado, naraujillo, tarumd, tarumd espirudo,
tarumén

Citharexylum myrianthum Cham. -- cag voro, carvoeiro, fruta
de macaco, pau de viola, pimenteira, pirazu rembiu,
primenteira, sarrid, turuman

Citharexylum pentandrum Vent. -- bois de guiterre, cotelet &
cing étamines, cotelet & feuilles molles

Citharexylum pernambucense Moldenke -- salgueiro

Citharexylum Poeppigii Walp. -- mullu-caspi, palo de
chaquiras

Citharexylum quitense Spreng. -- muyuyu, muy uyu del monté

Citharexylum Rosei Greenm. -- del ciervo a Sn. Juan

Citharexylum scabrum Sessé & Moc. -- jito siropo, panothillo

Citharexylum Schottii Greenm. -- chacni-bach, iximche,
ixtatakche, palo de violfn, tatakche, ahiobenches yerba

Citharexylum spinosum L. -- arbol de Santa Maria, bois carré,
bois cotelet, bois cotelet, bois cotelet carré, bois
cotelette, bois de cotelette, bois de fer blanc, bois
de guitare, bois fidele, bois fidéle, bois guitare,
bois guitarin, cotelette, cutlet, fairy, fiddlewood,
fiddle-wood, fig bush, guayo blanco, juniper-berry,
penda, savannah wattle, susanna, susanna tree, white
fiddle-wood

Citharexylum teclense Standl. -- café de érbol

Citharexylum tristachyum Turcz. -- agracejo, guayo blanco,
guayo roble, la calerio, mari de las Indias, palo
blanco, roble guayo

Citharexylum viride Moldenke -- corrimiente, corrimiento

Citharexylum sp. -- bois puitarin

Clerodendrum Burm. -- clérodendron, glorybower, glorybowers,

glory tree, Loosbaum, Losbaum, lotboom, pagoda-flower,
péragu, péragut

Clerodendrum aculeatum (L.) Schlecht. -- amourette, amnourette
de St. Cristophe, boesie droifi, boschhopfie, boton de
oro, chuc chuc, clavellina aspinosa, crab prickle,
crab=prickle, escambron blanco, gratte jambes,
Haugenush, madampolam, pree-bree, prickly myrtle,
privet, the bord de mer, wild coffee, wild-coffee,

- gamourette

Clerodendrum aculeatum var. gracile Griseb. & Moldenke --

clavellina espinosa

98 POH Yat O4,0 Gk Vol .°25° rigace

Clerodendrum adenophysum H. Hallier -- boerta-boerta, kajoe
boerta-boerta

Clerodendrum barba-felis H. Hallier -- badi, tcaucui

Clerodendrum Bethuneanum Low -- anoran, antutufigau-puld,
biniuang, guanton, kali-kali, maitiim, matdé&-kud, udan-
udan

Clerodendrum Blumeanum Schau. -- aoepaloslan mahina, kémbang

boegang, kolon ranteh, maroerang, mata ajam, panggil-
panggil, singoep, tadjoer, tintinga, waroe dojong

Clerodendrum brachyanthum Schau. -- haminddng, kayomkom,
lusib, mangha, samanpait, talabogting

Clerodendrum Buchholzii Gtirke -- bakoréné, fafa-hinei, fafe,
male, mosongo-songo
Clerodendrum Bungei Steud. -- camelia americana, chau shi

mut li, flor de la rosa muerte, fragrant clerodendron,
rose glorybower, sa sha ping

Clerodendrum calamitosum L. -- Baum des Slendes, kajoe
gambir, k&mbang boegang, klimmerlicher Losbaum, rampige
totboom

Clerodendrum canescens Wall. -- kwai. tim foh

Clerodendrum capitatum (Willd.) Schum. & Thonn. -- agbul u
uwagh,, ayeti, feremdmi, firi-fore, fuem8mi, furu-fure,
illiri, iye, korlejiga, mashayi, ny&@kpe, obranmotuan,
pfufulla, pipe-tree, taasen dua, tab&to, tataba, tr&men

Clerodendrum Colebrookianum Walp. -- kadungbi
Clerodendrum cubense Schau. -- hiel de gallina, magitire
cimarrona

Clerodendrum Cumingianum Schau. -- salumget

Clerodendrum cyrtophyllum Turcz. -- tzou tsing tsoi

Clerodendrum deflexum Wall. -- baboon's fat, balong ayam,
big wood blumea, black ixora, buffalo's tongue, cheret
hutan, chuchohgambar, cock's comb, decline wood, hill
haunted plant, kayu sampu, kayu sampu kSlau, kelusam
jantan, 14mak bérok, lidah kérbau, lidah k&rbau b&tina,
mérambong bukit, p&chah p¥riok hitam, s&kacha lima
jantan, s&kati lima, s&#kati lima jantan, s¥mbong hutan ~
-jantan, sétawar hutan, woodland s&tawar

Clerodendrum disparifolium Blume -- anting-anting,
chél8guri, chinaguri, guriam, kecholan, lampin badak,
lampin budak, 18lampang badak, p&ncholam, puding,
quilau, s&18guri, s@l&guri bétina, s%lulang bukit,
sSmbong, s*mpayan pitu, tampan put@ri jantan, tudong
roman, ubat tumboh, uloh-ulai, unting-unting

Clerodendrum formicarum Glirke -- taasen dua

Clerodendrum fortunatum L. -- fortuné, gelukkige boom,
gelukkige lotboom, glilicklicher Losbaum, Gliicksbaum

Clerodendrum fragrans (Vent.) R. Br. -- exhilaration tree,

glory tree, higantong, Javanese jasmine, m¥lor jawa,
mil flores, nang yam, pak yat pak, pelegrina,

: 1944 Moldenke, Common Names | 99

peregrino, pokok rabu k&mbang, rompok, sabuka, s¥tumpok -
Clerodendrum fragrans var. pleniflorum Schau. -- ela de
angel, boca amelia, bocamelia, boramelo, bridal-
bouquet, camellia, chau pin tung, cologne-plant, flor de
muerte, flor de muerto, Goehagan bush, herbe a Mad.
Villaret, herbe puante, hortensia, japana, jasmin,
jasmin de Amelia, jasmin de Italia, jasmin del muerto,
jasmin de] muerto de perro, jasmin del perro, jasmin de
muerto, jasmin de perro, jasmin hediondo, jazmin de
Amelia, jazmin de "spana, jazmin de [talia, karu,
madam-polan, madan=-polan, marabella, metrocedar, mil
flor, mil flores, milki hoedoe, misteriosa olorosa,
Nassau-rose, Nessau-rose, odorous clerodendrum,
Spanish-jasmine, verbena, vinda alegre, viuda alegre,
vulcana, wild jessamine
Clerodendrum glabrum E. Mey. -- bush clerodendr m, peo de
perico
Clerodendrum heterophyllum (Poir.) R. Br. -- bois cabri,
bois cabril, bois cabris, bois chenilles, bois de bouc,
bois de chenilles, gros bois de chenilles
Clerodendrum indicum (L.) Kuntze -- arnah, arnf, bd&man-hati,
baminhetti, baranai, bead-flower, bharangi, bhdrangi,
bhérgi, bidoejoek, brahmen-patta, brahma yashtika,
brahmunf, brahmunu yushtiki, chingari, daoen apioen,
daun apium, daw-di-mubarik, gandja, ganja, ganja-ganja,
géndje, memadatan, m@madatan, naijamphé ti, p¥natoh,
ronggo dine; sarum cutur, s&kar petak, siphonanthus,
tow yai-mon, tube flower, Turk's-head, Turk's-turban
Clerodendrum inerme (L.) Gaertn. -- afg-angri, ariya, |
baliseng, baén-jai, ban-juen, batraj, biring djéné, bon-
joi, bunga pawang, bun-join, bun-jumat, busel- pieeks
chia bam, eru-pichecha, eru puchcha, eti pisinika, foo
long shue, gambir laoet, goo yis hai, gulinda,
isamdhér{, kakoli, k¥mbang boegang, kémbang lygang,
ketoewer, koi a koi, koli, kundah, lagoendi alas, 1én-
jai, limau 1@lang, mangotngot, manoeroe dowongi, manor
oetan, m#lati oetan, m¥lati utan, nalla~kupi, nir-
notsjil, parian solojon, penni ka, pinari, ping-
shengam-kuppi , pirolai. kyont, pisangi, pishinika,
pisingha, samin-afga, sang-kupi, sd&ng-ktpi, oueroe
kupi, ‘shangam-kuppi , shengan-kuppi, sorcerer's flower,
tabangofigong, tak-kdélapu-chettu, t®rong gambul, tulang-
tulang, Ubi chettu, vana-jai, verevere, wal- guranda,
wal-gurenda, wiri salo
Pierodendrum infortunatum L. -- parni, bhandfra, bhéndira,
bhant, bhantaka, bhanti, bhat, bockada, bujiphyv,
chitu, gas-pinna, ghentu, infortuné, ka~aungeyl, kf if
. basuiti, kari, kari, kdung, khaoung gyf, kharbari,
kulamarsal, lukunah,. ongelukkige boom, ongelukkige

100 FY. Ts Ol OnGe ek Vol. 2, noe 3

lotboom, peragu, péragu infortuné, piene, pinne gala,
ungliicklicher Losbaum, Unglticksbaum, varni

Clerodendrum intermedium Cham. -- aloksok, asuangai,

balantana, bantana, dagtung, human g » iginga, kalalauan,

kasopénigil, kasupangil, katufigétun , kolokolog, laroan-
anito, libintano, pakapis, salinguak

Clerodendrum japonicum (Thunb. ) Sweet -- wan hon na wan

njari

Clerodendrum Kaempferi (Jacq-) Sieb. -- pak yat hung,
s€@panggil hutan

Clerodendrum Klemmei Slim. -- luag

Clerodendrum laciniatum Balf. f. -- bois cabri, nasty tree

Clerodendrum lanuginosum Blume -- antutungau-taluk,
magalablab, salumpapait, takipan, tanogo

Clerodendrum ligustrinum (Jacq. ) R. Br. -- itzimte, mosté,
muste, snake-tree, y'imte

Clerodendrum ligustrinum var. nicaraguense Moldenke --
jasmin, si me miras

Clerodendrum Lindenianum A. Rich. -- roble guayo, turquesa

Clerodendrum macrostegium Schau. -- agboligan, aktolfgan,
bagauak, baugak, kas opaéngil-na-puti, magboligan,
nakboligan, payi-payi

Clerodendrum minahassae Teijsm. & Binn. -- amamboligan,
ambuligan, ayam~ay am, bagalbak, bagduak, bagduak-itim,
bagéuak-na-puti, bakébak, boenato, danata, kasopangil-
gubat, ku-ku, 1¥i1%m in asoe, 1#%i1%m in taloen,
sunkol, tabiigok

Clerodendrum mindorense Merr. -- bagab, bagduak
Clerodendrum multibracteatum Merr. -- palutan
Clerodendrum myricoides (Hochst.) R. Br. -- surbattri
Clerodendrum nutans Wall. -- canastiilé, fire-bush,

martinica, misteriosa, ramo de novias, Santa Alda

Clerodendrum paniculatum L. -- baéh zitang, bunga mara,
bunga tinggal, danger flower, nom sawan, pagoda-
flower, pangil-pangil, p&manggil, p&ngkilai, p¥pangil.
pingdang, sapanggil, s€pangil, tabut

Clerodendrum phiomoides |. f. -- airan, airanamula, arni,
gharayt, irun, nellie, panjot, pirun, taludala,
taludalel, tekkali, telaki, teleki, tilaka, urni,
vato-ghni, wada madichi

Clerodenarum phyllomega Steud. -- kojoe lampam, ramo daging
lalaki

Clerodendrum Picardae Urb. -- jéte bois pin

Clerodendrum Pittieri Moldenke -- espino
Clerodéndrum polycephalum J. G. Baker -- apord, taasen dua

Clerodendrum puberulum Merr. -- urang-urang
Clerodendrum quadriloculare (Blanco) Merr. -- bagéuak,

bagsuak-na-pula, baligtaérin, baliktaran, salinguak,
uak~udk

mw Ae Se

ar

1944 Moldenke, Common Names 101

Clerodendrum Rumphianum De Vriese -- acepaloetan hahoela,
. pboenga panggil, boenga pluim, boenga poean, loloet
Clerodendrum serratum (L. Moon -- angar, baikyo, baranai,

barbara, bebya, bharang, bhérange-mula, bharangi,
bharangi, harungi, brah-mari mari, cheru tékka, chiru
dekku, chia, gandu-bhdrangi, gant-bahdrangt, gantu=
bhérangi, gunti paringsie, jeru-hka, kanta-bharanni,
kenhenda, ken-henda, kertasé, lampin budak, mata
_k¥sang, népélu, nirisa, pinggir.tosék, sagoenggoe,
saram lutur, s@nggoegoe, sénggugor, shimtek, simar
baoengkoedoe, singgoegoe, srigoenggoe, sunga tasek,
taman tasek, tambun tasek, ténjal tasek, timba tasek,
tindjaoe handak, tinjal tasek, pet cesta vatham
addakki

Clerodendrum serratum var. Wallichii ©. B. Clarke -- nirisa

Clerodendrum speciosissimum Van Geert -- coral, glorybower,
herba a Madam Villaret, herbe a Mad. Villaret grand,
jazmin rojizo-corazon, red honeysuckle, Santo Domingo,
scarlet clerodendrum

Clerodendrum spinosum (L.) Spreng. -- chichara, quédec

Clerodendrum splendens G. Don -- adabi, afifia omya,
ek&nyieya, geakoi, opé-éshi, ra-bina, ufuchi, widow of
last year

Clerodendrum ternifolium H.BeK. -- puta de noche

Clerodendrum Thomsonae Balf. f. -- ala de angel, balao de S.
Jose, bleeding heart, bleeding-heart, brinco de danea,
broedae nahatti, clara lisa, clematida, corazon horido,
Cornell-flower, crendolinda, egwa, enredadera flori-
rosada, fucsia, jamaiquina, lazo de amor, mata vaine,
misteriosa, Mrs. Thompson's clerodendrum, oviakuku,
pasion de Cristo, posidn de Cristo, secreto de amor,
southern bleeding-heart

Clerodendrum Thomsonae var. delectum Hort. -- lagrima de
Cristo

Clerodendrum tomentosum (Vent.) R. Br. -- Oumberland-tree,
downy clerodendrum

Clerodendrum trichotomum Thunb. -- harlequin glorybower,
kusagi, tungau

Clerodendrum umbellatum Poir. -- firi-fore, furu-fure,
honawai, hwana wulie, kajie, ti dah

Clerodendrum umbellatum var. speciosum (Dombrain) Moldenke
-- bleeding heart

Clerodendrum umbratile King & Gamble -- 1éruntoh, mali-mali
bukit, m#royan kabut, pianggu
Clerodendrum villosum Blume -- boerta boerta, buffalo's

foot-print, chapa, chapaneng, chémp@ning, daun bubut,
gampir roesa, kalopang pait, kasap jantan, labu-labu,
1&mpong hutan, milki hoedoe, p&chah p¥riok babi, pigs
ixora, pokok kasap, rough plant, tapak ké@rbau, zottiger

102 P BE CT DR OrG TF se Vol. 2,5 noe 3

Los baum

Clerodendrum violaceum Glirke -- ish&-dtin, work is sweet

Clerodendrum viscosum var. nilagiricum H. Hallier -- hukre-
mara

Clerodendrum volubile P. Beauv. -- dagba, &benote

Clerodendrum Williamsii lm. -- dibalai

Congea tomentosa Roxb. -- enredadera santa hoji-rojiza,
japonesa, ka-yan,. tamakanwe

Cornutia Plum. -- agnantes, agnanthe

Cornutia grandifolia (Schlecht. & Cham.) Schau. -- azari,
cuatro caras, cucaracho, palo cuedrado, zopilote

Cornutia grandifolia var. intermedia Moldenke -- cucaracho,
flor lila, hoja de jope, morcielago, murciélago

Gornutia grandifolia var. normalis (Kuntze ) Moldenke --
cuatro caras, morcielago, murciélago, palo cuadrado

Cornutia prandifolia var. guadrangularis fgrst. & Moldenke --
pavilla

Cornutia latifolia (H.B.K.) Moldenke -- chialche, loth-ché,
matasano, tzultesnuk

Cornutia obovata Urb. -- nigua, palo de nigua

Cornutia odorata (Poepp. & Endl. ) Poepp. -- dona, tal, ulape

Cornutia pyremidata L. -- agnanthe 4 fleurs en grappe,
agnanthe & fleurs en grappes, bois a cotelettes, bois
cac, bois cagne, bois care, bois cassau, bois cassave,
bois cdtelet, bois cotelet quarré, bois de caque, bois
de l'ancre, bois de saban, bois de savane, bois de
savanne, bois guarri, fiddlewood, mouri debout, penda,
purple fiddlewood, salvilla

Cornutia pyramidata var. isthmica Moldenke -- latche,
lattche, pangage, pangage, x oltexnuc

Diostea juncea (Gill. & Hook.) Miers -- cau-cau-mamill,
retama, retamilla, retamo

Diostes scoparia (Gill. & Hook.) Miers -- clavelillo del
campo, escobilla de] campo

Duranta L. -- golden-dewdrop, pigeonberry, skyflower, sky-

flower
Duranta argentea Lodd. -- silvery duranta i:
Durenta coriacea Hayek -- naranjuelo .

Duranta costaricensis (Donn. Sm.) Standl. -- ufia de gato

Duranta Mutisii L. f. -- limoncillo, long-fruited duranta,
Mutis's duranta

Duranta repens L. -- adonis, adonis blanco, adonis morado,
arisgo, azota-ceballo, blue plumeria, campo-koche,
celosa, celosa cimarrona, chulada, coralillo rosado,
cuenta de oro, cuento de oro, durancia, duranta,
duranta de Plumier, durante de Plumier, ®lis's
duranta, espina blanca, espina de paloma, espino
negro, fructea de jacu, fruta de jguana, fruta de
paloma, garbancillo, golden dewdrop, golden-dewdrop,

~—1944 Moldenke, Common Names 103

golden dewdrops, granjenillo, heliotrope bush,
heliotrope tree, heliotropio, heliotropio morado ,
hombocoche , kempok6-ché, kan poco che , kanpdko~ché ,
kanpo ko=ché, kanppocoche, kanppocoché, kanppocgché,
lila, limoncillo-.cimm, lluvia, Toray pensamiento,
.- pigeon berry, BEgoae herr ys Plumier's duranta,
skyflower, troene d! Amérique, unarmed duranta,
vanilier, vanilla, vanillier, varita de San José,
_villatermin, violeteira, violetina, Xalapa duranta,
xcambocoché, x kambocoche, yellow hat tree
Duranta repens var. alba (Masters) L. H. Bailey -- forget-
me-not, heliotropio blanco, nomeoluides, no-me-oluides,
varita de San José
Duranta revens var. canescens Moldenke -- fruta de paloma
Duranta serratifolia (Griseb.) Kuntze -- pala blanca, tala
- blanca, tala blanco
Duranta Skottsbergiana Moldenke -- judu casha
Duranta triacantha A. L. Juss. -- chisnan
Faradaya ovalifolia (A. Gray) Seem. -- ngakawa, wakarovungi,
wakorovundi, wa vatu
Faradaya vitiensis (A. Gray) Seem. -- wa vatu
Geunsia Cumingiana (Schau. ) Rolfe -- danasi, gagayug,
magilars malataba&ko, manabako, nago, sambuyut,

sobsoganbogo
Geunsia flavida (Elm. ) H. J. Lam -- layaupan, madolau,
~_ pananagok
Geunsia pentandra (Roxb.) Merr. -- layaupan
Ghinia Boxiana Moldenke -- cardero, coast broom
Ghinia curassavica (L.) Millsp. -- flor morada, spiny-
r fruited vervain
F Ghinia curassavica var. yucatanensis Moldenke -- chan-ko-
~ xnuk, chanxnuk
Gmelina L. -- gmelin, heilpeeren
Gmelina arborea Roxb. -- at-demmata, bolko bak, chimman,

cummi, gamari, gaméri, gambari, gambhar , gemhar ,
gomari, gumadi, gumar , gumar-tek, gumbar, gumbhar,
gumbhar , gumbhari, gumhar , gumhér , gumher , gumudu
gumudu téku, kdékddumbdéri, kembhar, kasamar, kasmar,
kdemari, kasmaryamu, kasmiri, kassamar , khamer ,
khambhéri , khammara , kall, kimér, kumbhdr, kumbulu,
kaimhér , kurse, kyunboc, kywon-pho, pedda gomru, pedda
gumudu téku, ramani, sag, sewan, shewan , shewney ,
shewun, shivan, shivani, shiwali, shiwun sripmari,
tagumida, tegegummadu, yemanai , y émené

Gmelina asiatica L. -- badhdéra, bhedaira, biddari, bulangan,
challa-gumidu,coumelon, gamudu, gatta-demmatta, gmelin
asiatique, udu, heilpeeren, kal-shivani, kavva-

< gumudu, 1a&han shivan, nilak-kumazh, nilak-kumizh
Gmelina asiatica var. villosa Bakh. -- boelangan, boewah

3
q

ee eo

104 Pe TO G-0.6.t & Vol. 2, %Gece

kerandjang, kananga woeba, kemandiang, loewarang,

| ponranga, sarogang salaki, waréng k¥tan, w8wenganga

Gmelina elliptica J. %. Sm. -- bangana, b¥longeh, bohol,
bulang, bulangan, bulang gajah, bulang hutan, bulang
k¥chil, bulbuol, dadiangas, danhafgas, kalufmgun, kang
mao, kemandiang, nom méo, puhtng, pukeng mata hari,
saonad, talauan, talvngud, talungun, tanlungun,
tulungun, tufigdinol, waréng, wareng k&tan

Gmelina Leichhardtii (F. Muell.) F. Muell. -- beech,
Queensland beech, white beech

Gmelina macrophylla Wall. -- kaju titi, kaju titie, kaju
tittie

Gmelina moluccana (Blume) Backer -- titi, toehoe, toeroe

Gmelina philippensis Cham. -- aliplng, alipufga, baga-babui,
betebet, bosel-bosel, bulangan duri, ching chai,
kalulut, kumbil, paniktik, sousou, tuldfgau

Gmelina racemosa (Lour.) Merr, -- shek tzi shu, song tsio.
gun °

Holmskioldia Retz. -- holmskioldia, holmskioldie

Holmskioldia sanguinea Retz. -- chapeau chinois, Chinese hat
plant, Chinese hat-plant, Chinese-hats, kapni, kul
tolia, kumaon, mandarin's-hat, misiwahchil, paraguita
de China, parasol-flower, rithoul, sombrero chino

Hosea Lobbii (C. B. Clarke) Rid1. -- d'dap mira

Junellia asparagoides (Gill. & Hook.) Moldenke -- tomillo

Junellia bryoides (R. A. Phil.) Moldenke -- culesoro, pata
de perdiz

Junellia Lorentzii (Niederlein) Moldenke -- matorro moro

Junellia seriphioides (Gill. & Hook.) Moldenke -- espina de
pescado, tomillo macho .

Junellia tridens (Lag.) Moldenke -- mata negro

Lachnostachys Cliftoni F. Muell. -- big flannel plant

Lampaya medicinalis R. A. Phil. -- lampaya, lampayo

Lantana L. -- bergsalbei, camara, ‘cambara, cambaras, capitaéo
do campo, cha de pedreste, lantana, lantane, Vehl baum,
Schwalke, shrub-verbenas, Wandelbltite, Wandelrose, wild
sage

Lantana aghyranthifolia Desf. -- cariaco de San Juan,
cariaquito blanco, frutilla blanca, frutillo

Lantana bahamensis Britton -- Bahama lantana, golden-rod
Lantana balsamifera Britton -- Inagua sage-bush
Lantana Camara L. -- adelamanyi, akotongm¥, amerikanischer

Mehistrauch, ananse d&Skono, ananse dua, ananu -komi,
ananu kin-tsho, angelmund, Bahama tea, bahig-bahug,
Boenga pagar, bohdé-bohd, cabard-cad, camara, camara,
camara & feuilles de melisse, camara piquant, cambard,
carrioquito, cinco negritos, common deep orange lantana,
corbeille d'or, coronitas, dame cubre galanos, der
surinamsche Thé, “nglish sage-bush, &w8n adé1é, &w’n

‘4

1944 Moidenke, Common Names 105

agogo, filigrana, flor de duenda, flor de sangre, flor

di sangur, galaba, gekroonde lantana, herbe 4 caiman,

herbe & plomb, hunter-does-not-eat-it, Jamaica mountain

sage, jaral, jarilla, kajoe singapore, kemantjo,
k¥mbang saték, k¥mbang talék, kiskeete, koorsoe

Wiwierie, koorsoe wiwiri, koorsoe wiwirie, koortsruid,

; korso-wirie, lantana, lantana & feuilles de mélisse,

i maintjo, Marie crabe, Mehlbaum, mora de caballa, oblo,
poejéngan, poetj®ngan, red sage, red sage-bush, sage,
Salbeystrauch, salijara, salijéré, sauge & feuilles
rondes, sauge de montagne, Soldaten Thee, soterre,
spider's kenki, surinamischer Thé, Surinam tea plant,
tahi ajam, tai hajam, tai kotok, tamandjho, tea plant,
tamb¥1ek, t¥mb#lékan, t¥t¥rapan, tjenté, tres colores,
venturosa, vieille-fille, viorne d'Amérique, waoeng,
wilde salbey, wild sage, wiléran, yellow sage, yemo
sigba, yerba de la muestranza, yerba mora

Lantana Camara var. aculeata (L.) Moldenke -- boenga-in tah,
bunga asam senyur, bunga pagar, bunga tahi anjing,
bunga tahi asu, bunga tahi ayam, bunga tahi ayam busok,
camara de espinho, camara espineux, cariaquillo,
chénté, comyon lantana, dog's dung flower, hedge
flower, kamkung, katu-hinguru, k%mbang satek, k¥mbang
telek, lantaine arguillonense, lantan, lantana, lilac
lantana, oblo, paka krawng, pink sage, prickly lantana,
prickly sage, puch&ngan, puySngan, saliyara, saliyéré,
stachelige Lantane, stachlige lantane, tahi ayam munai,
tai hayam, tai kotok, témbY1ék, t&mb#lékan, t#t¥rapan,
tokalau, waiwai, waung, wiléran

Lantana Camara var. flava (Medic.) Moldenke -- lantanna,
yellow lantana

Lantana Camara var. hybrida (Neubert) Moldenke -- dwarf
lantana, lantanna ;

Lantana Camara var. mista (L.) L- H. Bailey -- cinco
negritos, common lantana, @nglish sage bush, hairy
lantana, lantana, lyre, oema koorsoe wiwirie, solande,
West Indian coast bramble, wild sage

Lantana Camara var. mutabilis (Hook.) L. H. Bailey --
lantana, lilac lantana

Lantana Camara var. nivea (Vent.) L. H. Bailey -- camard de
flor branca, lantanna, white-flowered lantana, white
Lantana

Lantana Camara var. sanguinea (Medic.-) L. H. Bailey --
lantanna, red lantana

Lantana Camara var. varia (Kuntze) Moldenke -- harlequin
lantana, lantanna

Lantana Chamissonis (D. Dietr.) Benth. -- cambaré >

Lantana citrosa (Small) Moldenke -- hiervade javillas,
oregano xiu, sac-chili, toronjil.

co

“a

a 2

a

m.

4
*

106 PHYTOLOGIA Vole 2, noe 3

Lantana frutilla Moldenke ~~ frutilla

Lantana fucata Lindl. -- Brazilian lantana, camara roseo,
cariaco morado

Lantana glandulosissima Hayek -- cinco negritos, cinco-
negritos, cinco nigritos, confituria amarilla,
confiturilla amarilla, frutilla, guaquita, oregano
silvestre, oregano xiu, sincuria, tres colores, xo-
pec

Lantana hispida H.B.K. -- chinkuro, jaral, mora de caballo,
orozuz del pais, soterre blanco, toltolquelite

Lantana horrida H.B.eK. -- bunch-berry, calico bush, cinco-
a, confiturilla, hierba de Christo, lantana,
palabra-de-muger, yerba del Cristo

Lantana insularis Moldenke -- lantana

Lantana involucrata L. —- andornblattrige Lantane, baume de
la grande terre, big sage, button sage, camara &
feuilles obtuses, common sage-bush, filigrana, lantaine
involucrée, monjol, montjoli, montjoli de Cayenne,
sage, sage bush, sage tree, Santa Maria, Santa Maria de
playa, te de la playa, wild sage, zicilhaxiu

Lantana Langlassei Moldenke -- toronjil

Lantana macropoda Torr. -- mejorana

Lantana Mearnsii Moldenke -* ananse kono, ananu komi, ananu
kon-tsho, eleku, hunters! scent, hunters!’ spice, kimbar
mahalba, urdi loho'be

Lantana montevidensis (Spreng.) Briq. -- cabard-cad, camard,
filigrana, lantana, polecat-geranium, salvia morada,
trailing lantana, weeping lantana, weeping-lantana,
yeung ue yi fa

Lantana Moritziana Otto & Dietr. = cariaquito, flor de

sangre

Lantana ovatifolia Britton -- ovate-leaved lantana

Lantana rugulosa H.B.K. -- venturosa

Lantana scorta Moldenke -- frutilla, frutilla para comer,
jantana, yerba de tres colores

Lantana tiliaefolia Cham. -- cambard, common lilac lantana,
Violet King Lantana, yellow lantana

Lantana trifolia L. -- benturosa morada, bunga pagar puteh,

lantana, oregano, sweet sage, venturosa, yellow sage

Lantana urticaefolia Mill. -- drap d'or

Lantana velutina Mart. & Gal. -- confite, confite blanca,
confiturilla blanca, frutilla, oregano xiu, rosa
blanca, salvia alta, sarza mora, tolochocho,
tolonchocho

penten’ sp. -- bois de sauge, katu-hinguru

Lippia Houst. -- lippi, lippia, Lippie, 2itronenlippi

Lippia adoénsis Hochst. -- afurati, bahé, bahé-bahé, borom-
borom, diohuli, efinrin-gogara, fasau, fetfetti,
Gambia tea, gane ba, guilel guéri, kani ba, kimbo,

Ne A

3 1944 Moidenke, Common Names 107

kingkilli ba, mbalhat, mbormbor, ng&su, ng&suru, nydna,
saa-nunum, sisiling hy amo

Li ppiae affinis Schau. -- orégano, oregano di burro, poleo

Lippia alba (Hill. ) N. ©. Br. -- alfronbrilla, beukes bosjie,
blakka tiki ment, bushy lippia, cidrera, graveelkruidje,
hierba del negro, hierba negra, jueanilema, leppie rude,
malmequer do mato, malva, malva thee, oregano, palisado,
pampa oregano, pan poregano, poleo, poley, Saint Mario,
Balvia, salvia morada, sidraera, tarete

Lippia Berterii Spreng. -- orégano, orégano de burro, poleo

ue ppis Briquetii Moldenke -- amogre
Lippia callicarpaefolia H.B.K. -- salvia real
Lippia cardiostegia Benth. -- chiligua, orégano montés
while Geisseana Solered. -- orégano
Lippia graveolens HeB.oK. -- grégano, hierba dulce, oregano,
oregano, oregano cimarron, tabay, tarbay, té del pais,
xakilche
Lippia Grisebachiana Moldenke -- salvia morada
Lippia Hoehnei Moldenke -- atiaci
Lippia hypoleie Briqe -- cutujume, maste, tah
Lippia integrifolia (Griseb.) Hieron. -- manzanillo, poleo,
pulco, yerba de Inca
Li ppia micromera Schau. -- orégano, origanum, Spabish-thyme
Lippia micromera ware Helteri (Britton) Moldenke --mejorana,
oregano, orégano
Lippia myriocephala Schlecht. & Cham. -- tatascame, vera
blanca
Lippia oxyphyllaria (Donn. Sm.) Standl. -- caragra
Lippia Palmeri S. Wats. -- orégano, origano
Lippia Palmeri var. spicata Rose -- origaro
Lippia Pringlei Brig. —= bacaton, batayaqui, choila, chokili,
matayaki, tabaquilla, talabao, talakao
Lippia Pseudo-thea (A. St. Hil.) Schau. -- camara faux thé,
capitao do mato, cha de pedreste, faux thé, thé de
piéton
Lippia Recolletae Morong -- malvena
Li ippia a scabra Hochst. -- mu-tswane
Lippia substrigosa Turcz. -- salvia santa
Lippia Torresii Stand]. -- caragra, caragra negra, caragre
Lippia turbinata Griseb. -- poleo, té del pais
Lippia turbinata f. angustifolia Osten -- poleo

ippia umbellata Cav. -- droceria, drosira
ippia yucatana Loes. -- salvia poblana, xolténuuc

Nashia Panne Millsp. -- moujean tea

Neosparton ephedroides Griseb. -- chinquillo, pichanilla
retamo

Peronema canescens Jack -- djati sabrang, ki sabrang,
loeroes, soengkai, soengkai m¥lajoe

Petitia Jacq. -- black-fiddlewood, Petitie, petitier

108 PHYTOLOGIA ‘Vol. 2, noe a

Petitia domingensis Jacq. -- bastard stopper, bastard-
stopper, black-fiddlewood, black-heart fiddlewood, bois
de fredoche, bois d'ortie, bois pelé, bois sans écorce,
capé&, capa amarillo, capd& amarillo, capa-blanca, capa
blanco, capa& de sdbana, capdé sabanero, capa savannah,
chen a gren, chéne calebasic, chéne calebassic, chien a
gren, fiddlewood, fiddle-wood, fidéle, guayo, guayo
prieto, petitia, petitier de Saint-Domingue, roble
guayo, spur tree, westindische Petitie

Petraeovitex multiflora (J. @. Sm.) Merr. -- hahiat, seroe
wari, tali boeboe

Petrea Houst. -- pétrée, purplewreath, whitewreath

Petrea Andrei Moldenke -- chivovo gueb

Petrea arborea H.B.K. -- blue tree petrea, lilac, tosatido,
tostadito

Petrea arborea var. Broadwayi Moldenke -- bridal-wreath

Petrea aspera Turcz. -- bejuco de hajo, biura, flor de la
cruz, flor de mayo, Santa Lucia, tostadito, viuda

Petrea bracteata Steud. -- hajauballi saléroe, hayariballi,
parapo, petraea, sandpaper-vine

Petrea glandulosa Pittier -- penitente

Petrea Kohautiana Pres] -- bridal wreath, liane rude, lilas,
purple reef-plant, purplewreath, sandpaper-flowers,
tree petrea, white petrea

Petrea Kohautiang var. alba (Freeman & Williame ) Moldenke --
bridal-wreath

Petrea macrostachya Benth. -- moronea

Petrea peruviana Moldenke -- sanango sacha
Petrea pubescens Turcz. -- pluma
Petrea racemosa Nees -- flor de S. Miguel, flor de viuva, 1a

pétrée grimpante, purple wreath, purple-wreath, touca
de viuva, twining petraea, viuvinha

Petrea rugosa H.B.K. -- chaparillo, chaparrillo, mamoncillo

Petrea volubilis L. -- adelfa, adolfina, bejuco de caballo,
bejuco del caballo, buird, carbonera del monte,
chaparrito, chaparro, choreque, ci contre, coamecate
azul, colacion, estrella Bz s flor de Jesus, flor de
papel, flor de Santa Marfa, hoja chigue, jazmin, jazmin
azul, 18 pétrée grimpante, lengua de vaca, liane de Str
Jean, liane rude, liane Saint-Jean, opp-tzimin, potrea,
piocha viejo, purple wreath, purple-wreath, queen's-
wreath, raspa sombrero, sandpaper-vine, Santa Rita,
soltero, stapelia-flowered petrea, tortills tortada del
caballo, tortilla tostada del caballo, totopostillo,
twining petreea, yoch opp tzimin, yoxop-9imin

Petrea volubilis var. pubescens Moldenke -- chorreque, cuera
de zapo, flor de Jesus, raspa-guacal

Phyla Lour. -- fog-fruits, frog-fruit, frog-fruits

Phyla cuneifolia (Torr.) Greene -- vhapparal, Mexican

He a
EO a ee ee ee a ee ae eee meee ee, See

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e
“5
A

1944 Moldenke, Common Names | 109

heliotrope, wedgeleaf frog-fruit, wedge-leaved fog-
fruit
Phyla incisa Small -- fog-fruit, frog fruit, spatulate-
leaved fog-fruit, wedgeleaf frog fruit, weighty fog-
fruit
Phyla lanceolata (Michx.) Greene -- fog fruit, fog-fruit,
frog-fruit, spatulate-leaved fog-fruit
Phyla nodiflora (L.) Greene -- ana-coluppa, bhui-okra,
busbusi, cape-weed, chhota okra, cidron, creeping
lippia, fog fruit, fog-fruit, fox-fruit, fraise de mer,
frog fruit, Godet's-weed, godon kada, herimena-kola,
hierba de la Virgen Marfa, 1ién fuen, lopu-lopu, lopu-
lupu, naculad, nakulad, orozus, sarad buti, sirik puto,
sirik puyo, spatulate-leaved fog-fruit, spepetun,
sprain bush, te cimarrén, turkey tangle, verveine,
verveine du pays, verveine sauvage
Phyla nodiflora var. canescens (H.B.K.) Moldenke -- hierba
de hormiga
Phyla nodiflora var. reptans (H.B.K.) Moldenke -- buttonweed,
hierba de hormiga, larger creeping lippia, spepetun,
wedge-leaved fog-fruit
Phyla nodiflora var. rosea (D. Don) Moldenke -- yerba-de la
Vir gen
Phyla scaberrima (A. L. Juss.) Moldenke -- corronchocho,
hierba buena, hierba dulce, honey-herb, malba,
neuctixihuitl, orozul, orozus, orozuz, orozuz de
latierra, orozuz del pais, salvia santa, yerba dulce
Phyla stoechadifolia (L-) Small -- cabalyaxnic, cabal-
yaxnic, marsh lippia, té del pais, te negro, thé del
paiz, yerba Luisa Elena
Premna L. -- anderése, Bocksmtille, bokkeblad, premme

Premna acuminatissima Merr. -- sai ko din nuang
Premna adenosticta Schau. -- kalanggiduan, kalipdpa-madam,

kla, lanabau, lifigo-lifigo, magupai, mulduinaso, sasalit

Premna barbata Wall. -- bakar, bakharcha

Premna. bengalensis C. B. Clarke -- dhaoli, gabbu nelli,
"_ gohora, gwyheli, sungna

Prema congesta Merr. -- alakdag

Premna cordifolia Roxb. -- amboeng-amboeng lacet, baroewas,
baroew%h, baroh, b%roewas, boewas—-boewas, si baroeweh

Premna corymbosa (Burm. f.) Rottl. & Willd. -- agetha,
agnimantha, aguy abat, aloalo, andarése, appel, arbre a
la migraine, arbre de la migraine, arni, babon,
bakarcha, balabi, b&buas, b&buat, bmit-bhirari,
Bocksblatt, bois de bouc bois sureau sauvage,
bokkeblad, chah leud, chamdari,daoen kambina, ganiari,
ganikérike, ganniari, ghebu-nelli, gineri, goemira,
headache tree, indjaro, inrelo, xarnike, ki pahang,
ki seungit, middf-gass, muiney kfray, munnay, munni-vayr,

110 PH Y-T:0 2-0: GE.4 Vol « 2, nose

naérvel, pinne-nelli, sajor kambing, singkil, singkil
alas, toung-than-gyee, ustabunda

Premna corymbosa var. obtusifolia (R. Br.) Fletcher --
adgau, agau, pedau, alagau, alagéu-blanko, alagdu-
dagat, alalgéu, anobrang, aragéu, argdu, naga

Prema corymbosa var. sambucina (Wall.) Moldenke -- kaikoa,
~~. kuabalon, qarovo, remako, siba, talitue

Premna Cumingiana Schau. -- plageu, banaba, magalas,
magilig, magilik, maladpi, manabé, maparai, mulmagan,
palaudiduan

Prema divaricata Wall. -- akar buas tunggal, akar jutong,
akar mSlor padang, akar p&rindu, akar rachun tikus,
bois de bouc, buas-buas, field jasmine, lingue bJanc,
rat-poison climber, t®nung rimau puteh

Premna foetida Reinw. -- ambong-ambong laut, buas—buas,
‘ drekhout, karuana, kua, pokok buru hantu, remako, sibo,
varo
Prema Gaudichaudii Schau. -- ahgao, ngurunguru, waro, waro
ndamu

Premna hispida Benth. -- bilankuru fida, kaf¥i, kafi

Premna. Latifolia Roxb. -- chambadi , chambari, dauli, gineri,
" gondona, michapgong, middi, nella, nelli, nellf kira,
pedda-nella-kira, padda nellf kira

Premna latifolia var. cuneata C. B. Clarke -- dangra seya

Premna latifolia var. mucronata (Roxb.) C. B. Clarke --
agniu, agnium, bakar, bakarcha, bakhurchu, bankar,
basota, ganhile, ganhin, gfan, jhatela, tumari

Premna membranacea var. cordata Merr. -- nago

Prema. nauseosa Blanco -—- agrau, alagau-gubat, ananghit,
~ anghit, angsuen, ansuan, areu, mala~mulauin, muléuin-

aso

Premna octonervia Merr. & Metc. -- sai ko din nuang, tai ip
~ shan po

Premna odorata Blanco -- aagdu, abgdu, adgdu, adiyo, agbdu,

OP SUE OSE ¢

~ agdau, alagdu, anobran, argau, atingi, duragdu,
guachal, lagau, lassi, pumuhat, tangli, tibangiigen

Premna parasitica Blume -- areuj ki hoedjan, djati areuj

Premna quadrifolia Schum. & Thonn. -- dengS, gyengya
aforowa, ogboso-tsho, ogbosu

Premna stellata Merr. -- manaba

Premna subglabra Merr. -- adgdu, agdu, alagséu, alagéu-
blanko, aridu, salipdpa

Premna subscandens Merr. -- alagdu-baging, anangget,
anobrang-ng-limanut, sikir-fNg-purau, uradgau

Premna taitensis Schau. -- nici, tavotavo, warowaro, yaro

Premna taitensis var. rimatarensis F. H. Bre -- aloalo,
rauvula, tavu, yaro

Premna tomentosa Willd. -- b¥boelang handak, b&buas,
bird's nest, boelang, boengboelang, boenglang, bulang;

eta
‘

— 1944 Moldenke, Common Names 111

bungbulang, bunglang, gadoengan, gadungang, g&mboelang,
gémbulang, lajas-lajas, 1¥%ban chapo, 1¥ban tjapo,
oenit, piat, pisang-pisang, sarang burong, t¥mbaroh,
unit .

Premna trichostoma Miq. -- buas, m¥dang palu

Priva Adans. -- Drehling, Eisenhart, velvet bur

Priva adhaerans (Forsk.) Chiov. -- arabischer Bisenhart,

hamsched

Priva aspera H.B.K. -- chile gieco, chirrite, churrite,
pengua, salvia alta .

Priva cordifolia (L. f.) Druce -- obeera, scharfblattrige
Drehling, wotray cheddy

Priva cordifolia var. abyssinica (Jaub. & Spach) Moldenke --
nassak

Priva lappulacea (L.) Pers. -- amor seco, berbenilla, bur
vervain, bur-vervain, cadillito, cadillo, cadillo de
bolea, calluncay, carbroom, cat's-tongue, cayuncay,
cola de alacrdn, collant, costdén, globito, guanaboa,
guerit tout, heal-all, herbe 4 l'angine, klettenartiger
Eisenhart, mozote, mozote de gallina, mozote de pollo,
mozotillo, pedagoso, pegajosa, pega~pega, pega pollo,
Sonderahre, stick-tight, styptic bur, tzalluntzay,
velvet bur, velvet-bur, verbena, voekoe voekoe tolman,
xpakunpak, 2a] lunaay

Priva mexicana (L.) Pers. -~- mexikanischer Bisenhart,
verveine du Mexique

Priva rhinanthifolia (Mart. & Gal.) B. L. Robinson --

' pionillo

Pseudocarpidium ilicifolium (A. Rich.) Millsp. --
‘chicharron, navaja de verraco, pico de cotorra

Pseudocarpidium mitidens (Urb.) Moldenke -- chicharron

Pseudocarpidiun Wrightii Millsp. -- chicharron, Wright's
pseud ocarpidium

Pygmaeopremna herbacea (Roxb.) Moldenke -- bhuijam,
bhumijambu, bhimi-jambuka, huniyan, kada met, méla
niredu

Pygmaeopremna humilis Merr. -- huniyan

Rehdera penninervia Standl. & Moldenke -- palo blanco

Rehdera trinervis (Blake) Moldenke -- llayo, sacuisilche,

eee eee
saquilzciche

Rhaphithamnus Miers -- citarexilon

Rhaphithamnus spinosus (A. L. Juss.) Moldenke -~ amydn
macho, arayan de espino, arrayaén de espino, arrayén
espinudo, arraydén macho, espino, espino blanco espino
negro, -guayun, nayun, prickly-myrtle, repu, repu mayun

Rhaphithamnus venustus (R. A. Phil.) Be Le Robinson --
arayan macho, arrayén macho, espinillo, juan bueno

Sphenodesme barbata (Wall.) Schau. -- aga lumut, akar
chabane lima, akar lumut, akar m&ruan, five-points

aie POW Y.T 040-4. f° # Vole 2, no. 3

Climber, hutat, lLembu-lembu, lilimbo

Sphenodesme borneénsis Merr. -- sumpin

Sphenodesme pentandra Jack -- akar k&tu-k&tu, akar lintang
ruas, akar subang, akar tanak rimau, bunga kértas, ear-

_ stud climber, lentang ruas, paper flower, shan pak tang

Sphenodesme triflora Wight -- akar bisa, akar bisar, akar
katup-katup, akar m@mali, akar pinang gusi, akar risa,
akar sambu, akar s&mpuleh

Stachytarpheta Vahl -- bastard vervain, Dichta&hre, Eisenbart,
Eisenhart, Fettahre, gervao, ogervaéo

Stachytarpheta angustifolia (Mill.) Vahl -- chilillo,
esponjilla

Stachytarpheta australis Moldenke -- gervao, tinho

Stachytarpheta Calderonii Moldenke -- verbena

Stachytarpheta cayennensis (L. C. Rich.) Vahl -- akojoe
malakali, akojoe mala kali, burr vine, cola de millo,
corocillo, gerbdao, ervao, Java verbena, kaka -kankan,
kuka kankang, 1a che rat, large leaf, man kaka kakkan,
Nuenu-pichada, ojediballi, ojédiballi, snake-rattle,
ucullucui-sacha, ucullucuy sacha, verbena, verbena
ancha, verbene blanca, verbena falsa, verbena negra,
vervaine, water vine

Stachytarpheta elatior var. Jenmani Moldenke -- esponjilla

Stachytarpheta Frantzii Polak. -- cola de alacran, cola de
armado, mazote, verbena, verbena celeste, verbena

morada
Stachytarpheta fruticosa (Millsp.) B. L. Robinson -— Bahama
vervain

Stachytarpheta guatemalensis Moldenke -- camag o1al, San
Diego, verbena

Stachytarpheta indica (L.) Vahl -- brasilianischer Tee,
Brazilian tea, djarong lalaki, Tast Indian false
vervain, herbe & chenilles, jarbao, jarung lalaki,
ngadi réngga, orgibao, queue de rat, roemdjaroem,
sSlaseh dandi, s¥laseh hutan, spotted basil, thé du
Brésil, vervain, verveine bleue, verveine queue de
rat, woodland basil

Stachytarpheta jamaicensis (L.) Vahl -- abontennua, agba,
&agOgo igun, albaka, angkasa-angkasa, ankasa, bastard
vervain, berbena, bilu-bilu, biron, blue-flower,
bolomoros, Brazilian tea, Brazilian-tea, devil's coach
whip, djarong, g&wongan, ibinxiu, irt aldngba, irt
amure, Jamaica vervain, kaka kairkau, kaka kankan,
kandikandildan, karoménal, large leaf, limbagat,
lizard's tail, mes, mouse's bowstring, nagabo-so,
nsunsu, oi, Spd para, pig's dung grass, queue de rat,
rat's tail, rumput tahi babi, s&kar laroe, s%laseh
dandi, spotted basil, tabonsu, talche, tanagya,
tanodza, tsarkiyar kusu, tuetu, verbena, verbena azul,

Sg ee AN RE TM TT RT ee

1944 Moldenke, Common Names| 113

verbena cim, verbena de playa, verbena manza, vervain,
verveine, verveine queve de rat, water vine, wesussi,
Wutsiyar 'bera, wutsiyar kadangare, wutsiyar kusu
Stachytarpheta Maximiliani Schau. —- ger vao
Stachytarpheta Mexiae Moldenke -- pedrésy
Stachytarpheta mtabilis (Jacq.) Vahl -- balunakuta, bois de
chenilles rouge, djarongan, jarongan, ki meurit beur scum,
laler méngéng, rém&k g&tih, rumput puti, verveine A
fleurs rouges
Stachytarpheta orubica (L.) Vahl -- aristate bastard-vervain
Stachytarpheta polyura Schau. -- gervao
Stachytarpheta Robinsoniana Moldenke -- xtalché
aera straminea Moldenke -- verbena negra
Stachytarpheta trinitensis Moldenke -- verbena

Stachytarpheta urticaefolia (Salisb.) Sims -- albdka, balu-

nakuta, biana ana blau, bilu-bilu, biron, blue rats tail,
bolomoros, comasi, djarong, jaih babi, kandikandildan,
karoménal, kena-qele-yago, limbagat, maukakarawa,
mautofu tala, mautofu vao, mofalu, mokaukarau kedra,
motofu, nettle-leaved bastard-vervain, nettle-lLeaved
cymburus, ngadi-renggo, sekar-laru, tumbutumbu, turu-
Levu

Stilbe Berg. -- Heidenmiille, stilbé

Svensonia laeta (Fenzl) Moldenke -- marib

Symphor ema involucratum Roxb. -- gubba ddéra, konda tekkali,
nwey-sat, surudu

Symphorema luzonicum (Blanco) Fern.-ill. -- baldbai,
malabulaon, malasiad, malasiag, malaskog, mul auing-
b&ging, pamuleklakin

Tectona L. f. -- Indian oak, teak, teak tree, teak wood,
tectona, teka, Tekbaum, tektone, thek, Theka-baum, ~
Tiek-baum, Tik-baum

Tectona grandis L. f. -- edaritéku, bois de teck, bois de
tek, cajaten-hout, gaka, cay-g0-gia tri, chene des
Indes, chéne du Malabar, chingjagu, dalanang, dalandon,
dati, d31%¢, djati, djati, Djatibaum, djatiboom,
djatti-boom, djattie, djattie-boon, dodolan, Vast
Indian oak, Eisenholzbaum, fati, hadlaydti, hati,
iattie, Indian oak, indische Eicha. indische eik,
indische Teak, jadt, jati, Jatibaum, jatiboom, kalayati,
khaka, kyun, kywon, loheru, pedda téku, Bag, sag,
sagon, sagun, sagin, sagin, ségun, saguna, sagunyati,
sagvén, SagWan, Sagwan, sagwan, sagweni, saigun, 8aj,
séj, sak, saka, sdka, sa&khi, sal, sal, segun, segun,
sigwan, singuru, sipna, teak, Teakbaum, teakboom, teak
tree, teak tree of India, teak wood, teca, téca, teck,
teck des Indes, téga, tepgina, tégu, tek, ték, teka,
Tek-berm, teke, tekka, tekka-maram, tekku, tékku,
tékkumaram, téku, téku-mdnu, theca, theck, Theka Baum,

114 <p PEP OL OGL A Vol. 2) nos 3

Tihk Baum, tydgada mara, ydati, Zimmer Baum
Tectona Hamiltoniana Wall. -- dwarf teak, ta-hat, ta-nap
Tectona philippinensis Benth. -- bunglas, malaparngit
Teijsmanniodendron Ahernianum (Merr.) Bakh. -- agugs
amamahit, dalipapa, sangula, didigkalin, dilipdpak,
dufigila, galipdpa, igang, kalipépa, kolipdpa, kulipdpa,
malatgang, mamahit, mongpong, pamagsen, sasalit,
sasilit, sasulit, tayupuk

Teijsmanniodendron longifolium (Merr.) Merr. -- atikoko,
mamanau, sikukok —

Teijsmanniodendron pteropodum (Miq.) Bakh. -- médang poedi, -
pokok agak paya, sepoendang, sSpoegang, s&pugang,
sSpundang, sipanoeh, tj&mpana

Verbena L.-- camaradinhas, cambards, Eisen-hart, Eisen-
hort, Eisen-klette, Eisenkraut, EBisenweich, herva
cidreira, ijzerhard, iron-weed, jujubas, shah's
favourite, sha~passan, verbena, verbenas, vervain,
vervains, verveine, vervene, wijkruid, yepau

Verbena Abramsi Moldenke -- common vervain

Verbena bipinnatifida Nutt. -- common verbena, Dakota
verbena, purple verbena, small-flowered verbena, sweet
William, vervain

Verbena Blanchardi Moldenke -- blue vervain, false vervain
Verbena bonariensis L. -- hoi, South American vervain,

sudamerikanisches Eisenkraut, verbena, verveine,
verveine de Buénos~Ayres, verveine sauvage

Verbena bracteata Lag. & Rodr. -- bracted vervain, large-
bracted vervain, large-bracted vervaine, large-bracted
vervane, large-leaved vervain, lavender ground-flower,
prostrate vervain, verbean vervain, verbena, vervain

Verbena canadensis (L.) Britton -- cut-leaved vervain,
flowering verbena, large flowered verbena, large-
flowered verbena, large flower verbena, perennial
verbena, pubbeeten, purple verbena, Rocky Mountain
vervain, rosenrotes Eisenkraut, rose verbena, rose
vervain, sweet William, verbena, verbenen, vervain,
verveine & bouquets, verveine citronnelle, verveine de
Drummond, verveine de Miquelon, verveine de Miquelon
naine magenta, verveine de Miquelon naine rose,
verveine des Indes, verveine du Canada, verveine du
Pérou

Verbena canescens yar. Roemeriana (Scheele) Perry --
vervain, wild verbena

Verbena carolina L. -- verbena

Verbena ciliata Benth. -- alfornbrilla, moradilla

Verbena corymbosa Rufz & Pav. -- correcaballito, verbene

Verbena crithmifolia Gill. & Hook. -- bichicho

Verbena delticola Small -- kamiyo, moradia

Verbena dissecta Willd. -- margarita morada

anetil

1944 | Moldenke, Common Names 115

Verbena domingensis Urb, -- verbena
Verbena elegans var. asperata Perry -- moradilla
Verbena Engelmannii Moldenke -- nettle-leaved vervain

Verbena ephedroides Cham. -- jaqueca, verbena

Verbena Gooddingii var. nepetifolia Tidestr. -- kamiyo

Verbena Halei Small -- blue vervain, slender vervain,
vervain

Verbena hastata L. -- American vervain, blue vervain, blue
vervian, blue verbena, blue-verbena, clowne's all—heal
of New England, false vervain, iron-weed, purple
verbena, purvain, shades, simplers' joy, simpler's
joy, verbena, vervain, wild hyssop

Verbena hispida Ruiz & Pav. -- verbena

Verbena hybrida Voss -- Apple Blossom, Beauty of Oxford,
Blue Defiance, Blue Sentinel, Blue Shades, Burnett's
Scarlet, Cameo Pink, Cardinal, Carmine Ball, Carmine
Rose, Carter's Dwarf Coerulea, Carter's Dwarf Compact,
Carter's Holborn Mammoth, Cerise Queen, Chamois,
Coccinea, Coccinea Dwarfer, common garden verbena,
common verbena, Crimson Seedling, Crown Prince,
Dannebrog, Dark Blue, Vefiance, Defiance Re-selected,
Dwarf Coerulea, Dwarf Compact, Elfin Scarlet, Elite
Ellen Willmott, Etna, Fairy Queen, Fireball, Floradale
Beauty, Fordhook Famous, Giant Srect Mixed, Giant Pink,
Giant Salmon Pink, Golden Queen, Grandiflora Blue,
Grandiflora White, Holborn Mammoth, Lavender Glory,
Lucifer, Luminosa, Lutea Improved, Majestic Rose
Shades, Mamath, Mammoth Mixed, Mammoth Rose Queen,
Mammoth Scarlet Queen, Mammoth Snow Queen, Miss
Willmott, New Red, Oxford Pink, Pfitzer's Giant, Pink,
Pink and Rose Shades, Pure White, Purple Garnet, Red,
Rose Cardinal, Rosea Stellata, Royal Bouquet, Royale,
Salmon Defiance, Salmon Pink, Scarlet, Scarlet
Defiance, shah's favourite, sha-passan, Snow Queen,
Snowball, Snowdrift, Spectrum, Spectrum Red, verbeina,
verbena, verveine, Violacea Stellata, Violet Bouquet,
White, White-eye Blue Verbena

Verbena incisa Hook. -- margarita punzo, pubbeeten, verbenen

Verbena integrifolia Seasé & Moq. -- verbena

Verbena laciniata (L.) Briq. -- margarita morada, moss
verbena, sandia lahuen, verbena

Verbena lasiostachys Link -- common vervain

Verbena litoralis H.B.K. -- false vervain, verbena, verbena
blanca, verbena blanca serrana, vervena

Verbena menthaefolia Benth. -- bercul, weyhooli

Verbena moechina Moldenke -- hoary verbena

Verbena officinalis L. -- Altarblume, aristereon, Aschlepius
alceas, ashthroat, berbena, berbine, camaradinha,
chamelicos, columbina, columbine, common vervain,

116

PAY tO oy Geek Vol. 2, noe 3

creisetta, curetis fersefomon, cyparissos, demetrias,
diose lacete, echtes Eisenkraut, Hisen-bart, Eiseén-dek,
Bisen-hart, eisen-hendrik, Bisen-herz, Eisen-kraut,
eisen krokt, Hisenrsich, Hisenweich, eiserich, eisern,
eisernhart, eisewich, eisewig, enchanter's plant,
1 rd ¢ 7 s 2
enchanter’s-plant, erba de bermena, erbo 4 touti li
man, Buropean-verbena, European vervain, eysencrut,
eysenhart, eysenkruyd, eyssen-chrawtt, eyssen-kraut,
ferraria, gemeines Hisenkraut, geweihet kraut, Hahnen-
kampf, Hahnen-kopf, hardijzer, heiliges Kraut,
Heiligkraut, herbe & tous les maux, herbe aux
enchantements, herbe aux sorciéres, herbe de sang, herbe
du foie, herbe sacrée, herb-grace, herb-of-the-cross,
herb o' grace, herva de ferro, hierabotane, hierobotane,.
holy herb, holy-herb, iherabotane, iisercruyt,
ijsenkruyd, ijserhard, ijserkruyd, ijzerhard,
ijzerkruid, irenharde, isarna, isernina, isecruyt,
isekrut, isena, isenarre, isen-bart, isen-brut, isen-
hard, isen-kraut, iserbart, isercruyt, isere, iseren-
bart, iseren-hard, iseren krut, iserhart, iserhert,
iserich, isern, isernehart, isern Hendreck, isern krut,
isin-chlete, isin-ina, isin~-un, isirn, isirnwurz, isni,
issernhar, issinkraut, Juno's herb, Juno's tears,
Juno's-tears, Junothréinen, jururuba, kardita,
Katzenblut, kerckkruyd, kerkkruid, kuma-tsuzura,
licinia, lung nga ts'o, lustam, ma pin ts'o, Mercurie's
moist blood, Mercury's moist blood, Merkurblut,
militarem, ngoh sat na, Opferbraut, pamikh, pancremon,
pempentar, peristerion, perstereona, pigeon's grass,
pigeon's-grass, pigeon's grasse, pitagosas, planta de
sorte, reich-hard, reich-hart, Segen-kraut, shamuki,
shop vervain, sideritis, simpler's joy, simpler's-—joy,
sirpina, Stahl-kraut, Tauben-kraut, Taubenlieb, tialu,
tigrodion, varveino, varveyn, Venusader, verbena,
verbenam, vertiperdum, veruaine, vervain, vervayne,
vervein, verveine, verveine commune, verveine
officinale, verven, vervena, vervene, vervin,
Weihsprossen, wilder eisewig, ysen-hard, ysen-krute,
ysercruit, ysere, yseren, yseren-hard, yseren-hart,
yser-hard, yser-hart, ysern, ysernhard, ysinina

Verbena origenes R. A. Phil. -- hierba del incordio,

ricarrica

Verbena peruviana (L.) Britton -- brennende Liebe,

Feuerverbene, garden verbena, margarita punzo,
mar garita punzo, roode verbena, verbena, verbenas

Verbena phlogiflora Cham. -- camaradinha
Verbena platensis Spreng. -- piedra, verbena, verbena

blanca, verveine odorante

Verbena pumila Rydb. -- bartanucha, hairy verbena, kawiyo,

1944 Moldenke, Common Names LY,

pink verbena, pink vervain

Verbena rigida Spreng. -- hardy garden verbena, large-veined
vervain, stiff verbena

Verbena scabra Vahl -- rough vervain

Verbena simplex Lehm. -- bur-vine, narrow-leaved vervain,
pigmy vervain, vervain

Verbena stricta Vent. -- blue verbena, blue vervain, bur-

vine, fever-weed, hoar vervain, hoary verbena, hoary-
verbena, hoary vervain, mullen-leafed vervain, mullen-
leaved vervain, thimble-weed, verbena, wild hyssop

Verbena stricta f. albiflora Wadmond -- hoary vervain,
white-flowered verbena

Verbena Teasii Moldenke -- Albion Verbena, Bellaire Verbena,
Ceres Verbena, Madge Roberts Verbena, Rowena Verbena,
Ruth Verbena, Teas Hybrid Verbena

Verbena tenera Spreng. -- Italian verbena, verveine
délicate, verveine gentille, verveine jolie

Verbena tenera var. Maonetti Regel -- Italian verbena

Verbena tenuisecta Briq. -- amor de hombre, amor de hombre,
fumaria, hierba del incordio, lazo de amor, margarita
morada, moss verbena, perajil, rouen, sandia lahuen,
sandialahuén, verbena, verveine élégante, violette,
yerba del incordio

Verbena urticifolia L. -- bur-vine, common vervain, common
white vervain, nettle leaved vervain, nettle-leaved
vervain, nettle leaved Virginian vervain, verbena,
white verbena, white-verbena, white vervain

Verbena urticifolia var. leiocarpa Perry & Fernald --
nettle-lLeaved vervain, short-hair white vervain, white

vervain :
Verbena xutha Lehm. -- vervain
Verbenacese J. St. Hil. -- Eisenhartgewéchse,

eisenkrauta&mliche Gewachse, Eisenkrautartigen,
Eisenkrautgewachse, Verbenaichtigen, verbénacées,
Verbena Family, verbenas, Verbena Tribe, verbenes,
Vervain Family

Vitex Tourn. -- chaste tree, chaste-tree, chaste-trees,
gatilier, gatillier, gattilier, hemptree, Kenschbaum,
Keuschbaum, Keuschlamm, kuischboom, Maria preta,
Monchspfeffer, monks-pepper-tree, Mille, mullen,
Millen, rage tree, samaw hin, taruma, taruma, vitex,
vitice

Vitex Agnus-castus L. -- Abraham's balm, Abraham's balm
fruit, Abrahamsbaum, Abrahamsboom, Abrahamstrauch,
agneau chaste, agnocasto, agnus castus, agnus-castus,
alecrim d'Angola, aloch, arbre au poivre, arbre de
poivre, artenhewe, boom der kuisheid, borst-saame,
borst-samen, chaste lamb, chaste lamb tree, chaste
tree, chaste-tree, common chaste-tree, europische

118 PHYTOLOGIA Vole 2, no. 3

kuisboom, faux poivre, raux poivrier, gatilier,
gattilier, gattilier commun, grattilier, hemp tree,
hemp-tree, herbe chaste, Indian-spice, Keuschbaum,
Keuschbaummullen, Keuschlamm, Keuschlammstrauch,
keuschlamp, Klosterpfeffer, kuisch-boom, kuisch-lam,
Kiinschbaum, kuysche boom, Monchspfeffer, monks pepper-
tree, monk's pepper tree, monnikspeper, mullen, Miillen,
myrrh tree, pébré, pébrier, pepe di monaci, peperboom,
pepper taste, petit poivre, Pfefferbaum, pimienta de
Guinea, poivre des moines, poivre petit, poivre
saurage, poivre sauvage, sage tree, sanzgatillo,
Schaeffmille, Schaefsmtihle, schaapsmiul, Schaf-milch,
schaf-milte, Schaf-miile, Schaf-miille, Schaf-miillen,
schafs-milben, schafs-millen, Schafs-mlihlen, Schafs-
mulben, schafs—-mullel, Schafs-miillen, schafwulle,
totsane, tree of chastity, true chaste-tree, Virginia
sage, wild lavender, wild-pepper, zeowilg, uy Yds,
hiya ped, vyds, Avyss, tros, olcos ~

Vitex altissima L. f. -- ahay, balgay, banalgay, gua,
_kadamanakku, maila, milla, millilla, myrole, namilf
adogu, nauladi, sampaga-pala, sapu-milla, simyange.

Vitex Balbisii Spreng. -- bois des savannes, gri-gri, mata
~~ becerro, palo perrito

Vitex bankae H. J. Lam -- kajoe melati, mlak

Vitex barbata Planch. -- ba-kudu-ne, kuru, kutu-fingo |

Vitex brevipetiolata Moldenke -- ubaia

Vitex calothyrsa Sandw. -- tarume

Vitex capitata Vahl -- aceituno, escobillo, five-leaf
~~ £4ddlewood, guarataro, piedrero, totumillo, white
fiddlewood

Vitex chrysocarpa Planch. -- ba-kudu-ne, balamagnian kan,
~ 4nsuo-koto, kuru

Vitex cofassus Reinw. -- panafat, banohoeba, beso, biti,
~ boepasa, gawasa, gofasa, gofasa batoe, gofasa gaba- ~
gaba, gofasa mrah, gofasa p¥rampoean, gofasa tikar,
katonde, katondéng, pasal, vasari, wila

Vitex columbiensis Pittier -- acietuno
Vitex c compressa Turcz. -- aceituno-totumillo, acetuno,

acietu, acietuno, acietuno blanco, alasoabo, apokotja,
arbor procera, boschkalebas, bosch kalebas, fruta de
gonzalo, hakia-balli, leon pintado, taroema, totumillo,
white fiddlewood

Vitex Cooperi Standl. -- cua-ja, cuajada, raja bien

Vitex coriacea C. B. Clarke -- banton, connaropsis laurel,
“jali batu, m&dang pupoi, m&royan batu, pupoi, rock
m&royan , urat rusa

Vitex cymosa Bert. -- aceituno, acietuna, azietuno, guazuy,
taruma, taruméd, tarumé de varzea, taryma

Vitex Degeneriana Moldenke -- mannua cachorro

Sao, Vk me TER TA Sorter Phere

1944 - Oe ' Moldenke, Common Names 119

Vitex divaricata Sw. -- bastard fiddlewood, black
fiddlewood, bois agouti, bois d'agati, bois d'agouti,
bois de lézard, bois lézard, bois manive agouti, cote
lizard, higuerillo, manive agouti, palo de pendula,
pendola, pendulo blanco, timber fiddlewood, totumillo,
totumillo blanco

Vitex divaricata var. cubensis Urb. -- pable: guayo

Vitex Doniana Sweet -- abisoa, ada, adaga, ade, afetewa,
African oak, African teak, angalem, aranga, black
plum, burzun, dinchi, 'dinya, 'dumniya, 'dunya, dyob,
ebisaa, edin, &ji, ele-ele, f8, f¥-ti, FY yi, f¥ yi-ti,
fSyi-tsho, galbihi, galbije, gidjiko, heul, ingari, ink
tree, koto, kudu,kukpweli, kukui, kurugh, lubei,
lugbei, luwu-wului, makwaiwe, narenge, ngalbihi, nya,
nyerina, Scha koro, ofén; Sri, Sri-nla, Sri-Sdan,
panyer’, punyo-tsho, samanibir, song-sho, sd-tsho,
tschangbaio, tschangmaro, tschingmara, ucha koro, um-
digulgul, um-dugulgul, udli

Vitex Duckei Huber -- tarumdé, taruman

Vitex erioclona H. J. Lam --. lako

Vitex excelsa Moldenke -- taruma

Vitex ferruginea Schum. & Thonn. -- tiogbi

Vitex flava Ridl. -- jampang laki

Vitex flavens H.B.K. -- acietuno, mameira, pechiche, tarumé,
taruman -

Vitex Fosteri C. H. Wright -- akwakora~gyahini, ibang, koro
~ Koronta, obuban, ogi, Sri-&ta.

Vitex Froesii Moldenke -- taruma de mata

Vitex gamosepala Griff. -- cooked rice leban, 1¥ban nasi,
~~ 18ban pachat, leech's 1&8ban, p&long, s@tulang, sulong
chong

Vitex Gardneriana Schau. -- jaramataia

Vitex Gaumeri Greenm. -- barbas, barrabas, blue blossom,
blue flower, dachnik, fiddlewood, flor azul, flor
azule, jocote de mico, locote de mico, matasamo, monkey
fiddle, yaaxnic, yaax nic, yashnik, yash snick, yaxnic

Vitex gigantea H.B.K. -- moconto, pechiche

Vitex glabrata R. Br. — ampapalut, amuldon, ashwal ,
~ bhodiya, bihboel, bihbul, boetboet, bongdg, bofigig,
bonkolion, g®ntileng, gentileng ketileng, goda,
horina, htoukehar, kalipdpa-aso, kamolduan, karril,
k8tilwng, ki now, laban k&til8ng, laban k&til¥ng
hléng, longarbis.thiras, luki, neva-lédi, sasalit,
sengeni, senkane, sheras, sherasa, songarbi, tdlang-
puso, tilgng, tokra, topas

Vitex grandifolia Gtirke -- abisoa, ada, adaga, adefia,
~ afatewa, aranga, bofuluk, Sbisaa, £¥, f¥-ti, FY yi, FY
yi-ti, f8yi-tsho, God's coconut, ingari, ink tree,
kukpweli, kukui, lubei, lugbei, lugnei, luwu-wmulul,

120 Peay? 0 [+0'C 2A ~' Volks 25 toca

narenga, nya, nyam@1e-kukwe, nyarina, ogikhimi,
okurutu, orabia, Gri, oriri, ovuruburu, Swenkundigbon,
punyo-tsho, samanibir, song-sho, s06-tsho, uruahu

Vitex guianensis Moldenke -- hakiaballi

Vitex Hemsleyi Briq. -- capulin blanco

Vitex Kuylenii Stand). -- barabaés, barbés, flor azul

Vitex leucoxylon L. f. -- kaddunochchi, nebedda

Vitex lignum-vitae A. Cunn. -- lignum-vitae, Queensland
lignum-vitae

Vitex longisepala King & Gamble -- flowery 1¥ban, gading
~ kahua, halban, hearth-frame plant, kahua, 1%ban bunga,
foe kunyit, 1%ban nasi, pokok galang dapur, tumeric
1éban

Vitex lucens T. Kirk -- Neuseelandteak, New Zealand oak, New
Zealand teak, New Zealand teak tree, puriri, puriri
tree

Vitex madiensis Oliv. -- kuru kudulé ‘

Vitex Masoniana Pittier -- quajado

Vitex Mexiae Moldenke -- ipé branco, maminha, Maria preta

Vitex micrantha Giirke -- andofiti, djin-akwa, feve, fevei,
sah-sah

‘Vitex mollis H.B.K. -- agualamo, aguamalario, agiiilate,
ahuilote, atuto, coyotamate, coyotomate, flor de tila,
huhwwali, mate, nanche de perro, negro coyote,
obalamo, oovalama, tescalama, torete, ualama, uvalama,
uvatano, uvulama, valama, walamo

Vitex montevidensis Cham. -- bracuy, echter Tarumén,
guabiroba brava, tarumaé, tarumé, tarumé dure, taruma
guazu, taruman, tarumdn de ley, tarumfo, tatuman

Vitex Negundo L. -- ai toeban, amalu, aslag, banj-angasht,
~ bankahd, banna, barf, bari, beguniyd, bey guna, binna,
biuna, cannellier & feuilles de niekegas, chineesche
kuischboom, ehuri, fanjangasht, faux poivrier, filfil,
five-leaved chaste-tree, gattilie de Chine, gattilier
négundo, hobaro, ké&trf, kiyow-bhén-bin, kiyubdén-bin,
ko ling ngio, lagoendi laoet laki-laki, lakki, lakki-
gidd, lakkle, 1%ban,- 1¥nggundi, lingir, man king shue,
marwa, marwan, marwandai, maura, mawe , mehrwan, mewri,
mora, morann, mordun, nagoda, nalla varili, nargunda,
negundo, nengar, ng chi fung, nika, nikka, nirgandt
nirgari, nirgiri, nirguda, nirgudi, nirgunda,
nirgundi, nirgindi, nirgundf, nirgur, nirgér, nir-
nochchi, nishinda, nishindd, nisinda, nisinda, noch-
chi, pén{-ki-sambhalu, panj-angusht, pasatia, samalu,
sambhal, sambhélu, sandke, sanbhalu, semdlu, shambalf,
shamélu, shanbdéli, shawali, shiwdéli, shiwari, shvéte-
surasa, shwari, simali, sindhuca, sindhuvara, sinduari,
sindwar, sindwor, sisbaén, sudi-nikka, swanjan, tella-

_ it ea

Rael ee ee

1944 Moldenke, Common Names 121

vavili, three-Leaved chaste-tree, torban, torbanna,
vavali-padu, vavili, vellai-noch-chi, vella-noch-chi,
vel-noch cn, veyala, vrikshana, wana, warmande,
zukhamsate-asdbea, zuikhamsatil ourdq

Vitex Negundo var. cannabifolia (Sieb. & Zucc.) Hand.-Mazz.
-- chaste-tree

Vitex Negundo var. intermedia (P'ei) Moldenke -- potentilla

Vitex odorata Huber -- petit arbre de campos

Vitex orinocensis H.B.K. -- tasajo

Vitex orinocensis var. multiflora (Miq.) Huber -- taruma,
taruma preto, totumillo

Vitex oxycuspis J. G. Baker -- fevei, kpar-seh

Vitex arvifiora A. L- Juss. -- ad gauon, alah, amaréun ,
amuauan, amugauan , amul ation , anja, bulauen, burikén,
edieu, foeli kad, hamolauen , hamoréon , hamulai,
hamul ation , hamuréon , hamuyéon , hulgh, kajoe koela,
kalipdpa, kalipdpa-batd, kauere, kulimpapa, kulipépa,
malabalinanau, marauin, maulalin-aso, molauin, molauin,
molave, molave, molave~batu, moldve de playa, muldon,
mulauin, mulaviin, muravin, sagad, sagat, salingkdpa,
sasalit, taga, topas, tugas, tugas-bufgogan, tugas—
lanhan

Vitex peduncularis var. Roxburghiana C. B. Clarke -- bhadu,
_ boruna, goda, hila auwal, krawru, kyetyo, marak',
navalddi, osai, shelangri

Vitex Perriana var. abludens Moldenke -- Maria molle

Vitex pinnata | L. -- ablas, alaban, aloban, amurdon, black
~_1&ban, busi, cooked rice leban, din, flowery 1%ban,
goelimpapa, halaban, halban, haleban, haniban, h@jas,
horn 16ban, htouk-sha, kalapapa, ka non, ki arak,
kiketaroe, kon samaw, kopiher, kulimpdpa, kyet yo,
kyet-yob, laban, laban kapoer, laban k#tilé@ng, laban
koenjit, laban soengoe, laban tandoek, labari, labhan,
16ban, 1¥%ban boenga, 1¥%ban bunga, 1%ban haniban, 1%ban
hitam, 1¥%ban koenjit, 1%ban kunyit, 18ban nasi-nasi,
18ban pantis, 1¥ban tandoek, 18ban tandok, 183ban
toengkak, 13bén, manéh, molave, moron, muria, myladi,
nevalad ugu manu, nevali adugu, n&ban, non, nowli
eragu, pagil, salingkdpa, samaw buang, samaw tin pét,
tin nok, tumeric 1éban

Vitex polygama Cham. -- Maria preta, taruma&, taruman,
tarumao, velame de campo

Vitex polygama var. Bakeri Moldenke -- mamma cachorro de
catinga

Vitex pseudolea Rusby -- aceituno del monte, anacahiute,
~ anacahuita, taruma, torumar, wild olive

Vitex pyramidata B. 4. Robinson -- hupari, negrito, negrito
coyote, querenda, querendereniqua, tescalama, uvalama,
uvulama

122 PAY T0450 G:T is Vol. 25 noe 3

Vitex quinate (Lour.) F. N. Will. -- arnrai, basal, boengis,
flowery 18ban , gofasa, hemulauin, horn 18ban, kajoe
s¥moet, kalipdpa-asu, kalipdpa-madam, ka liu tsoi,
kamalan, k&tiléng, ki bangbara, koefo-koefo, koetil¥ng,
kojoe semoet, laban, laban kétilang, laban koening,
laban s%moet, 18ban boenga, 1¥ban bunga, 1¥%ban tandok,
limo-lfmo, limpdpa, Lifigo-lifigo, magupai, mamali,
masarwet, m&dang giring, moldave-aso, niue, rimoewas,
sai tsio tau, saivonta, sace masarawet, saoce poeti,
sace réndai, saoce séla, tileng, tugas, woelas watoe

Vitex rapinoides Guillaum. -- incdic, nay-mof-si

Vitex Rehmanni Gtirke -- thorns

~ -bummere, 'dinyar biri, 'dunyar biri, idj&li, kuru,
nambalerri, panyer’ buda, plum-tree :

Vitex Stahelii Moldenke -- alasaobo, hackiaballi, panda :

Vitex triflora Vahl -- mama cachorra, mama de cachoira, | ‘
~ tahuari, taruma, taruma de terrafirma

Vitex triflora var. coriacea Huber -- tarumé

Vitex triflora var. Kraatzii Huber -- mammi-cachorri

Vitex trifolia L. -- ai toeban, asla, banj-angashte-abi,
dangla, dinsaw, doenoeko, galoemi, gendarasi, hand of
Mary, Indian privet, jala-nirgundi, kiyoubhén-bin,
konti saw, lagoendi, lagondi, lagundi, lagindi,
lagundian, lagunding-dagat, langghoendi, langgoendi,
lanra, lawarani, 1&goendi, 1¥gundi, 1@nggundi,
lilégoendi, lingei, lipuk, nfra-lakki-gidé, nir-noch-
shi, nir-noch-chi, nfru-vévili, pani-ki-sanbhalu,
pan{-ki-shanbalf, péni-samalu, panj-angushta-abi,
que-abi, sagarai, salah gundi, sangari, shiru-noch-
chi, shiru-varfli, silagundi, sisua noi, suf éd-
sanbhelu, sunrasa-vrikshashe, tella-vavili, tigau,
uljf-shanbéli, vaturu-nikka, wild pepper, yé-kiyuban-
bin

Vitex trifolia var. bicolor (Willd.) Moldenke -- dabtan,
~ dangla, dauhon lagondie, lagundf, lala tea, lingei,
sagarai®

Vitex trifolia var. simplicifolia Cham. -- agubarau,
— daldaldkd, dancundi, dangla-ti-baybai, dungléd,
legund{f-ddgat, Tecra dagat, lagunding gapang,
lagunding-gapdng, paak pui ip, paak pui man king,
pak=-muk-ying, polinalina, so pa, vulokaka

Vitex umbrosa Sw. -- bois lezard, boxwood, fiddlewood, West

Vitex rivularis Giirke -- eEnemor sere ie antelope‘s garden
egg, kataboawin, old man's shin-bone, Stwe-ntdrowa,
ububan
Vitex rufa A. Chev. -- kpar-seh
Vitex Schaueriana Moldenke -- taruman, taruman-sinho 3
Vitex Sellowiana Cham. -- Maria preta, taruman
Vitex simplicifolia Oliv. -- buji, bummehi, bummeji,
;

1944 Moldenke, Common Names 123

Indian boxwood, yellow fiddlewood

_ Vitex unifoliolata Merr. -- babako

Vitex velutina (Koord. & Val.) Koord. -- k&til8ng

Vitex venosa H. J. Lam -- kajoe kahomboek gaeling,
kerindjing daoen talang

Vitex vestita Wall. -- alaban baengat, bangas jantan, black
18%ban, chichah, flowery 1%ban, halban, haleban, horn
18ban, k¥payan, 1¥%ban, 14%ban bunga, 18ban hitam, 1¥ban
nasi-nasi, 18ban nasi rimba, 1¥ban pelandok, 14%ban
tandok, maramboeéng, marambu&ng, mouse-deer's 1¥%ban,
rice 1¥%ban, s&%pit, tampang b¥si, tumeric 1¥ban

Vitex Wilmsii var. reflexa (H. H. W. Pearson) Pieper -- ama-
kosikati

Vitex sp. -- bois de la morue, bois de savanne franc, hab-
ul-takad, kabel jauwhout, renu-kabij, sham-baloo kabij,
tukm-i-pan jan gusht.

Viticipremna philippinensis (Turcz.) H. J. Lam -- bofgogon,
hamurauon-asu, kalimantau, kamalan, lifngo-lingo,
linolfno, lino-lffio, ma gomo , mala-moldave, malamulduin,
malduing-&s0, mala-usd, mulduin, tugas, tugas-bufgogon,
vasun g

—-— -— =& = = @|— —-—§ = = = °

(1) Moldenke, H. N., An alphabetic list of common and
vernacular names recorded for members of the
Verbenaceae and Avicenniaceae. 34 pp. New York Botani-
cal Garden, August 31, 1939.

Moldenke, H. N., A supplementary list of common and ver-
nacular names recorded for members of the Verbenaceae
and Avicenniaceae. 24 pp. New York Botanical Garden,
February 25, 1940. :

Moldenke, H. N., Additional common and vernacular names
recorded for members of the Verbenaceae and Avicennia-
ceae, Phytologia 2: 65--89. 1944.

SUPPLEMENTARY NOTES ON THE ERIOCAULACEAE, AVICSNNIACEHAE,
AND VESRBENAGEAE OF TEXAS. I.

Harold N. Moldenke

Herewith begins a series of notes supplementary to my
discussion of the Sriocaulaceae, Avicenniaceae, and Verben-
aceae in Lundell's "Flora of Texas", volume 3, part 1, pages
1--87 (1942). Since the publication of that work 144 addit-
ional Texan specimens ef these groups have been examined,
representing 104 new collections and bringing to light 34

124 PHT OL0 Gta | Vol. 2, noe 5

new county records. The abbreviations used for the herbaria
in which these specimens are deposited are in conformity
with the ones used by me in all my independent works to date
and elucidated in my paper entitled "A list showing the 10-
cation of Li principal collections of Verbenaceae and Avi-
cenniaceae", pp. 1--5 (February 20, 1942 2). For convenience
the ones used in the present supplement are repeated here-
with: Bt = Butler university, Indianapolis, Indiana; Ca =
University of California, Berkeley; N = Britton Herbarium,
New York Botanical Garden, New York City; Ok = University of
Oklahoma, Norman; Tr = 8. M. Tracy Herbarium, Texas Agricul-
tural Station, College Station; Up = University of Pennsyl-
vania, Philadelphia; Ur = University of Illinois, Urbana;
and W = United States National Herbarium, Smithsonian Insti-
tution, Washington.

ERIOCAULON DECANGULARE L.
-Additional citations: Waller Co.: E. Hall 675 Sue

AVIC™NNIA NITIDA Jacq.
Additional citations: Cameron Co.: R. Run user 2 O77 (N).

ALOYSIA LIGUSTRINA (Lag.) Small

Additional citations: Brewster Co.: Cutler 4779 9 (N); G.
L. Fisher. s.n. [Marathon, July 11, 1927] (Bt). Cameron Coe
R. Runyon 2088 (N). Comal Oo. Lindheimer 1070 (Ok). Jeff
Davis Co.: Hinckley gene (July! '16, 1936] (N). Presidio Co-:
Hitchcock & Stanford 6811 (N).

ALOYSIA LIGUSTRINA var. SCHULZIL (Stand1.) Moldenke
Additional citations: Presidio Co.: O. M. Clark 4764 (0k)

ALOYSIA MACROSTACHYA (Torr.) Moldenke
Additional citations: Zapata Co.: Cory 35913 (N).

ALOYSIA WRIGHTII (A. Gray) Heller :
Additional citations: Jeff Davis Co.: Hinckley sen. [Aug.
22, 1939] (N). :

CALLICARPA AMERICANA L. 3
“Additional citations: Harris Co.: Fisher 8 en. :
[Houston, Sept. 14, 1930] (Bt), een. Banker 2307] (N)

Pree
tie eer Oey

Ge Le
[Herb.

CITHAREXYLUM BURLANDIERI B. L. Robinson :
Additional citations: Cameron Co.: Cory 36624 (N). Willa-
cy Co.: Tharp 1249 (Bt). =:

CITHAR=XYLUM SPATHULATUM Moldenke & Lundell, Contrib. Univ.
Mich. Herb. 8: 82--83. 1942. Citharexylum brachyanthum

os a acel anmeal

1944 Moldenke, Supplementary Notes 125

var. glabrum C. L. Hitchc. & Moldenke in Fedde, Repert. 37:
218. 1934; Lundell, Flora of Texas 3/: 75. 1942.

See the original publication of this species for a com-
plete description and discussion of its differences from OC.
brachyanthum. : .

Additional citations: Hidalgo Co.: Lundell & Lundell 9953
(Mi--type).

DURANTA REPENS var. ALBA (Masters) L. H. Bailey
Additional citations: Hidalgo Co. (cultivated): Cory

36181 (N). |

LANTANA CITROSA (Small) Moldenke
Additional citations: Cameron Co.: Cory 36729 (N).

LANTANA HORRIDA H.B.K.

An additional Texan reference to this species is in Anna
May Tarrance Davis, A study of Boscaje de la Palma in Camer-
on County, Texas, and of Sabal texana (thesis), pp. 33 & 61.
August, 1942. -- An additional synonym is Lantana rubra
Berland. in Teran. & Berland., Mem. Comision Limites 15.
1832. :

Additional citations: Cameron Co.: Cory sen. [11-14-]940]
(N). Gonzales Co.: Tharp sen. [Ottine, 5/1/35] (Bt). Llano
Co.: G. L. Fisher sen. [Llano, Apr. 21, 1930] (Bt). Travis
Co.: Tharp s.n. [Austin, 5/9/35] (Bt). Zapata Co.: Cory
35916 (nN).

LANTANA MACROPODA Torr.

Additional citations: Dimmit Co.: Texas Apr. Exp. Sta.
Herb. %xch. sen. [12-10-39] (Tr). Hidalgo Co.: Cory 36035
(N). Houston Co.: Tracy 9146 (Up). Starr Co.: Cory 3593
(N). Val Verde Co.: Cory 31715 (N), 38097 (N), 39746 (N).
Zapata Co.: Cory 35922 (N).

LIPPIA ALBA (Mill.) N. %. Br. |
An additional Texan reference is A. M. T. Davis, A study
of Boscaje de la Palma in Cameron County, Texas, and of
Sabal texana (thesis), pp. 33 & 62. August, 1942.
Additional citations: Cameron Co.: Nealley 113 (Up).
Hidalgo Co.: Cory 36288 (N).

LIPPIA GRAVEOLENS H.B.K.
Additional. citations: Houston Co.: Tracy 9158 (Up). Val
Verde Co.: Cory 38065 (N). Zapata Go.: Cory 359350 (N).

PHYLA CUNSIFOLIA (Torr.) Greene
Additional citations: Crockett Co.: Cory 18895 (N), 188
(N), 32749 (N), 39335 (N). Jeff Davis Co.: Hinckley 466 (N).

126 PHY 704-0 6.1.x Vol. 2, noe 3

Ochiltree Co.: Headlee 56 (Tr). Potter Co.: G. J. Goodman
3052 (N). Schleicher Co.: Cory 34447 (N).

PHYLA INCISA Small

An additional Texan reference is A. M. T. Davis, A study ©
of Boscaje de la Palma in Cameron County, Texas, and of
Sabal texana (thesis),p. 61. August, 1942.

Additional citations: Cameron Co.: R. Runyon 2085 (N).
Comal Co.: Lindheimer 1069 (Ok). Crockett Co.: Parks & Cory
18896 (N). Edwards Co.: G. L. Fisher sen. [Rock Springs,
July 19, 1955] (Bt). #1 Paso Co.: G..W. Dunn sen. [1 Paso,
July 20, 1887] (Up). Jackson Co.: Drushel 10260, in part
(Ok). Jefferson Co.: Wherry sen. [September 7, 1936] (Up).
Kinney Co.: Cory 33472 (N). Midland Co.: Cory 40613 (N).
Presidio Co.: Hinckley 691 (N). Tarrant Co.: Ruth 106 (Up).
Tom Green Co.: Cory 39602 (N). Travis Co.: Tharp s.n.
[Austin, 5-15-35] (Bt). Val Verde Co.: Cory 37997 (N). Will-
acy Co.: Cory 36726 (N).

PHYLA LANCEOLATA (Michx.) Greene

An additional Texan reference is A. M. T. Davis, A study
of Boscaje de la Palma in Cameron County, Texas, and of
Sabal texana (thesis), p. 61. August, 1942.

Additional citations: Liberty Co.: Langman 1945 (Up).

PHYLA NODIFLORA var. R=PTANS (H.B.K.) Moldenke

Additional citations: Cameron Co.: Cory 36468 (N); Tharp
1203 (N). Comanche Co.: Lindheimer 1071 25 Uvalde Co.:
Cory 39429 (N).

PRIVA LAPPULACTA (L.) Pers.

An additional Texan reference is A. M. T. Davis, A study
of Boscaje de la Palma in Cameron County, Texas, and of
Sabal texana (thesis), p. 61. August, 1942.

Additional citations: Cameron Co.: Cory 36622 (N).

TETRACLEA COULTERI A. Gray
Additional citations: Presidio Co.: Hinckley 1056 (N).

VERBENA AMBROSIFOLIA Rydb.
Additional citations: Cory 40367 (N).

VERBENA BIPINNATIFIDA Nutt

Additional citations: Bastrop Co.: Duval sen. (W). Comal
Qo.: Lindheimer 1073 (Ok). Crockett Co.: Cory 35469 (N).
Gonzales Co.: Friesner 10376 (Bt); Pladeck s.n. [near Gonza-
les; 3-30-1940] (Bt). Maverick Co.: M. ©. Jones 28300 (Ca). ©
Sutton Co.: Cory 39625 (N). Travis Co. ‘Rose & Russell 24129
(W); K. &. Smith s.n. [Austin, 5/1/35] (Bt); Tharp sen. [Au-

eee

1944 Moldenke, Supplementary Notes 127

stin, 4/12/35] (Bt). Waller Co.: Dixon 561 (W). County un-
determined: L. I. Davis 1 (N), 3 (N).

VERBENA BIPINNATIFIDA var. LATILOBATA Perry
Additional citations: Hidalgo Co.: L. I. Davis 199 (N).

VERBENA BRACTEATA Lag. & Rodr.
Additional citations: Presidio Co.: Hinckley 694 (N).
Sherman Co-: Wedver 17793 (Tr).

VERBENA CAM@RONENSIS L. I. Davis, Nature Leaflet 1: 1--3.

August 14, 1941. Verbena Lundelliorum Moldenke, Phytolog-
ia 2: 24. August 26, 1941. -- Careful comparison of publica-
tion records shows that Davis' name for this species was ac-
tually published and distributed to botanists and botanical
libraries twelve days before my name, so his name becomes
the valid name for the species. Anna May Tarrance Davis, in
her thesis entitled "A study of Boscaje de la Palma in Cam-
eron County, Texas, and of Sabal texana", pp. 31, 39, 41,
42, 43, & 61 (August, 1942), gives valuable additional in-
formation about this species and on pl. 10 a splendid illus-
tration of it. Unfortunately, she misspells the synonymous
binomial "Verbena lundellorum Moldenke".

Additional citations: Cameron Co.: L. I. Davis sn.

[March 7, 1942] (N), sen. [March 22, 1942] (N), sen. (N).

VERBENA CANADENSIS (L.) Britton

Additional citations: Culberson Co.: Hitchcock & Stanford
6782 (N). Harris Co.: G. L. Fisher sen. [Spring, May 4,
1924] (Ur), s.n. [Houston, Apr. 3, 1920] (Ur).

V@RBENA CANESCENS var. ROSMERIANA (Scheele) Perry

Additional citations: Edwards Co.: Parks & Cory 20841
(Tr). Kendall Co.: Parks & Cory 12929 (Tr). Llano Co.: Wolff
3064 (Tr). Uvalde Co.: H. R. Reed 53818 (N).

VERBENA CILIATA Benth.

Additional citations: Brewster Co.: Cory 35568 (N). Sd-
wards Co.t Oory 38871 (N). Hall Co.: Re We Bennett 44 (Tr).
VERBENA CILIATA var. LONGIDENTATA Perry

An additional Texan reference is A. M. T. Davis, A study
of Boscaje de la Palma in Cameron County, Texas, and of
Sabal texana (thesis), pp. 42 & 61, pl. 10. August, 1942.

Additional citations: Cameron Co.: Muenscher & Muenscher

14457 (N); Nealley 116 (W).

“VERBENA CILIATA var. PUBERA (Greene) Perry
Additional citations: Jeff Davis Co.: Tracy & Sarle 162,

128 PHYTOLOGIA _ Vol. 2, no» 57m
in part (Tr--isotype).

VERBENA CLOVSRI Moldenke ; :
Additional citations: Brooks Co.: Pladeck s.n. [near Fal-
furrias, 5-5-1940] (Bt).

VERBENA DELTICOLA Small

An additional Texan reference is A. M. T. Davis, A study
of Boscaje de la Palma in Cameron County, Texas, and of
Sabal texana (thesis), p. 61. August, 1942.

f
oa

VERBENA DELTICOLA f. LILACINA L. I. Davis in A. M. T. Davis, ©
A study of Boscaje de la Palma in Cameron County, Texas,
and of Sabal texana (thesis), p. 62 [as "lilaciana"]. Aug-
ust, 1942; Moldenke, Knowm Geogr. Distrib. Verbenac. Supple
1: 2, nom. nud. Novenber 15, 1943. -- The type of this color
form was collected by L. I. Davis on the banks of Resaca del ~
Rancho Viejo, Cameron County, Texas, in May, 1942, and is

deposited in the herbarium of the University of Texas.

VERBENA HALEI Small

An additional Texan reference is A. M. T. Davis, A study
of Boscaje de la Palma in Cameron County, Texas, and of ;
Sabal texana (thesis), p. 72. August, 1942. In regard to
Lindheimer 155, in part, cited on page 22 of my work in the ~
Flora of Texas as possibly from Palo Finto County, Dr. Geis—
er thinks that Lindheimer never collected in Palo Pinto a
County. The specimen may have come from some other county -- —
its label states merely "Dry prairies on the Brazos, July |
and August, 1843". . E

Additional citations: Anderson Co.: K. &. Smith s.n. [Pal-
estine, 4/21/35] (Bt). Bell Co.: Wolff 2948 (Tr), 3491 (Tr), —
3718 (Tr). Bexar Co.: G. Jermy 88 (W). Fayette Co.: Parks &
Cory 10072 (Tr). Grimes Co.: Weaver 1038 (N). Harris Co.: G.
L. Fisher sen. [Houston, Apr. 9, 1931] (Bt). Jim Wells Co: —
Muenscher & Muenscher 14391 (N). Llano Co.: Wolff 3067 (Tr,
Tr), 3825 (Tr). Nueces Co.: Tracy sen. [Corpus Christi, 3-
31-1905] (Tr). Travis Co.: Tharp s.n. [Austin, 5/2/55] (Bt). —

VERBENA NEOMEXICANA (A. Gray ) Small
Additional citations: Jeff Davis Co.: Hinckley sen. [Ju-
ly 5, 1936] (N). :

VERBENA NEOMEXICANA var. HIRTELLA Perry
Additional citations: Bexar Co.: O. M. Clark 7441 (Ok).

VERBENA NEOMEXICANA var. XYLOPODA Perry 4
Additional citations: Presidio Co.: Hinckley 1254 (N). ~~

Sa ae

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PHY TOLOGIA -

_ Designed to expedite botanical publication

Vor. 2 June, 1946 No. 4

CONTENTS

Published by H. A. Gleason and Harold N. Moldenke
The New York Botanical Garden
Bronx Park, New York 58, N. Y.

Price of this number, 65 cents; per volume, $5.00 in advance

Vol. 2, No. 3, was issued January 4, 1945

=

NOMENCLATURAL NOTES -- IIT

Harold N. Moldenke

Continued studies in the herbarium of the New York Botani-
cal Garden and in the field have brought to light several as
yet undescribed species, varieties, forms, and hybrids, and
have revealed the necessity of publishing certain new names
and combinations.

AEGIPHILA VALLENSIS Moldenke, sp. nov.

Frutex scandens; ramis dense lanatis; foliis oppositis;
petiolis dense lanatis crassis; laminis firme chartaceis vel
subcoriaceis ovatis longe acuminatis integris, ad basin ro-
tundatis, juventute lanatis, supra maturitate substrigoso-
pubescentibus, subtus dense sublanuginoso-villosis; inflores-
centiis terminalibus axillaribusque cymosis vel paniculatis
multifloris; pedunculis bracteisque bracteolisque dense
flavido-lanatis; limbo calycis 4-lobato.

Large woody vine; branches obtusely tetragonal, medium-
stout or slender, very densely lanate with matted cinereous
hairs; principal internodes 7--11 cm. long; leaf-scars and
buds very densely lanate like the branches; leaves decussate-
opposite; petioles stout, 10--15 mm. long, very densely len-.
ate with cinereous matted hairs; blades firmly chartaceous
or subcoriaceous, dull-green above, ashy beneath, ovate, 1/--
25 cm. long, 7--l2 cm. wide, long-acuminate at apex, rounded
at base, entire, substrigose-pubescent above, lanate when
young, very densely sublanuginous-villous with cinereous or
yellowish hairs beneath; midrib slender, slightly prominent
above, conspicuously prominent beneath; secondaries slender,
11--15 per side, arcuate-ascending, joined only by small
tertiaries at the margins, subimpressed above, prominulous
beneath; tertiaries abundant, subimpressed above, prominu-
lous beneath; inflorescence terminal and in the axils of the
uppermost pair of leaves, cymose or paniculate, many-flower-
ed; peduncles densely lanate like the branches, the axillary
ones 1--2 cm. long, the terminal ones to 7 cm. long; folia-
ceous bracts sometimes present beneath the terminal panicle,
densely yellowish-lanate or -villous; bractlets and prophyl-
la numerous, linear-filiform, 10--15 mm. long, twisted,
densely villous-lanate and almost hidden in the dense tomen-
tum of the inflorescence; calyx cyathiform, its tube abott 6
mm. long and 4 mm. wide, very densely villous with yellowish
forward-pointing hairs, its limb 4-lobed, the lobes about 2
mm. long, obtuse or acute, densely villous like the tube;
corolla infundibular or hypocrateriform, sulphur-yellow, its

129

1350 PH Y.TO 1:0 G4 2°4 Vol. 2, now 4

tube slender, 4--5 mm. long, glabrous, its limb 4-parted,
the lobes about 3 mm. long, obtuse at apex, glabrous; stam-
ens included; pistil long-exserted; fruiting-calyx incrass-
ate, to about 1 cm. long and wide, densely villous, its rim
deeply 4-lobed; fruit drupaceous, orange, globose, about 9
mm. long and wide, fleshy, glabrous.

The type of this species was collected by José Cuatrecas-
as (no. 15564) in woods at La Laguna , alt. 1250--1400 m., on
the left side of the valley of the Rio Sanquininf, on the
west slope of the Cordillera Occidental, El Valle, Colombia,
between December 10 and 20, 1943, and is deposited in the
Britton Herbarium at the New York Botanical Garden. The
species is obviously related to A. cordata Poepp.

CORDIA RANGELENSIS Moldenke, sp. nov.

Frutex vel arbor; ramis ramulisque gracilibus griseis
irregularibus, juventute dense adpresso-puberulentibus, sen-
ectute glabrescentibus; foliis alternis; petiolis crassius-
culis adpresso-puberulentibus canaliculatis vel sulcatis;
laminis coriaceis suborbicularibus vel ellipticis vel subob-
lanceolatis, acutis vel brevissime acuminatis, integris
subrevolutis, ad basin attenuatis, utrinque microscopice
scabrellis pernitidis.

Shrub or tree; branches and branchlets slender, gray, ir-
regular or even slightly zigzag, the youngest parts densely
appressed-puberulent with very short yellowish-brown hair
visible only under a hand-lens, the older parts glabrescent;
leaves alternate; petioles rather stout, 5--10 mm. long,
appressed-puberulent like the branchlets, canaliculate or
sulcate above, wrinkled in drying; blades coriaceous, vary-
ing from suborbicular to elliptic or slightly oblanceolate,
3.5--8.5 cm. long, 2--4.2 cm. wide, normally acute or very
short-acuminate at apex, often (apparently abnormally) ob-
tuse or even retuse, attenuate to the base, entire, subrevo-
lute, microscopically scabrellous on both surfaces, very
shiny on both surfaces, appressed-puberulent on the midrib
benesth; midrib slender, sharply prominulent above, promin-
ent beneath; secondaries, tertiaries, and veinlets forming a
dense and conspicuous reticulum which is conspicuous and ~—
prominulent on both surfaces; inflorescence axillary, soli-
_ tary -in one of the uppermost axils, about 16-flowered; ped-
uncles slender, 1.5 cm. long or less, several-branched, ap-
pressed-puberulent and also with a few longer spreading
hairs above; pedicels slender, about 1 mm. long, puberulent
and often also pilose with longer spreading hairs; calyx cu-
puliform, 3--4 mm. long, about 3 mm. wide, firm, appressed-
puberulent throughout, irregularly split into 2 or more
lobes toward the apex.

The type of this species was collected by Juan Tomas Roig

aa

1946 - Moldenke, Nomenclatural Notes - 131

y Mesa and Julian Acufla Galé (no. 4531) on the Sierra de
Rangel, Finar del Rfo, Cuba, between August 27 and 30, 1927,
and is deposited in the Britton Herbarium at the New York
Botanical Garden. Another collection was made by Acufia Galé
(no. 14119) at Taco-Taco, Rangel, in March, 1923.

CORNUTIA LATIFOLIA f. ALBA Moldenke, f. nov.

Haec forma a forma typica speciei corollis albis recedit.
-- This form differs from the typical form of the species in
having white corollas.

The type was collected by William Campbell Steere, with-
out number, in chaparral at Champoton, Campeche, Mexico, in
July, 1932, and is deposited in the herbarium of the Insti-
tuto Biologia at Mexico City.

XCRYPTOSTEGIA HYBRIDA Moldenke, nom. nov.

Cryptostegia madagascariensis x C. grandiflora Polhemus,
Hill, & Elder, U. S. Dept. Agr. Tech. Bull. 457: S721)» Plie

1 & 35--9. 1934.

DERMATOCALYX PANDURATUS Moldenke, sp. nov.

Arbor myrmecophila; ramis griseis glabris; foliis opposi-
tis, cicatricibus amplis suberosis elevatis glabris; laminis
firme membranaceis panduriformibus vel obovatis integris ac-
uminatis ad basin subamplexicaulibus utrinque glabris.

Tree to 8m. tall; trunk 13 cm. in diameter at breast
height; branches rather stout, gray, glabrous, prominently
lenticellate, rather angular, flattened at the nodes; leaf-
scars large, broad, elevated, with corky margins; leaves de-
cussate-opposite; petioles very thick, about 5 mm. long or
less, somewhat arched or humped, glabrous; leaf-blades firm-
ly membranous, panduriform or obovate, 13--28 cm. long, 5--
12 cm. wide, acuminate at apex, entire, subamplexicaul at
base, glabrous on both surfaces; midrib flattened or slight-
ly canaliculate above, stout and prominent beneath; second-
aries heavy, about 6 per side, ascending, arcuate near the
margins, flattened or slightly impressed above, very promin-
ent beneath; vein and veinlet reticulation subprominulous on
both surfaces, especially beneath; inflorescence axillary,
glomerate or fasciculate; flowers not seen, but said to be
white; fruiting-pedicels very heavy, about 5 mm. long, glab-
rous; fruiting-calyx very stout and heavy, campanulate-cupu-
liform, about 8 mm. long and 15 mm. wide when fully mature,
leathery, glabrous, somewhat 2-lipped and irregularly lobed,
often scarious-margined; fruit subglobose, about 1 cm. long
and wide, glabrous, hard, 2-celled; seeds very numerous on
both surfaces of the central placenta.

The type of this species. was collected by Elbert L.
Little, Jr. (no. 6317; U. S. Dept. Agr. Forest Service

130. FHYTOLOGIA Vol. 2, no. 4

96771) in partly cut wet tropical forest, alt. 10 m., 2 km.
south of San Lorenzo, Esmeraldas, Ecuedor, April 21, 1943,
and is deposited in the Britton Herbarium at the New York
Botanical Garden. The collector records the vernacular name
"mata palo" and reports that there are "black chambers from
ants all over."

ERIOCAULON ATABAFENSE Moldenke, Known Geogr. Distrib. Erio-
caul. 5 & 32, nom. nud. (1946), sp. nov.

Herba; foliis caespitosis firmis linearibus utrinque gla-
bris nitidis stramineis argute acutis vel mucronulatis; pe-
dunculis paucis costatis paullo contortis glabris; vaginis
firmis stramineis laxis glabris nitidis oblique fissis; cap-
itulis albis duris.

' Herb to 30 cm. tall; leaves cespitose, the fresh ones
few, firm,to 3.8 cm. long, linear, glabrous on both sur-
faces, shiny, stramineous, sharply acute or mucronulate at
the apex, not visibly fenestrate, the old ones thin-membran-
ous, numerous, translucent, very plainly fenestrate, to 12
em. long, very weak and flaccid, glabrous; peduncles one or
more per plant, 12--17 cm. long, several-costate, somewhat
twisted, glabrous; sheaths firm, stramineous, loose, about
3.8 cm. long, glabrous, shiny, obliquely split at the apex,
the lobes sharply acute; heads very hard, not flattening in
pressing, very white.

The type of this species was collected by Llewelyn Will-
iams (no. 13858) in sandy soil along a stream at Chamuchina,
alt. 280 m., Rio Atabapo, Amazonas, Venezuela, on January
19, 1942, and is deposited in the United States National
Herbarium at Washington.

ERIOCAULON CAPITULATUM Moldenke, sp. nov.

Herba pumila; foliis caespitosis paucis parvis glabris
non fenestratis; vaginis membranaceis laxis glabris fissis;
pedunculis solitariis glabris 5 cm. longis; capitulis grise-
is vel albidis; bracteis involucrantibus paucis obovato-
ellipticis pallide stramineis glabris, ad apicem rotundatis.

Plants very small, tufted; stems obsolete; leaves cespi-
tose, few, 1--2 cm. long, ampliate-sheathing at base, blunt
at apex, glabrous on both surfaces, not visibly ribbed nor
fenestrate; sheaths membranous, about 18 mm. long, glabrous,
split at the apex, loose; peduncles solitary, about 5 cm.
long, glabrous; heads about 5 mm. wide, grayish or whitish;
involucral bractlets few, obovate-elliptic, pale-stramineous
throughout, about 2.6 mm. long and 1.3 mm. wide, rounded and
slightly cucullate at the apex, glabrous and shiny on both
surfaces; receptacle subglabrate; receptacular bractlets ob-
ovate, brown towards the apex, about 1.8 mm. long and 1.3
mm. wide, concave within, convex on the outside, rounded at

1946 Moldenke, Nomenclatural Notes 133

apex, short-pilose at the apex but not tufted-bearded; stam-
inate florets: sepals 3, obovate-cuneate, spathaceous-con-
nate at the base, brown except at the united base, about
1.56 mm. long and O.7 mm. wide, slightly cucullate and apic-
ulate at the apex, glabrous on both surfaces; petals 5, uni-
ted into a slender hyaline tube about 0.6 mm. long, the free
apical lobes also about 0.6 mm. long, hyaline, densely white
bearded at the apex; stamens 6; anthers dark-brown or black;
pistillate florets: sepals 4, free and separate to the base,
dark-brown or nigrescent throughout, obovate, navicular,
carinate or short-alate on the back below the middle, about
1.56 mm. long and 1.1 mm. wide, short-acuminate at the apex,
white-pilose with very short hairs throughout on the back;
petals 3, separate to the base, oblanceolate, about 1.8 mm.
long and 0.5 mm. wide, hyaline, black-glanduliferous below
the apex, white-pilose on the inner surface; ovary 3-celled,
3-ovulate; style about 0.6 mm. long; stigmas 3, about 0.6
mm. longe ©

The typw of this dwarf species was collected by Carl Aug-
ust Ehrenberg (no. 219, in part) somewhere in Mexico and is
deposited in the Herbario Nacional of the Instituto Biologia
de Universidad Nacional de Mexico (no. 2608, in part) at
Mexico City. The heads superficially resemble those of E.
Ehrenbergianum Klotzsch, but differ in their essential char-
acters.

ERIOCAULON PARADOXUM Moldenke, sp. nov.

Herba perpumila; foliis paucis caespitosis glabris non
fenestratis; pedunculis solitariis 2 cm. longis glabris;
vaginis membranaceis laxis fissis glabris; capitulis grise-
is; bracteis involucrantibus pallide stramineis oblongis
glabris, ad apicem rotundatis.

Plants very small, tufted; stems obsolete; leaves few,
cespitose, 1--2 cm. long, 1.5--2 mm. wide, blunt at the a-
pex, glabrous on both surfaces, not visibly ribbed nor fen-
estrate, erect; peduncles solitary, about 2 cm. long, glab-
rous; sheaths membranous, loose, about 15 mm. long, split at
the apex, glabrous; heads grayish, about 5 mm. in diameter;
involucral bractlets pale-stramineous, oblong, 2--2.% mm.
long, 1--2 mm. wide, rounded at the apex, glabrous on both
surfaces; receptacle glabrate; staminate florets not well
developed (or past anthesis?), very few; receptacular bract-
lets cuneate-obovate, very dark-brown or black above the
middle, about 1.6 mm. long and 1.3 mm. wide, triangular-acu-
minate and cucullate at the apex, densely white-pilose with
short appressed hairs on the back and margins, glabrous
within, not tufted-bearded; sepals 3, hyaline, elliptic,
about 0.8 mm. long and 0.3 mm. wide, concave within, convex
On the back, brownish towards the obtuse apex, bearded at

1) PHYTOLOGIA Vol. 2, n@ere

the apex on the back; petals 3, united into a hyaline tube
about 0.26 mm. long, the apical free portions also about
0.26 mm. lorig, glabrous; stamens not seen; rudimentary pis-
til present; pistillate florets: sepals 4, free to the base,
dark-brown throughout, elliptic or subobovate, navicular, a-
bout 1.8 mm. long, O.7--1 mm: wide, acute at the apex, white
pilose with obscure appressed hair on the back, more con-
spicuously so on the upper margins and apex; petals 3, free
to the base, hyaline, oblanceolate, about 1.5 mm. long and
O.4 mm. wide, acute at the apex, attenuate to the base,
long-pilose on the back from the middle to the apex, black-
glanduliferous just below the apex; ovary 3-celled, Z-ovula-
te; style about 0.7 mm. long, glabrous; stigmas 3, about 0.2
mM e long.

The type of this very dwarf species was collected by Carl
August Ehrenberg (no. 219, in part) somewhere in Mexico and
is deposited in the Herbario Nacional of the Instituto Bio-
logia de Universidad Nacional de Mexico (no. 2608, in part)
at Mexico City. The species superficially greatly resembles
E. microcéephalum H.B.K. in its habit of growth, but differs
in the technical characters of its heads and florets. The
foliage of E. paradoxum and E. capitulatum is almost identi-
cal.

ERIOCAULON SIERRALEONENSE Moldenke, Known Geogr, Distrib.
Eriocaul. 21 & 40, hyponym (1946), nom. nov.
Eriocaulon pumilum Afzel. ex Korn., Linnaea 27: 621.

1856 [not E. pumilum Ref., Atl. Journ. 121. 1832].

ERIOCAULON TOGOENSE Moldenke, Known Geogr. Distrib. Erio-
caul. 21 & 41, hyponym (1946), nom. nov.
Eriocaulon xeranthemoides Van Heurck & Muell.-Arg. in Van
Heurck, Obs. Bot. 103. 1870 [not E. xeranthemoides Bong.,
Act. Petrop. Sci. Math., ser. 4, 1: 635. 1831].

GALPINSIA LAVANDULAEFOLIA var. GLANDULOSA (Munz) Moldenke,
combe nov.

Oenothera lavandulaefolia var. glandulosa Munz, Am.
Journ. Bot. 16: 705. 1929.

HALERPESTES CYMBALARIA var. SAXIMONTANUS (Fernald) Moldenke,
combe nOVe
Ranunculus cymbalaria var. saximontanus Fernald, Rhodora

16: 162. 1914.

HYPERBAENA CUATRECASASI Moldenke, sp. nov.

Frutex scandens; ramis pendulis gracilibus glabris; peti-
olis gracilibus sulcatis glabris, ad basin curvatis; laminis
coriaceis nitidis ovatis longe acuminatis integris utrinque

1946 Moldenke, Nomenclatural Notes ES 135

glabris, ad basin rotundatis, pli-nervatis; inflorescentiis
peniculatis.

Large woody vine, mostly with hanging branches; branches
slender, glabrous; leaves alternate; petioles slender, 4--
6.3 cm. long, glabrous, sulcate, incrassate and abruptly
curved at base; blades coriaceous, bright-green and shiny on
both surfaces, ovate, 7--13 cm. long, 2.7--6.5 cm. wide,
long-acuminate at the apex, entire, rounded at the base,
glabrous on both surfaces; venation pli-nerved, 2 secondar-
ies issuing at the very base of the blade, the other 2 issu-
ing 5--6 mm: above the base, prominent on both surfaces;
tertiaries and veinlet reticulation abundant, very slender,
prominulous on both surfaces; staminate inflorescence panic-
ulate, to 15 cm. long, many-branched, glabrous throughout;
flowers greenish-yellow; prophylla 3, narrow-lanceolate, a-
bout O.7 mm. long, sharply acute at apex, glabrous, membran-
ous-margined, alternate with the outer sepals; sepals 6, the
outer 3 elliptic, about 2.6 mm. long and 2 mm. wide, thick,
convex on the back, rounded at apex, glabrous, scarious-
margined, the inner 4 elliptic-rotund, very convex on the
back, about 2.8 mm. long and 2.2 mm. wide, cupped at the
base with inflexed margins, subacutely hooded at the apex,
glabrous,’ scarious-margined; petals 6, spatulate, whitish,
delicate, about 1.5 mm. long and 1 mm. wide, each enfolding
one stamen when mature, shallowly bilobed at the apex, glab-
rous; stamens 6, about 1.3 mm. long, separate; pistillate
flowers and fruit not seen.

The type of this species was collected by José Cuatrecas-
as (no. 16959) in the neighborhood of Palestina, alt. 5--50
m.e, on the Rfo San Juan, Choco, Colombia, between March 12
and 14, 1944, and is deposited in the Britton Herbarium at
the New York Botanical Garden.

JUNELLIA CONNATIBRACTEATA f. GLOMERATA (Monticelli) Molden-
ke, combe nov.
Verbena connatibracteata f. glomerata Monticelli, Lilloa
3: 358. 1938.

JUNELLIA CONNATIBRACTEATA f. ROSULATA (Monticelli) Moldenke,
comb. nov.
Verbena connatibracteata f. rosulata Monticelli, Lilloa ~

3: 558. 1938.

JUNELLIA LAVANDULIFOLIA var. COLCHAGUENSIS (R. A. Phil.)
Moldenke, comb. nov.
Verbena colchaguensis R. A. Phil., Anal. Univ. Chile 1873
521. 1873.

JUNELLIA ROSULATA Moldenke, sp. nov.

136 PHYTOLOGIA Vol. 2, no. 4

Suffrutex pumilus procumbens; ramis numerosis brevibus
procumbentibus radiatis; ramulis numerosissimis brevissimis
dense rosulatis; internodiis perabbreviatis dense foliatis;
foliis sessilibus rigidis tripartitis utrinque leviter pub-
escentibus, ad basin ampliatis amplexicaulibus, lobis linea-
ri-oblongatis obtusis revolutis; corollis rubellis.

Dwarf procumbent subshrub with heavy woody stems and num-
erous short procumbent radiating branches; branchlets and
twigs very numerous, very short, densely rosulate, the ster-
ile ones with extremely abbreviated internodes, very densely
leafy, forming a solid moss-like mat or cushion, the fertile
branchlets more elongate, to 5 cm. long, rather densely
short-pubescent, with internodes elongated to 8 mm., often
more abbreviated; nodes slightly ampliate, annulate by the
practically contiguous leaf-bases; leaves sessile, rigid,
opposite, 3-parted to the slightly ampliate and clasping
base, lightly pubescent on both surfaces with uniform, whit-
ish, forward-pointing hairs, the lobes practically equal,
linear-oblong, about 4 mm. long and 1 mm. wide (on the fert-
ile branchlets), obtuse or rounded at apex, the margins rev-
olute, much smaller on the sterile branchlets, bright-green
on both surfaces, the lower ones hidden from the light by
the densely matted upper ones but persistent, gray, brown,
or black and dry; peduncles abbreviated, to 1 cm. long,
densely short-pubescent with spreading white hairs; inflor-
escence terminal, densely many-flowered; rachis densely
whitish-pubescent; bractlets ovate-lanceolate, 5--6 mm.
long, 2 mm. wide at base, attenuate to the acute or acumin-
ate apex, strongly costate on the back, rather densely
whitish-pubescent with spreading hairs; calyx tubular, 6--8
mm. Long, densely short-pubescent with spreading white
hairs, 5-ribbed, the rim shortly 5-toothed; corolla pink,
conspicuously exserted, its tube to 10 mm. long, slightly
curvate, densely short-pubescent with spreading whitish
hairs outside, its limb 5-parted, its lobes 2--3 mm. long,
more or less short-pubescent outside, glabrous within.

The type of this remarkable species was collected by my
good friend and colleague, Teodoro Meyer (no. 9586), at
Tecka, Chubut, Argentina, on December 25, 1945, and is dep-
osited in the Britton Herbarium at the New York Botanical
Garden. The species is apparently related to J. minutifolia
(R. A. Phil.) Moldenke, but differs in its densely rosulate
habit and larger leaves. It may also be closely related to
J." Struthionum (Speg.) Moldenke, a species known to me only
from the original description.

JUNELLIA ROSULATA f. ALBA Moldenke, f. nov.
Heec forma a forma typica speciei corollis albis recedit.
-- This form differs from the typical form of the species in

= hy
et

mt
+

1946 Moldenke, Nomenclatural Notes 137

having white corollas.

The type of this form was collected by Teodoro Meyer (no.
9587) at Estancia "La Mimosa", Chubut, Argentina, on Decem-
ber 25, 1945, and is deposited in the Britton Herbarium at
the New York Botanical Garden.

JUNSLLIA TRIDACTYLA (R. A. Phil.) Moldenke, comb. nov.
Verbena tridactyle R. A» Phil., Anal. Mus. Nac. Chile
Bot. 1891: 59. 1891.

LANTANA HINTONI Moldenke, sp. nov.

Frutex; ramis ramulisque gracillimis obtuse tetragonis
brunnescentibus parce strigillosis; petiolis gracilibus
strigoso-pubescentibus; laminis subchartaceis ovatis longe
acuminatis serratis, ad basin acutis, supra strigilloso-
scabridis, subtus strigillosis vel puberulis densissime
punctatis; inflorescentiis axillaribus; bracteis ovatis.

Shrub; branches and branchlets very slender, obtusely
tetragonal, brunnescent, rather sparsely strigillose-pilosu-
lous; principal internodes 4--6.5 cm. long; leaves decuss-
ate-opposite; petioles slender, 4--8 mm. long, rather abund-
antly strigose-pubescent with appressed whitish hairs;
blades subchartaceous, uniformly bright-green on both surf-
aces, ovate, 4--7 cm. long, 2--4 cm. wide, long-acuminate at
apex, acute at base, rather coarsely but regularly serrate
except at the very apex and base, strigillose and scabridous
above, strigillose and very densely punctate beneath or pub-
erulent; midrib slender, plane above, prominulent beneath;
secondaries slender, 3 or 4 per side, arcuate-ascending, not
at all pli-nerved, plane above, subprominulous beneath, usu-
ally rather densely puberulent beneath; vein and veinlet re-
ticulation abundant, but apparent only under a hand-lens;
inflorescence axillary, 1 or 2 per node, shorter than the
subtending leaf; peduncles very slender, 2.5--4 cm. long,
strigillose; heads many-flowered, densely capitate, about 1
om. long and wide, not elongating after anthesis; bracts o-
vate, the lowest to almost 1 cm. long and 6 mm. wide; acum-
inate at apex, strigose-puberulent and densely punctate;
corolla-tube about 5 mm. long, its limb about 3 m. wide,
the outer portion of the limb and the exserted portion of
the tube densely puberulent.

The type of this species was collected by George B.
Hinton (no. 9307) at I. R. F. Placeres, alt. 1050 m., Zihua-
queo, Mina district, Guerrero, Mexico, on August 21, 1936,
and is deposited in the Britton Herbarium at the New York
Botanical Garden. It was first identified by me and distrib-
uted as L. canescens H.B.K., and later as L. Langlassei Mol-
denke, from both of which species it proves to be abundantly
distinct. The collector records the vernacular name

138 PHYTOLOGIA Vol. 2, no. 4
"toronjil."

LANTANA MINASENSIS Moldenke, sp. nov.

Frutex erectus; ramis gracilibus obtuse tetragonis parce
aculeolatis vel inermibus stramineis obscure pulverulento-
puberulis vel glabrescentibus; ramulis acutiore tetragonis
inermibus pulverulento-puberulis; nodis annulatis; petiolis
gracillimis obscure puberulis vel glabrescentibus; laminis
chartaceis griseo-viridibus lanceolatis vel lanceolato-ell-
ipticis longe acuminatis serrulatis, ad -basin acutis vel
plerumque acuminatis, supra scabris, subtus leviter puberu-
lis; inflorescentiis axillaribus.

Erect shrub, freely branching; branches rather slender,
obtusely tetragonal, sparingly aculeolate or unarmed, stram-
ineous, very finely and obscurely pulverulent-puberulent or
glabrescent; twigs more acutely tetragonal, unarmed, pulver-
ulent-puberulent; nodes annulate; principal internodes 1--3
cm. long; leaves decussate-opposite, numerous; petioles very
slender, 3--7 mm. long, very obscurely puberulent or glab-
rescent; blades chartaceous, gray-green, lanceolate or lan-
ceolate-elliptic, 3--7 cm. long, 1--3 cm. wide, rather long-
acuminate at apex, acute or usually acuminate at base, fine-
ly serrulate with blunt appressed teeth from apex to base,
pronouncedly scabrous above, lightly puberulent and not
scabrous beneath; midrib very slender, plane or slightly im-
pressed above, prominent beneath; secondaries very slender,
about 5 per side, arcuate-ascending, extending to the very
margins, plane or subimpressed above, prominulous beneath;
inflorescence axillary, usually borne only in the 2 or 3
uppermost axils of the twigs, shorter than the mature sub-
tending leaves; peduncles 1 or 2 per axil, very slender, 1.5
--2.5 em. long, tetragonal, minutely puberulous; heads many-
flowered, the canescent-puberulous rachis elongated to 1 cm.
during anthesis; bractlets lanceolate-ovate, about 2 mm.
long, 1 mm. wide at base, densely canescent-puberulous,
sharply acute at the apex; calyx about O./7 mm. long, canes-
cent-puberulous outside; corolla bright-pink, its tube very
slender, about 6 mm. long, pulverulent-puberulent outside,
its limb about 3 mm. wide.

The type of this species was collected by Ynes Mexia (no.
44488) in openings of cut-over woods, alt. 690 m., at about
km. 2 along the road from Vigosa to Barroso, Minas Geraes,
Brazil, on March 9, 1930, and is deposited in the Britton
Herbarium at the New York Botanical Garden.

MALVA MOSCHATA f. ALBA Moldenke, Am. Midl. Naturalist 35:
336, hyponym (1946), f, nov.
Haec forma a forma typica speciei corollis albis recedit.
-- This form differs from the typical form of the species in

|

ee ee

1946 Moldenke, Nomenclatural Notes 139

having white corollas. The type was collected by H. N. Mol-
denke (no. 15166) in a grassy field about 2 miles west of
Warren, Warren ren County, Pennsylvania, on July 1, 1943, and is
deposited in the herbarium of the Carnegie Museum at Pitts-
burgh. The form is quite common in this locality.

MARIPA CUATRECASASI Moldenke, sp. nov.

Frutex scandens; ramis ramulisque glabris; petiolis cras-
sis curvatis giabris; laminis coriaceis ellipticis magnis
longis acuminatis integris, ad basin rotundatis vel obtusis,
utrinque glabris; inflorescentiis axillaribus corymbiformi-
bus paucifloris; pedunculis fructiferis crassis glabris;
pedicellis fructiferis glabris; calyce fructifero glabro.

large liana; stems and branches glabrous; internodes ap-
parently quite variable in length; leaves alternate; petio-
les stout, 2--2.5 cm. long, curved, glabrous, wrinkled in
drying; blades coriaceous, clear-green on both surfaces, el-
liptic, 22--25 cm. long, 8.5--9 cm. wide, acuminate at apex,
entire, rounded or obtuse at base, glabrous on both surf-
aces, not particularly shiny; midrib plane or very obscurely
impressed above, very prominent beneath; secondaries slend-
er, 7 or 8 per side, arcuate-ascending, not reaching the
margins, mostly arcuate-joined about 5 mm. from the margins,
plane above, prominent beneath; veinlet reticulation rather
sparse, indiscernible above, only the largest portions dis-
cernible beneath; inflorescence axillary, corymbiform, few-
flowered; flowers not seen; fruiting-peduncles very stout, 6
--7 cm. long, glabrous, its branches few, 1--1.5 om. long,
glabrous; fruiting-pedicels stout, 1--1.5 cm. long, glab-
rous; fruiting-calyx incrassate, glabrous throughout, the
sepals suborbicular, 5--6 mm. long and wide, rounded at a-
pex; fruit elliptic, 3.5--4 cm. long, 2--2.5 cm. wide, yel-
low, acuminate at apex and base, its pericarp rather tough,
enclosing 2 seeds in a dark sweet gelatinous endocarp.

The type of this species was collected by José Cuatrecas-
as (no. 16531) at La Trojita, alt. 5--50 m., Rfo Calima, in
the region of the Choco, El Valle, Colombia, between Febru-
ary 19 and March 10, 1944, and is deposited in the Britton
Herbarium at the New York. Botanical Garden. The species is
quite distinct, differing at once from all the species of
the genus listed in Martius, Flora Brasiliensis 7: 205--210
(1871) by being a large liana with a perfectly glabrous
calyx, and from all the species listed by Gleason in the
Bulletin of the Torrey Botanical Club: 56: 107--111 (1929) by
its axillary inflorescences, very large non-cuneate leaves,
and lack of pubescence. The generic determination was made
by Joseph Monachino.

_MEGAPTERIUM BRACHYCARFUM var. WRIGHTII (A. Gray) Moldenke,

140 PRY Ono Fa Vol. 2, no. 4

comb. nov.
Oenothera wrightii A. Gray, Pl. Wright. 2: 57. 1853.

PAEPALANTHUS BRITTONI Moldenke, Known Geogr. Distrib. Erio-
caul. 5 & 45, hyponym (1946), nom. nov.
Paepalanthus montanus (Britton) Moldenke, Rev. Sudam.
Bot. 4: 17. 1937 [not P. montanus Alv. Silv., Fl. Montium

76. 1928].

PAEPALANTHUS COUTOENSIS Moldenke, Known Geogr. Distrib. Eri-
ocaul. 11 & 47, hyponym (1946), nom. nov.
Paepalanthus barbulatus Alv. Silv., Fl. Montium 211, pl.
139 & 140. 1928 [not P. barbulatus Herzog in Fedde, Repert.
20: 83. 1924]. ;

PAEPALANTHUS GLEASONII Moldenke, Known Geogr. Distrib. Erio-
caul. 6 & 49, hyponym (1946), nom. nov.
Paepalanthus robustus Gleason, Bull. Torrey Bot. Club 58:
330. 1931 [not P. robustus Alv. Silv., Flor. Serr. Mineiras
53. 1908]. =e a Bag

PAEPALANTHUS NEOCALDENSIS Moldenke, Known Geogr. Distrib.
Eriocaul. 14 & 51, hyponym (1946), nom. nov.
Paepalanthus caldensis Alv. Silv., Fl. Montium 186, pl.

120. 1928 [not FP. caldensis Malme, Bihang till K. Sv. Vet.

Akad. Handl. 27, Afd. 3, no. 11: 29, pl. 2, fig. 3. 1901].

PAEPALANTHUS NEOFPULVINATUS Moldenke, Known Geogr. Distrib.
Eriocaul. 51 & 61, hyponym (1946), nom. nov.
Peepalanthus pulvinatus Alv. Silv., Fl. Montium 37, pl.

18. 1928 [not P. pulvinatus N. E. Br. in Thiselton-Dyer, Fl.
Trope Afr. 8: 263. 1902].

PHYLA YUCATANA Moldenke, sp. nov.

Herba procumbens; ramis gracilibus plerumque sulcatis
canescento-strigillosis; petiolis plerumque alatis obscure
canescento-strigillosis vel glabrescentibus; laminis ovatis
vel ovato-ellipticis viridibus valde dentatis, ad apicem ro-
tundatis vel acutis, ad basin acuminatis, utrinque dense
canescento-strigillosis. .

Procumbent herb, freely branched from the base; branches
slender, rooting at the nodes, obtusely and rather irregu-
larly tetragonal, often deeply and irregularly» sulcate, of-
ten reddish or purplish toward the base, canescent-strigill-
ose with closely appressed antrorse hairs, the tips ascend-
ing or erect; secondary branches more slender, stramineous,
erect; nodes annulate; principal internodes 1--5 cm. long;
leaves decussate-opposite, numerous; petioles 1--5 mm. long,
mostly winged and merging into the base of the blade, rather

1946 Moldenke, Nomenclatural Notes 141

obscurely canescent-strigillose like the twigs or becoming
glabrescent; blades ovate or ovate-elliptic, mostly conspic-
uously widest below the middle, uniformly green on both sur-
faces, 1.5--4 cm. long, 0.5--2 cm. wide, rounded or acute
(in outline) at apex, acuminate at base and prolonged into
the petiole, the margins conspicuously and regularly dentate
with sharply acute or apiculate broadly triangular rather
divergent teeth from the apex to the widest part, the marg-
ins of the teeth rather thick and often more or less invo-
lute, both surfaces rather densely canescent-strigillose
with short closely appressed hairs usually visible only mic-
roscopically; midrib slender, plane above, very strong and
prominent beneath; secondaries slender, 4--6 per side, plane
above, very strong and prominent beneath, extending conspic-
uously to the leaf-margin and ending in the sinus between
two teeth, often with 1 or 2 short branches issuing almost
at the apex and extending to the apiculation of the nearest
tooth; tertiaries and veinlet reticulation not visible; in
drying, the leaves become almost plicate; inflorescence ax-
illary, capitate; peduncles slender, 2.5--5.5 cm. long, usu-
ally only one per node, deeply sulcate, rather densely can-
escent-strigillose or glabrescent; heads densely many-flow-
ered, 4--8 mm. long; bractlets ovate, about 3 mm. long, 1.5
mn. wide at the base, sharply acute at apex, densely canes-
cent-strigose, strongly costate; calyx minute; corolla about
3 mm. long in all, its limb about 1.5 mm. wide.

The type of this species was collected by Percy Gentle
(C. L. Lundell 4780] along the Corozal-Pachacan road, Brit-
ish Honduras, on nm July 20, 1933, and is deposited in the
Britton Herbarium at the New York Botanical Garden. The spe-
cies was hitherto confused with P. nodiflora var. reptans
(HeB«K.) Moldenke, the type of which is Venezuelan.

PHYLA YUCATANA var. PARVIFOLIA Moldenke, var. nov.

Haec varietas a forma typica speciei omnibus partibus
minoribus et densiore canescento-strigosis recedit. -- This
variety differs from the typical form of the species in
being smaller in all its parts and more conspicuously canes-
cent-strigose throughout. The stems arid branches are often
slightly woody; the internodes are often reduced to 1 em. or
less; the petioles are obsolete or only 1--2 mm. long; the
leaf-blades are usually less than 1 cm. long and wide, with
the venation plainly impressed above and prominent beneath,
imparting a decided plicate appearance to the leaves, con-
spicuously canescent-strigose.

The type of this variety was collected by George B. Hin-
ton (no. 6024) on & llano at Mal Paso, Heutamo district,
Michoacan, Mexico, on May 9, 1934, and is deposited in the
Britton Herbarium at the New York Botanical Garden. The col-

142 Puy TU Loom = Vol. 2, no. 4
lector records the vernacular name “hierba de hormiga."

PRIVA GRANDIFLORA (Ort.) Moldenke, comb. nov.
Verbena grandiflora Ort., Hort. Matr. Dec. 2. 1797.

SVENHEDINIA TRUNCATA Moldenke, sp. nov.

Frutex vel arbor; ramulis crassiusculis glabris valde an-
nulatis medullosis; petiolis crassis glabris marginatis, ad
apicem biglanduliferis; laminis coriaceis ovatis magnis, ad
apicem obtusis, integris subundulatis, ad basin late trunca-
tis, utrinque glabris pernitidis; reticulo venularum valde
perspicue utrinque prominente.

Shrub or tree; branchlets rather stout, glabrous, plainly
annulate sat the nodes, the annulation confluent with the up-
per margin.of the petioles, very pithy; principal internodes
1--4.5 cm. long; leaves alternate; petioles stout, 5.5--8
cm. long, glabrous, flattened above, with two corky margins
which terminate in two closely adjacent obtuse glands at the
apex; bladés coriaceous, ovate, to about 16 cm. long and 13
cme wide, obtuse at apex, entire but slightly undulate along
the margins, broadly truncate at base, glabrous and very
shiny on both surfaces; midrib plane above, very prominent
beneath, branching into 12 or more seconderies per side and
‘very numerous tertisries from the midrib, the secondaries,
tertiaries, and veinlets forming a very abundant and beauti-
fully conspicuous prominent reticulum on both surfaces, the
reticulum equally prominent on both surfaces; inflorescence
not seen.

The type of this species was collected by Julian Acufia
Galé (no. 14069) at Alto Babiney, Sur del Turquino, Oriente,
Cuba, on on August 1, 1935, and is deposited in the herbarium
of the Estacion Experimental Agronomica at Santiago de las
Vegas. The species is obviously related to S. minor (Urb.)
Urb., which differs notably in it very variable but always
acute or attenuate leaf-blade bases.

SYMPHOREMACEAE Moldenke » nom. nov.
Symphoremacees Van Tieghem, Journ. de Bot. 12: 359--365.
1898.

SYNGONANTHUS MINUTULUS (Steud.) Moldenke, Known Geogr. Dis-
trib. Eriocaul. 18 & 37, hyponym (1946), comb. nov.
Eriocaulon minutulum Steud., Syn. Pl. Cyp. 2: 270. 1855.

TOXICODENDRON SUCCEDANEA (L.) Moldenke, comb. nov.
Rhus succedanea L., Mant. 2: 221. 1767.

TOXICODENDRON VERNICIFLUA (Stokes ) Moldenke, combe nove
Rhus verniciflua Stokes, Bot. Mat. Med. 2: 164. 1812.

— =“. > a /

1946 Moldenke, Nomenclatural Notes 143

URGINEA SCILLA f. RUBRA Moldenke, f. nov.

Haec forma a forma typica speciei bulbis rubris recedit.
-- This form differs from the typical form of the species in
having red bulbs. The two forms of the species are kept sep-
arate in the trade, where the species is widely handled as a
crude drug, and, according to my fried, Joseph Monachino,
natives of Sicily invariably distinguish them. Up to the.
present time, however, I cannot find that the red form has
ever received botanic recognition.

VARRONIA ACUNAE Moldenke, sp. nov.

Frutex; ramis ramulisque subgracilibus subangulato-sulca-
tis dense puberulo-furfuraceis, juventute adpresso-strigill-
osis; foliis alternis numerosis; petiolis subobsoletis vel
usque ad 2.5 mm. longis strigilloso-puberulis; foliis sub-
coriaceis anguste ellipticis argute acutis valde revolutis,
ad basin attenuato-acutis, supra obscure pilosis vel glab<=
rescentibus, subtus dense fulvo-strigillosis; inflorescent-
iis terminalibus capitatis.

Shrub; branches and branchlets rather slender, somewhat
angulate-sulcate, densely puberulent-furfuraceous with fulv-
ous hair, the younger ones more distinctly appressed-strig-
illose with incanous hair; leaf-scars elevated, corky-marg-
ined; leaves alternate, numerous; petioles to 2.5 mm. long
(or almost obsoiete on young leaves), rather densely strig-
illose-puberulent with fulvous appressed hair; leaf-blades
subcoriaceous, narrow-elliptic, 1--4 cm. long, 4--10 mm.
wide, sharply acute at apex, attenuate-acute at base, the
margins very pronouncedly revolute, smooth to touch above
and very obscurely scattered-pilose, glabrescent in age,
rather densely strigillose-puberulent with fulvous hair be-
neath; inflorescence terminal, capitate; peduncles similar
to the branchlets in texture, shape, and color, 3--4.5 cm.
long, densely appressed-puberulent or strigillose with more
or less fulvous hair, or slightly incanous toward the apex,
several sulcate; heads globose, about 1 cm. in diameter,
many-flowered; calyx campanulate or cupuliform, about 3 mm.
long, rather sparsely strigose, its rim distinctly 5-lobed,
each lobe terminating in a filiform twisted densely strigose
appendage about 4 mm. long, the many contorted appendages
very conspicuous in the flowering and fruiting heads; fruit
elliptic, about 4 mm. long and 1.8 mm. wide, beaked at the
apex, glabrous, somewhat reticulate.

The type of this species was collected by Julian Acufia
Galé (no. 12687) along the highway at Delta No. 1, Moa,
Oriente, Cuba, on April 17, 1945, and is santero in the
herbarium of the Estacion Experimental Agronomica at Santia-
go de las Vegas. The species is obviously related to V.
globosa Jacq., which has similar flower-heads, but entirely

144 PHYTOLOGIA Vol. 2, no. 4
different leaves.

VARRONIA CORIACEA Moldenke, sp. nov.

Frutex; ramulis gracilibus griseis dense sotu lone heeeaane
is, pilis ad basin bulbosis; internodiis valde abbreviatis;
petiolis gracilibus denge setuloso-hispidis; leminis subcor-
jiaceis ellipticis vel lanceolato-ellipticis, ad apicem ob-
tusis, ad basin rotundatis, irregulariter denticulatis
revolutis; supra juventute dense setuloso-hispidis, senectu-
te scabris, subtus setuloso-hispidis; costa venisque venu-
lisque valde supra impressis, subtus prominentibus; inflor-
escentiis terminalibus capitatis setuloso-hispidis.

Shrub; branchlets slender, gray, densely setulose-hispid,
with bulbous-based hairs. which, upon being rubbed off, leave
the branchlets conspicuously verruculose; principal inter-
nodes greatly abbreviated, usually 1--1.5 cm. long; leaf-
scars elevated on corky circular sterigmata; petioles slen-
der, about 1 cm. long, densely setulose-hispid; blades sub-
coriaceous, elliptic or lanceolate-elliptic, 5.5--5.5 cm.
long, 1./--3 cm. wide, obtuse at apex, rounded at base,
rather irregularly denticulate except at the base, but the
margins so much revolute that the teeth are not obvious in
the dried state, scabrous above with short white bulbous-
based hairs, setulose-hispid beneath (and above when young)
with longer more slender and not so plainly bulbous-based
hairs, the hairs on the larger venation more plainly bulb-
ous-besed; midrib deeply impressed above, very prominent be-
neath; secondaries and tertiaries deeply impressed above,
giving the leaf a very bullate aspect, prominent beneath;
inflorescence terminal, capitate, varying from slightly to
very densely setulose-hispid with long white spreading
hairs, densely many-flowered; peduncles slender, 1l--2 cm.
long, densely setulose-hispid; calyx campsnulate, about 6
mm. long, puberulent and also more or less setose, its rim
regularly 5-lobed, the lobes sharply acute and about 2 mm.
long; corolla exserted, glabrous, its limb about 1 cm. wide.

The type of this species was collected by my good friend
and colleague, Julidn Acufia Galé (no. 14120) [an isotype is
labelled "Van Herman 14120"] at Lengua de Pajaro, Mayari,
Oriente, OQuba, in March, 1943, and is deposited in the herb-
arium of the Estacion Experimental Agronomica at Santiago
de las Vegas. The species is apparently related to V. lima
Desv., which differs in its non-hispid branches and ~inflor-
escences, few-flowered heads, and leaf-blades acute or att-
enuate at the base, and to Varronia Grisebachii (Urb.) Mol-
denke [Cordia Grisebachii Urb., Symb. Ant. 4: 477. .1908],
which differs markedly in its leaf-blades being acute at the
base and in its non-hispid pubescence on branchlets, peti-
oles, and inflorescences.

1946 Moldenke, Nomenclatural Notes 145

VARRONIA MOENSIS Moldenke, sp. nov.

Frutex; ramis gracilibus griseis glabris; foliis alter-
nis; petiolis gracilibus glabris; laminis subcoriaceis lanc-
eolatis vel suboblanceolatis utrinque glabris nitidis brun-
nescentibus integris subrevolutis, ad apicem acutis vel
breviter acuminatis, ad basin longe attenuatis; inflorescen-
tiis axillaribus capitatis; pedunculis filiformibus valde e-
longatis nutantibus glabris.

Shrub; branches slender, gray, glabrous, marked with
scattered elevated lenticels; leaves alternate; petioles
slender, 2--5 mm. long, glabrous, flattened or slightly can-
aliculate above; leaf-blades subcoriaceous, lanceolate or
suboblanceolate, 1.2--3.8 cm. long, 4--9 mm. wide, glabrous
on both surfaces, shiny above, brunnescent in drying, acute
or short-acuminate at apex, entire and somewhat revolute a-
long the margins, long-attenuate at base; inflorescence ax-
illary, capitate; peduncles filiform, 1.5--4.5 cm. long,
mostly greatly elongated and nutant, glabrous, slightly amp-
liate at the apex; receptacle club-shaped, very sparsely
short-pilose. .

The type of this species was collected by Julian Acufia
Galé (no. 12686) along the road at Delta No. 1, Moa, Oriente,
Cuba, on April 17, 1945, and is deposited in the herbarium
of the Estacion Experimental Agronomica at Santiago de las
Vegas. The species is apparently related to ¥. longipeduncu-
lata Britton & P. Wils., from which it may «st once be dis-
tinguished by its much narrower leaf-blades and its filiform
nutant peduncles.

XVERBENA BEALEI Moldenke, nom. nov.
Verbena hispida x litoralis Dermen, Cytologia 7: 164,
169, 170, 171, & 175~- 1936.

X VERBENA BINGENENSIS Moldenke, hybr. nov.

Herba mediocriter magna, ut videtur hybrida naturalis;
ramis sublignosis obtuse tetragonis, juventute pubescenti-
bus; ramulis argute tetragonis dense albido-pubescentibus;
petiolis alatis brevibus; laminis firme chartaceis fragili-
bus ovatis irregulariter inciso-laciniatis utrinque substri-
goso-pubescentibus, supra scaberrimis; inflorescentiis sim-
plicibus vel pauce ramosis axillaribus.

Medium-sized herb, apparently a natural hybrid between
V. bracteata Lag. & Rodr. and V. lasiostachys var. septen-
trionalis Moldenke; stems medium, slightly woody at the
base, obtusely tetragonal, brown, often blotched, rsether a-
bundantly pubescent with soft weak white hairs about 1 m.
long, wearing off in age; nodes annulate; principal inter-
nodes short, 2.5--5 cm. long; branches numerous, slender,
more sharply tetragonal and more densely white-pubescent

146 PAY 7.0 BOA Vol. 2, now 4

with soft hairs; leaves decussate-opposite; petioles short,
2--5 mm. long, winged, grading imperceptibly into the blade;
blades firmly chartaceous, brittle in drying, very rough a-
bove (when the finger is drawn downwards) and slightly so
beneath, ovate in outline, to 5 cm. long, mostly about l-- —
1.5 cm. wide, the largest to 3 cm. wide at base, irregularly
incised-laciniate, the two lowest laciniae often lobe-like
and spreading on the largest leaves, abundantly substrigose-
pubescent on both surfaces, the hairs very variable in
length beneath and densest on the venation; venation impres-
sed above, prominent beneath; inflorescence abundant, simple
or few-branched, in the axils of all the upper leaves; ped=-
uncles slender, 1--5 cm. long, sharply tetragonal, densely
short-pubescent; rachis slender, densely pubescent with
rather stiff forward-pointing white hairs of various lengths
spikes to about 10 cm. long, densely many-flowered, appar-
ently setting seed very poorly, the mature calyxes rather
distant; brectlets large and conspicuous, very variable in
size, 4--8 mm. long, lanceolate, the lowermost slightly fol-
iaceous, attenuate-acuminate, about 1 mm. wide at base, more
or less strigose, not keeled except when very old, mostly
greatly exceeding the calyx; calyx 3--4 mm. long, densely
‘strigose; corolla-tube slightly. surpassing the calyx, dense-
ly puberulent outside, its limb about 2 mm. wide.

The type of this natural hybrid was collected by Wilhelm
Ne Suksdorf in bottomlands near Bingen, Klickitat County,
Washington, on July 9, 1898, and is deposited in the Britton
Herbarium at the New York Botanical Garden.

VERBENA BRACTEATA f. ALBIFLORA (Cockerell) Moldenke, comb.
nove
Verbena bracteosa f. albiflora Cockerell in Daniels, Fl.
Boulder Colo. 204. 1911.

XK VERBENA COVASII Moldenke, nom. nov.
Glandularia santiaguensis x laciniata Schnack & Covas,
Darwiniana 7: 74. 1945; Rev. Argent. Agronom. 12: 228. 1945.

VERBENA CUMINGII Moldenke, sp. nov.

Herba; ramis procumbentibus stramineis tetragonis albo-
hirsutis; petiolis late alatis; laminis valde dissectis vel
bipinnatifidis utrinque strigosis, lobis subacutis revolutis
tenuiter chartaceis; inflorescentiis solitariis toerminali-
bus; pedunculis elongatis rectis hirsutulis.

Herb; stems apparently procumbent, ascending at the tips,
stramineous, tetragonal, hirsute with long white spreading
or reflexed hairs; principal internodes 2--3.5 cm. long;
nodes annulate, densely hirsute; leaves decussate-opposite,
usually with an abbreviated branch in each axil, the branch

1946 Moldenke, Nomenclatural Notes 147

very leafy and very hirsute; petioles about 5 mm. long or
slightly longer, broadly winged and indistinguishable from
the lamina; blades deeply dissected or bipinnatifid, strig-
ose on both surfaces, to 4 cm. long, the lobes subacute, the
margins slightly revolute, uniformly green on both surfaces,
thin-chartaceous; midrib and secondaries very slender, ob-
scure above, prominulous beneath; inflorescence solitary at
the end of each stem; peduncles erect, elongate, 15--17 cm.
long, rather more sparsely hirsutulous than the stems but
similar in color and texture, terminated by a pair of oppo-
site flowers with their subtending bracts and, about 1 cm.
beyond them, a dense head of flowers; bractlets lanceolate,
8--9 mm. long, about 1 mm. wide at the base, long-attenuate
at apex, hirsute-ciliate along the lower margins and short-
strigillose on the back and toward the apex on the margins;
calyx tubular, 10--11 mm. long (including the teeth), stri-
gose with appressed whitish hairs of several lengths, its
rim irregularly S-toothed, the teeth long-attenuate, 1--2
mm. long; corolla large, showy, its tube projecting about
2--3 mm. beyond the calyx, smooth outside, its limb about 12
mm. wide, the lobes deeply bilobed at apex; anther-append-
ages large, black, conspicuously exserted.

_ The type of this species was collected by Hugh Cuming
(no. 908) somewhere in Chile, sent to the Martius Herbarium
in 1835, now deposited in the herbarium of the Jardin Botan-
ique de 1'Etat at Brussels.

VERBENA CURTISII Moldenke, sp. nov.

Herba; ramis tetragonis stramineis leviter strigillosis;
petiolis gracilibus parce pilosis; laminis leviter chartace-
is lanceolatis acutis vel subacutis regulariter serratis, ad
basin attenuatis, utrinque strigillosis; spicis gracilibus
numerosis multifloris elongatis.

Herb; stems tetragonal, stramineous, lightly strigillose;
nodes annulate, with a transverse band of short white hairs;
leaves decussate-opposite, only the upper ones in and just
-below the inflorescence seen; petioles slender, 1--10 mm.
long, sparsely scattered-pilose with short white hairs;
blades thin-chartaceous, light-green, lanceolate, 1.5--4 cm.
long, 7--20 mm. wide, acute or subacute at apex, attenuate
into the petiole at base, rather regularly serrate from the
widest part to the apex with blunt or acute broadly triangu-
lar forward-pointing teeth, scattered-strigillose on both
surfaces. with short appressed whitish hairs, more densely so
along the venation beneath; inflorescence abundant; spikes
slender, numerous, two from each upper node, three at the a-
pex of each branch, 4--15 cm. long, rather'densely flowered
and fruited, often branched toward the. base, the branches
subtended by reduced leaves; peduncles slender, usually ab-

148 PHYTOLOGIA Vol. 2, now 4

breviated to about 1 cm., lightly strigillose; bractlets
lanceolate, about 2 mm. long, long-attenuate at apex, cilia-
te-margined, otherwise glabrate, persistent after the fruit
and fruiting-calyx have fallen; fruiting-calyx about 2 mm.
long, strigillose, especially toward the apex, the teeth
coming together over the apex of the fruit and forming a
distinctly sharp point until the calyx is ruptured; fruit 1
mm. long, smooth, long-coherent; corolla very small, the
tube about 2 mm. long, the limb slightly exserted.

The type of this species was collected by Moses Ashley
Curtis somewhere in "Carolina" [probably North Carolina] and
is labelled "Verbena caroliana L." It is Herb. G. Geete no.
5702, now deposited in the herbarium of the Botaniska
Tridgard at Goteborg, Sweden. The species reminds one of V.
carolina L. in its general aspect, but differs markedly in
its minute pubescence. It also resembles V. urticifolia L.,
but its fruiting-calyxes are more like those of V. scabra ~
Vahl, from which its non-scabrous leaves at once distinguish
it. It differs from V. riparia Raf. and V. urticifolia in
its dense fruiting-spikes and from the former in its non-
lobed leaves. Its full fruiting-calyxes seem to preclude V.
Engelmannii Moldenke or any other hybrid origin.

XVSRBENA DE&RMENI Moldenke, nom. nov.
Verbena hispida x bonariensis Dermen, Cytologia 7: 164,
165, 170, T71, & 175. 1956.

VERBENA DISSECTA f. GLANDULIFERA (Sanzin) Moldenke, comb.
nOVe

Verbena erinoides var. glandulifera Sanzin, Anal. Soc.
Cientific. Argent. 68: 131, fig. 34b. 1919.

VERBENA HAYEKII Moldenke, nom. nov.
Verbena procumbens Hayek in Engl., Bot. Jahrb. 42: 163.
1908 [not V. procumbens Forsk., Fl. Aegypt. Arab. 10. 1775].

VERBENA HOOKERIANA (Covas & Schnack) Moldenke, comb. nov.
Glandularia Hookeriana Covas & Schnack, Rev. Argent. Ag-
ronom. 12: 57, fig. 1. 1945.

XVERBENA KONDAI Moldenke, nom. nov.
Verbena racemosa x erinoides Dermen, Cytologia 7: 163.

1936.
XVERBENA LECOCQI Moldenke, nom. nov.

Verbena hispida x prostrate Dermen, Cytologia 7: 170.
1936.

VERBENA MONACENSIS Moldenke, sp. nov.

1946 Moldenke, Nomenclatural Notes 149

Herba; ramis prostratis ramosis acute tetragonis pilosis;
ramulis fere submarginatis dense adpresseque pubescentibus;
petiolis alatis strigosis; laminis chartaceis valde trifidis
incisis, lobis obtusis ellipticis vel oblanceolatis utrinque
strigosis, marginibus subrevolutis; inflorescentiis solitar-
iis terminalibus dense multifloris senectute elongatis.

Herb; stems apparently prostrate, ascending toward the
tips, branched, sharply tetragonal, brownish, lightly and
irregularly pilose, less so in age; branches more sharply
tetragonal (almost submargined) and more densely appressed-
pubescent; nodes annulate; principal internodes 2--7 cm.
long; leaves decussate-opposite, often bearing abbreviated
and very leafy branches in their axils; petioles to 1 cm.
long, usually much shorter, winged and almost indistinguish-
able from the rachis of the lamina, strigose on both surf-
aces; blades uniformly green on both surfaces, chartaceous,
deeply trifid, the divisions again incised, the individual
lobes mostly obtuse at apex and elliptic or oblanceolate in
outline rather than linear or oblong, strigose on both sur-
faces, the margins slightly revolute, the midrib and second-
sries slender, obscure above, prominulous beneath; inflor-
escence solitary st the end of each stem and branch, at
first congested, later elongating to 4 cm. or more, densely
meny-flowered; peduncles slender, 1.5--6.5 cm. long, densely
strigose or appressed-pubescent, conspicuously tetragonal
like the branches; bractlets lanceolate, about 6 mm. long, 1
mme wide at the base, densely short-pubescent with subap-
pressed whitish hairs, densely white-ciliate along the marg-
ins, long-attenuate at apex; calyx tubular, 8--9 mm. long
(including the teeth), strigillose, 5-costate, its rim short
ly 5-toothed, the teeth triangular end usually less than 1
mm. long; corolla large, showy, its tube projecting about 5
mm. beyond the calyx, glabrous outside, its limb about 1 cm.
wide, the lobes shallowly bilobed at the apex; anther-appen-
dages not exserted.

The type of this species is a specimen from the Martius
Herbarium now deposited in the herbarium of the Jardin Bot-
anique de 1'Ztat at Brussels, said to have been collected
from cultivated plants at Munich, Germany.

VERBENA NOACKI Moldenke, nom. nov.
Verbena hispida x hastata Dermen, Cytologia 7: 170. 1936.

VERBENA PARODII (Covas & Schnack) Moldenke, comb. nov.
Glandularia Parodii Covas & Schnack, Rev. Argent. Agro-

nom. ll: 94, fig. 3. 1944.

VERBENA FPERAKII (Covas & Schnack) Moldenke, comb. nov.
Glandularia Ferakii Covas & Schnack, Rev. Argent. Agro-

150 PY TO} Ot Grdd Vol. 2, no. 4
nom. ll: 89, fig. 1. 1944.

VERBENA FERENNIS var. JOHNSTONI Moldenke, var. nov.

Haec varietas a forma typica speciei ubique dense hirtel-
lis recedit. -- This variety differs from the typical form
of the species in having its stems, leaves, and rachis den-
sely spreading-hirtellous, the leaves to 5 cm. long, the
lowermost often with several linear lobes.

The type of this variety was collected by L. R. Stanford,
K. L. Retherford, and R. D. Northceraft (no. 915) among var-
ied vegetation of large shrubs, small trees, and herbs, in
broad damp river-beds, alt. 1950 m., 12 km. northwest of
Falmillas, on the road to Miquihuana, Tamaulipas, Mexico, on
August 14, 1941, and is deposited in the Britton Herbarium
at the New York Botanical Garden. It was originally dis-
tributed as "Verbena Shrevei Johnston."

_ VERBENA RIGIDA var. REINECKII (Briq.) Moldenke, comb. nov.
Verbena venosa var. Reineckii Briq., Ann. Conserv. &

Jard. Bot. Genev. 3: 164. 1899.

VERBENA SANTIAGUENSIS (Covas & Schnack) Moldenke, comb. nov.
Glandularia santiaguensis Covas & Schnack, Rev. Argent.
Agronom. ll: 92, fig. 2. 1944. |

xX VERBENA SCHNACKII Moldenke, nom. nov.

Glandularia peruviana x megapotamica Schneck & Covas,
Rev. Argent. Agronom. 12: 227--228, pl. 12, figs. 1--3.1945.

X VERBENA SUKSDORFI Moldenke, hybr. nov.

Herbe alta ut-videtur hybrida naturalis; ramis mediocri-
ter crassis tetragonis breviter pubescentibus scabris, pilis
ad basin bulbosis; petiolis late alatis; laminis crassiuscu-
lis firmis scaberrimis fragilibus valde trifidis vel lacini-
ato-incisis ovatis, ad basin saepe 2-lobatis, marginibus
revolutis, utrinque breviter pubescentibus, pilis ad basin
bulbosis; spicis perelongatis dense multifloris.

Tall herb, apparently a natural hybrid between V. offic-
inalis L. and V. lasiostachys var. septentrionalis Moldenke;
stems rather stout, obtusely tetragonal below, sharply ang-
led above, light-colored, rather abundantly short-pubescent
toward the base with stiff whitish often bulbous-based hair
less than 1 mm. long, less densely so towerd the apex and on
the branches, peduncles, and rachis, but sufficiently abund-
ant throughout to impart a very rough and harsh feel to
these organs especially when the finger is moved downwards,
the uppermost portions of the stem with margined angles, the
stems, branches, and peduncles also very densely fine-puber-
ulent with microscopic (almost punctiform) hairs, the rachis

——

aS!

1946 Moldenke, Nomenclatural Notes 151

more conspicuously puberulent with longer whitish forward-
pointing hairs; principal internodes mostly elongate, 3--8
em. long; leaves decussate-opposite, usually with clusters
of smaller ones on greatly abbreviated branches in their ax-
ils; nodes plainly annulate; petioles 1--2 cm. long, broadly
winged and not plainly distinguishable from the blades into
which they grade; blades rather thick and firm, very harsh
to touch on both surfaces, very brittle in drying, to about
9 cm. long, varying from deeply 3-fid to undivided and
merely laciniate-incised along the margins, ovate in out-
line, the two basal lobes on the largest leaves usually
wide-spreading, giving the blade a4 width of 5.5 cm. at the
base, incised-laciniate, the margins more or less revolute,
more or less densely short-pubescent on both surfaces with
stiff forward-pointing bulbous-based hairs, more densely and
lastingly so on the venation beneath; inflorescence copious-
ly branched at the apex of the stem; spikes greatly elong-
ate, usually 15--4O cm. long, densely many-flowered, but ap-
parently not setting fruit; peduncles mostly abbreviated,
acutely tetragonal, margined; brectlets ovate, about 2 mm.
long, O.5--1 mm. wide at base, ciliate-margined to the apex,
puberulent on the back, acuminate, keeled; calyx slightly
longer than the subtending bractlet, about 3 mm. long,
whitish-strigose; corolla-tube equalling the calyx, its limb
about 2 mm. wide.

The type of this remarkable plant was collected by Wil-
helm N. Suksdorf in a garden at Bingen, Klickitat County,
Washington, on November 21, 1904, and is deposited in the
herbarium of the State College of Washington at Pullman. The
plant seems definitely to be a hybrid because the greatly e-
longated spikes in the only two specimens seen contain
thousands of mature calyxes, but not a single fruit. The
other described natural hybrids in the genus are also noted
for their paucity in fruiting, but none has the lack of
fruit as complete as this one!

VERONICA SERPYLLIFOLIA f. ALBIFLORA Moldenke, Am. Midl.

Naturalist 35: 376, hyponym (1946), f£. nov.

Haec forma a forma typica speciei corollis albis recedit.
-- This form differs from the typical form of the species in
having pure-white corollas. The type was collected by H. N.
Moldenke (no. 17293) in a field at Cranbrook Farm, North
Warren, Warren County, Fennsylvania, on May 8, 1945, and is
deposited in the herbarium of the Academy of Natural Scien-
ces at Philadelphia.

XCHRYSANTHEMUM CULTORUM Moldenke, nom. nov.
Chrysanthemum morifolium Ram. x C. sibiricum Fisch. ex
Rehd., Man. Cult. Trees & Shrubs, ed. 2, 862. 1940.

SUPPLEMENTARY NOTES ON THE ERIOCAULACEAE, AVICENNIACEAE, AND
VERBENACEAE OF TEXAS. II

Harold N. Moldenke

Since the publication of my discussion of these three
families in Lundell's “Flora of Texas", volume 3, part l,
pages 1--87 (1942) and the first in my series of supplement-
ary notes thereto in Phytologia 2: 123--128 (1945) nine hun-
dred and sixty-three additional Texan specimens have been
examined. These additional specimens have yielded several
scores of new county records and even three new species and
varieties for the state. This new material has come to me
from thirty-three herbaria, the abbreviations for which as
employed hereinafter are as follows: Al = New York State
Museum, Albany, New York; Au = University of Texas, Austin,
Texas; Br = Jardin Botanique de 1'Etat, Brussels, Belgium;
Bt = Butler University, Indianapolis, Indiana; Cm = Carnegie
Museum, Pittsburgh, Pennsylvania; Cn = University of Cin-
cinnati, Cincinnati, Ohio; Du = Dudley Herbarium, Leland
Stanford University, California; Fe = Colorado Agricultural
& Mechanical College, Fort Collins, Colorado; Ga = Georgia
Agricultural Experiment Station, Experiment, Georgia; Go =
Botaniska Tridgard, Géteborg, Sweden; H = Duke University,
Durham, North Carolina; Hp = H. Hapeman herbarium, Minden,
Nebraska; Hs = Crispus Attucks High School, Indianapolis,
Indiana; I = Langlois Herbarium, Catholic University of Am-
erica, Washington; It = Cornell University, Ithaca, New
York; Ka = Kansas State College, Manhattan, Kansas; Kr = B.
A. Krukoff herbarium, New York Botanical Garden, New York
City; Ky = University of Kentucky, Lexington, Kentucky; la ®
University of California at Los Angeles, Los Angeles, Calif-
ornia; Ll = Lloyd Library, Cincinnati, Ohio; Me # Instituto
de Biologia, Universidad Nacional de México, Mexico City; Ml
= Instituto Miguel Lillo, Tucuman, Argentina; N = Britton
Herbarium, New York Botanical Garden, New York City; Ok =
University of Oklahoma, Norman, Oklahoma; Pl = State College
of Washington, Pullman, Washington; Po = Pomona College,
Claremont, California; Pr = Princeton University herbarium,
New York Botanical Garden, New York City; Se = University of
Washington, Seattle, Washington; St = Oklahoma Agricultural
& Mechanical College, Stillwater, Oklahoma; Up = University
of Fennsylvania, Philadelphia, Pennsylvania; Vt = University
of Vermont, Burlington, Vermont; W = United States National
Herbarium, Smithsonian Institution, Washington; and We #
West Virginia University, Morgantown, West Virginia. I am
deeply grateful to the directors and curators of these herb-

152

1946 Moldenke, Supplementary Notes 153
aria Tor permitting me to examine and annotate this material.

ERIOCAULON COMPRESSUM Lam.

Hardin Co.: Ee Je Palmer 9563 (Du); Tharp sen. [July 20,
1929] (Au). Jefferson Co.: Hooks sen. [Beaumont, 5/3/1930]
(Au, N), sen. [Beaumont, 5/30/34] (Au).

ERIOCAULON DECANGULARE L.

Anderson Co.: LeSueur & Smith s.n. [7/7/35] (Au, N), sen.
17/7/38] (Au). Austin Co.: Tharp 44347 (N), 44348a (N), sen.
[6/28/42] (Al, Au). Freestone Co.: G. W. Goldsmith s.n. [6/
15/41] (Au, Au). Hardin Cos: Tharp sen. [July 20, 1929] (au,
Au), sen. [7-21-42] (Au). Henderson Co.: Tharp 2880 (Au).
Jasper Co.: G. L. Fisher 32101 (Au, Au). Jefferson Co.: Mrs.
Smith sen. [Beaumont, July '15] (Au). Newton Co.: Thar

(Au, N). Robertson Co.: F. A. Barkley 1340 (Al), l

N). Rusk Co.: Vinzent 47 (Br). Smith Co.: J. Reverchon 27

(Po), 4359a (Po). Tyler Co.: Tharp 44345 (Au, N). Waller Co.:
E. Hall 675 (Po).

a

ERIOCAULON KORNICKIANUM Van Heurck & Mill.-Arg.
Polk Co.: Tharp 42-6 (N), 42-7 (N).

ERIOCAULON TEXENSE Korn.

Austin Co.: Tharp sen. [5/4/40] (Au); Warnock 224 (N).
Leon Co.: F. A. Barkley 13556 (N). Milam Co.: Tharp 4434¢
(Au, N); 4540 (N), LASALb (N). Robertson Co.: F. A. Barkley
13543 (N); Painter & Barkley 13540 (N).

LACHNOCAULON ANCEPS (Walt.) Moron

Jasper Co.: Whitehouse s.n. [6/10/1951] (Au, Au, N). Jef-
ferson Co.: Hooks sen. [5/30/34] (Au), sen. [6/7/36] (Au).
ga Co.: Tharp 443546 (Au, N). Tyler Co.: Tharp 44343 (au,
N).

AYICENNIA NITIDA Jacq.

An additional vernacular name for the species from Texas
is "mangle negro", recorded by Runyon.

Cameron Co.: Parks 1724 (Au); R. Runyon 4031 (Au). Nueces
Co.: Tharp 8 eNe [July Bs 1939] (Au).

ALOYSIA LIGUSTRINA (Lag.) Small

Bexar Coe: Ge L. Fisher s.n. [San Antonio, July 11, 1921]
(Hp, Vt); Headley sen. [April 15, 1907] (1); Metz 62 (Hp, I,
Se), 65 (1), sen. [October 23, 1933] (I); H. B. Parks 2524
(Au). Brewster Co.: C. H. Mueller s.n. [July 12, 1932] (Au):
L. T. Murray sen. [Garden Springs, May 21, 1928] (It); Nels-
on & Nelson 5108 (Au); Warnock 66, in part (Au), 20675 (Au),
20689 (Au), W.288 (Au). Cameron Co.: R. Runyon s.n. [Browns-

154 Pn y TO ro Gran Vol. 2, no. 4

ville, 1930] (Hp). Comal Co.: Lindheimer 275 (Ka), 1070 (Br,
Me, Me, Me, Me). Concho Co.: J. Reverchon sen. [Curtiss
1965*] (I, Vt). Culberson Co.: U. T. Waterfall 4629 (N),
5091 (Au, N), 5458 (N). Dallas Co.: J. Reverchon sen. [Aug.
1877] (Vt). Frio Co.: Griffen & Barkley 13909 (Au). Hidalgo
Coe: Clover 10, in part (Ti Me). Hudspeth Co.: U. T. Water-
fall 4963 (N). Jeff Davis Co.: F. A. Barkley 141637 (Au);
Tracy & Barle 184 (Cm). Kleberg Co.: J. Fe Sinclair sen.
(Kingsville, Spring, 1940] (Au). La Salle Co.: Mauermann 2
(Au), 21 (Au). Llano Co.: Tharp sen. [8-17-40] (Fl). Mason
Co.: Nelson & Nelson 5195 (Au). Mitchell Co.: Tracy 8508
(Cm). Pecos Co.: Tharp 43-794 (Al, Au). Presidio Co.: Hitch-
cock & Stanford 6811 (Pl, Po, Se); M. S. Young sen. [Sept.
7, 1914] (Se). Travis Co.: CO. 0. Albers,32018 (Au); Fe. As
apes LO 2 (Au); Letterman 390 (Ka), sen- [Austin, July,

1882] (Al); R. H. Painter 29 (Ka); Tharp sen. [Austin, 7/25/
41] (Pl, Se). Val Verde Co.: W. H. Rhoades sen. [Delrio,
Aug. 1932] (Hs). County undetermined: A. S. Hitchcock s.n.

(Ka); Lindheimer 502 (Br, Ka).

ALOYSIA LIGUSTRINA var. SCHULZII (Standl. ) Moldenke
Brewster Co.: Warnock 66, in part (Au). Hidalgo Co.:
Clover 10, in part (Me). Pecos Co.: Hinckley sen. [June 30,
1941 J (Au); Tharp 435-793 (Au). Val Verde Co.: Cory 39092

(Au); Gs L. Fisher 32233 (Po); Munz 1446 (Po).

ALOYSIA MACROSTACHYA (Torr.) Moldenke

Hidalgo Co.: Clover 1075 (I); Mrs. E. J. Walker 20 (Au,
Au); T. R. Walker sen. [summer, 1938] (Au). Live Oak Co.:
Owens 1717 (Au); H. B. Parks 2043 (Au).

ALOYSIA WRIGHTII (A. Gray) Heller

Brewster Co.: Warnock 308 (Au, Au), 308b (Au), 20051
(Au), 20438 (Au), 21272 (Au). El Paso Co.: Shiner 40171
(Au). Jeff Davis Co.: M. S. Young sen. [Davis Mts., Aug. ll,
'14] (Se). Pecos Co.: Tharp 43-795 (Au), 2535 (H). Presidio
Co.: Hinckley s.n. [July 8, 1941] (Au).

BOUCHEA SPATHULATA Torr.

Brewster Co.: He C. Hanson 718 (Ka); Moore & Steyermark
3446 (Du). oe ee

CALLICARPA AMERICANA L.

Aransas Co.: Cory 45742 (Au). Bexar Co.: A. A. Heller
1832 (Se); Metz 273 (Se). Comal Co.: Lindheimer 297 (Ka),
1067 (Me, Me, Me). De Witt Co.: M. Riedel sen. (6-3-42]
(Au). Fayette Co.: Forshey s.n. [Rutersville, 1857] (Ka).
Gregg Co.: C. L. York sen. [Fall, 1937] (Au). Nacogdoches
Co.: Crausley sen. [July 18, 1944] (Au). Polk Co.: Girvin

e
7.

1946 Moldenke, Supplementary Notes 155

101 (Au). Robertson Co.: F. A. Barkley 13590 (Au). Travis
Co.: McKee & Wesley 3860 (Au); R. H. Painter sen, [Austin,
8/6/23] (Ka); Ripperton & Barkley 14524 (Au); Tharp 44416
(Au), sen. [Austin, 7-21-40] (Pl, Se); York & Wolf 46 (Au).
Trinity Co.: Goodrum s.n. [June, 1936] (Au). Washington Co.:
C. C. Albers 32022 (Au); Brackett 253 (Au), sen. [7/1/39]
(Au). Victoria Co.: Ferris & Duncan 3258 (Du).

CALLICARPA AMERICANA var. LACTEA F. J. Muller
Chambers Co.: G. L. Fisher s.n. [Anahuac, Sept. 18, 1931]
(Du).

CITHAREXYLUM BZRLANDIERI B. L. Robinson
Cameroh Co.: Clover 1237 (Du); Ferris & Duncan 3051 (Du);

G. L. Fisher 41195 (Hp); Owens & Parks 8.1715 (Au), R.1714

(Au); I. Shiller 659 (Au); Tharp 1652 (N). Willacy Co.:

Tharp 1227 (N), 1249 (N).

CLERODENDRUM FRAGRANS var. PLENIFLORUM Schau.
Gonzalez Co. (cultivated): Cory 29602 (Au).

CLERODENDRUM INDICUM (L.) Kuntze
Cultivated: Drushel, Tharp, & Barkley 13A163 (Au).

LANTANA CAMARA L.
Travis Co.: Warnock 11 (Au).

LANTANA CAMARA var. MISTA (L.) L. H. Bailey
Travis Co.: J. Le. White 47352 (Au).

LANTANA HORRIDA H-B.K.

Aransas Co.: Cory 453681 (Au), 45740 (Au). Bexar Co.: Metz
57 (Se); H. B. Parks 15596 (Kr). Cameron Co.: Re Runyon sen.
[TBrownsville, 1930] (Hp). Comal Co.: Lindheime 354 (Ka),
1068 (Me, Me, Me). De Witt Co.: M. Riedel sen. [7-27-41]
(Au). Gonzales Co.: F. A. Barkley 13882 (Au); Tharp sen.
{Ottine, 5/1/35] (St). Harris Co.: G. L. Fisher s.n. [Hous-
ton, May 16, 1916] (Hp). Hays Co.: Heald & Wolf 911 (Au);
Straudtmann sen. [San irra Apr. 23, 1937] (Au). Hidalgo
Co.: Walker & George 153 (Au). Medina Co.: Tharp sen. [Dev-
ine] (Au). Nueces hee Heller 1386 (Pl). Travis Co.:
C. C. Albers 32017 (Au); Harpin, Waldorf, & Barkley 13076
(Au); R. He Painter 85 (Ka); Tharp 44155 (Au), sen. (Austin,
5/9/35] (St). Victoria Co.: P. 0. Schallert 555, in part
(H). Washington Co.: C. C. Albers 34012 (Au); Brackett sen.
(July 1938] (Au). Willacy Co.: Tharp 1197 (Au). County unde-
termined: Nealley s.n. [s.w. Texas, 1888] (Fe).

|

LANTANA MACROPODA Torr.

156 PHY TOLOGIA Vol. 2, no. 4 ;

Mr. V. Le Cory offers the following valuable corrections
to my key to the Texan species of Lantana: "No. 455922 is
Lantana macropoda Torr. to me is correct; but from your key
im FLORA OF TEXAS I would not place it there. In that key
the separation from L. citrosa is unsatisfactory so far as
our material is concerned. 'Lesf-blades sharply serrate’ is
not true for the material of 55922, and it might stretch it~
some to be considered true for the material from Starr Coun-
ty (No. 35934), although there is a point to the teeth here
and this is lacking in the material from Zapata County and
on up the Rio Grande. I have written in my book this correc-
tion of your key 'lesf-blades broadly crenate to serrate’.
Now as to L. citrosa (No. 36729), the 'Leaf-blades finely
crenate or subentire' to me also is misleading, for the
leaves here to me are more nearly serrate than they are in
L. macropoda. However, there are other differences in the
leaves of these two species that would not be misleading."

Brewster Co.: C. He Muller 32016 (Au), sen. [Chisos Mts.,
7-17-32} (Au); Warnock 253 ae 831, in part (Au), sen.
(May 2, 1937] (Au, Au). Cameron Co.: Small & Wherry 11841
(Ml). Duval Co.: F. A. Barkley 13889 (Au). Hidalgo Co.: M.
L. Walker 105 (Au). Kinney Co.: He B. Parks PX.002 (Au).
Presidio Co.: Hinckley 1502 (Au, N). Reeves Co.:;Tharp 8852
(N). Starr Co.: Clover 1595 (I). Val Verde Co.: Gory 58096
(Au), 39745 (Au); G. L. Fisher s.n. [Davila River, July 14,
1927] (Hp)s Me. E. Jones 26218 (I); Rose-Innes & Moon 1292
(Au); Rose-Innes & Warnock 602 (Au). 3

LIPPIA ALBA (Mill.) N. E. Br.

A. De Je Meeuse in "Blumea", vol. 5, pp. 68--69 (1942)
claims that the correct name for this plant is L. javanica
(Burm. f.) Spreng. However, until I am able to examine the
type specimen and complete certain bibliographic inquiries
I am reserving judgment and am continuing to use Brown's
name»

Cameron Co.: A. M. Davis sen. [Palm Grove, Sept. '41]
(Au); G. L. Fisher s.n. [Brownsville, Aug. 16, 1924] (Hp),
sen. [Apr. 20, 1941] (Au, Au); R. Runyon 228 (N), sen.
[Browneville, 1930] (Hp). Hidalgo Co.: M. L. Walker 54 (Au)

LIPPIA GRAVEOLENS H.B.K.
The species is said by Hanson to frequent canyons.

(Au). Hidalgo Co.: I. Shiller 746 (Au); Mra. E. J. Walker

bd

1946 Moldenke, Supplementary Notes 157

PHYLA CUNEIFOLIA (Torr.) Greene
Mr. V. Le Cory has sent me valuable corrections to my key
to the members of this very complex group in Texas, for
which corrections as well as all his other helpfulness I am
deeply grateful. He says "In your key to the genus Phyla I
find that your key as it applies to our species P. cuneifol-
ia and P. incisa is misleading. All of our material accord-
ing to this would be referred to P. incisa, whereas three-
fourths or more of it is really P. cuneifolia. However, I
found that the New Mexico material of P. cuneifolia really
does have the shorter peduncles. As far as we are concerned,
if the separation was made on the bracts particularly then
one could not go wrong in following the key. This would make
the key foolproof."
Brewster Co.: Warnock 20736 (Au), 20737 (Au), 21204 (Au),

21277 (Au), sen. [duly 23, 1940] (Au). Crockett Co.: Co

fap? (Au), 3 39354 (Au). Culberson Co.: U. T. Waterfall 4685
N). Howard Co.: Tracy 7998 (Cm). Midland Co.: Cory GO61S
(Au). Olgham Co.: M. W. Howard 25 (Au). Potter id Ge Je
Goodman 3052 (Se). “Reagan Co.: Cory 12540 (Au). Scurry Co.:
Tharp sen- [7/9/41] (Au, N). Taylor Co.: Tolstead 7547

Herb. Texas Agr. Exp. Sta. 42543] (Au).

PHYLA INCISA Small

Bell Co.: Cohn 1.21 (Au); L. Mclean sen. [Temple, 8-30-
34] (St); J. F. Normand s.n. [1928] (N, N). Bexar Co.: Metz
OAT), 88. (I, 1), 159 (1), sen. [Aug- 10, 1931] (Se). Brazos
Co.: Chenault ceniedine 15, 1937] (Au). Brewster Co.: War-
nock 20435 (Au), 20676 (an); 20741 (Au). Brooks Co.: Tharp
BoNe [672 6/41 J Per Cameron Co.: Ae M. Davis sen. [Palm
Grove, Sept. '41] (Au). Comal Co.: Lindheimer 262 (Ka), 288
(Ka), 1069 (Me, Me), 1071 (Me, Me). Dallas Co.: + M. Ae Hynes
‘gone (Dallas, 6/2/26] (Au). El Paso Co.: Cory 45 4s0S8 (Aus Fe
W. Johnson 1707 (Go). Fayette Co.: Be We Crawford sen.
[Colony, June, 1892] (Ka). Gelveston Co.: G. L. Fisher sen.
[Galveston, Aug. 31, 1919] (H). Galveston or Harris Co.: B.
C. Smith sen. [between Houston and Galveston, 5-2-1942]
(Fe). Grayson Co.: Schleuse 36008 (Au). Gregg Co.: Ce Le
York s.n. [9-2-39] (Au), sen. [Aug. 28, 1941] (Au). Harris
Coe: Ge G. Le Fisher sen. [Houston, May 4, 1918] (Vt). Hudspeth
Co.: U. T. Waterfall 4589 (N, Pl). Jeckson Co.: Tharp sen.
[Aug. 28, 1941] Gia N), sen. [Sept. 3, 1941] (Au, Au, N).
Kerr Co.: A. A. Heller 1920 (Se). Kleberg Co.: J. F. Sin-
clair sen. [Kingsville, Spring 1940] (Au). Nueces Co. A. Ae
Heller 1806 (N--isotype, Fl--isotype, Se--isotype). Presidio
Co.: Hinckley sen. [San Esteban Lake] (Au), sen. Leivaes oe
ly, 1936] (Au). Reeves Co.: U. T. Waterfall 4361 (N),
(N). Refugio Co.: Tharp sen. [Austwell, 57-1689) a an °
Robertson Co.: F. A. Barkley 13003 (Au). Tarrant Co.: Let-

158 PRY BOL OG Tas Vol. 2, nOsy™

terman 391 (Ka); Ruth 91 (L1), 106 (Cm, La). Taylor Co.:
Tolstead 7550 [Herb. Texas Agr. Exp. Sta. 42551] (Au, Au).
Tom Green Co.: Cory 39600 (Au). Travis Co.: Cohn & Barkle
193 (Au), 13245 Au); A, M. Ferguson sen. {Waller ree,

Au); E. Hall 436 (Pr); Heald & Wolf sen. [Austin, 4-30-09]
(Au); Straudtmann sen. [July 17, 1940] (Au); Tharp 44138
(Au), 44188 (Au), sen. [Austin, 5-15-35] (St), sen. [Austin,
5/10/38] (Pl), sene [7-18-41] (Au, Au, N), sen. [August,
1941] (Au); Waldorf 19 (Au). Victoria Co.: P. 0. Schallert
550 (H). Washington Co.: C. C. Albers 33022 (Au); Brackett
20 (Au), 253 (N), sen. (Apr. 30, 1939] (Au). County undeter-
mined: Lindheimer s.n. (Ka).

PHYLA LANCEOLATA (Michx.) Greene

Lundell reports the corolla as white with an orange-
yellow eye or pink with a rose eye.

Anderson Coe: F. A. Barkley 13586 (Au). Austin Co.: Tharp
sen. [6/28/42] (Au). Dallas Co.: C. L. Lundell 11656 [Plant.
Exsicc. Gray. 1276] (Al, Au, H, I, Ka, N, Fl, St, We). Gon-
zales Co.: Straudtmann sen. [Aug. 12, 1940] (Au); Thar
44182 (Au), sen- [6-10-40] (Pl). Harris Co.: Boon 53 (Au).
Jackson Co.: Tharp sen. [Aug. 27, 1941] (Au, N). Lubbock Co.:
E. L. Reed 3827 (I).

FHYLA NODIFLORA (L.) ee '

Aransas Co.: Cory 45759 (Au). Galveston Co.: H. Hapeman
sen. (Galveston, May 10, '94] (Hp); Therp s.n. [5-1-37] (Fl).
Harris or Jefferson Co.: Crockett 7002 [between Beaumont and
Houston] (Au). Kerr Co.: H. R. Reed 45998 (Au). Nueces Co.:
C. Ce Albers 32021 (Au).

FHYLA NODIFLORA var. REPTANS (H.B.K.) Moldenke

The following collections previously reported and cited
by me as this variety are actually P. incisa Small: M. A.
Hynes sen. (Dallas, 6/2/26], =. Beck 58, Hinckley sen. [Mar-

fa, July, 1936], sen. [San Esteban Lake], and Tharp sen.
{Austwell, 9-7-1929]. The following collections previously
cited by me as P. nodiflora var. reptans are actually F.

yucetana Moldenke: Cory 28215, Mrs. P. Cottrell s.n. [San

1204. The following collections previously cited by me as P.
nodiflora var. reptans are actually F. yucatana var. parvi-
folia Moldenke: Clover 119, Berlandier 867, 2287, and Lund-
ell & Lundell 9922.

Cameron Co.: R. Runyon sen. [Brownsville, 1930] (Hp).
Chambers Co.: Tharp sen. (4--7/10/36] (N). Comanche Co.:

1946 Moldenke, Supplementary Notes 159

Lindheimer 1071 (Me, Me). Edwards Co.: Cory 37901 (Au). Tom
Green Co.: Cory 39321 (Au), 42826 (Au). Travis Co.: Ce. C.
Albers 32030 ue & Armer 5532 (Au); Tharp sen. [7/10/39] (N)}
Uvalde Co.: Cory 38212 (Au), = s3ho8 (Au), 44511 (au). Wichita
Co.: Tharp sens Ket 5-28-22 ] (N)s

FHYLA YUCATANA Moldenke

Cameron Co.: Mrs. P. Cottrell sen. [San Benito, Feb.-Apr.
1931] (Au); A. Me fs parle Sen. [Palm Grove, Sept. '41] (Au);
Ferris & Duncan 3091 (N); H. C. Hanson 508 (N); Lundell &
Lundell 10013 (N); = Runyon 350 (Au, N), 2688 (N), sen.
[Harlington, May 11, 1941] (N); Seventh Grade ade Brownsville
SeNe genta April 1934] (Au, N); Small & Wherry 11892
(N)s Tharp 1203 (Au, N). Hidalgo Co.: Cory 28215 (N); Mre.
E. J. Walker s.n- [la Joya, April 1942] (Au); M. L. Walker

7

PHYLA YUCATANA var. PARVIFOLIA Moldenke

Hidalgo Co.: Clover 119 (N). Fresidio Co.: U. T. Water-
fall 4782 (N). Starr Co.: Lundell & Lundell 9922 (N).
County undetermined: Berlandier 867 (T), 2287 (C).

PRIVA LAPPULACEA (L.) Pers.

Mr. L. I. Davis reports that this species "is fairly
common in certain parts of Hidelgo County."

Cameron Co.: Co = (Au); Ae Me Davis sen. [Palm
Grove, Sept. ‘41

TETRACLEA COULTERI var. ANGUSTIFOLIA (Woot. & Standl.) A.

Nels. & Macbr.

Culberson Co.: U. T. Waterfall 3765 (N), 4457 (N).
VERBENA AMBROSIFOLIA Rydb.

Brewster Co.: Cory 44804 (Au); Rose-Innes & Moon 1172
(Au); Warnock 20121 aye 20921 (Au), T.66, in part (Au),
W.283, in part (Au). ‘ce se Coe: U. Te “T. Waterfall 4458 (N,
Pl). Hudspeth Co.: Tharp 43-798 (Au); U. T. Waterfall 4895
(N). Jeff Davis Co.: Cory 40365 (Au); Warnock 21676 (Au); U.
T. Waterfall 4722 (N). Pecos Co.: Tharp 43-796 (Al, Au),

43-797 (Au).

VERBENA BIPINNATIFIDA Nutt.

Baylor Co.: Bridge sen. [near Seymour, 6/16] (Cn). Bexar
Co.: Barkley & Farsons 1 (Au); Lindheimer 10 (Ka); Metz 79
(I, Se). Brewster Co.: Warnock W.284 (Au). Burnet Co.: | Ge Ce
Albers 38003 (Au, Au); Warnock W.1096 (Au). Caldwell Co.:

Sos sen. [Dec. 30, 1956] (Au). Comal Co.: We H. Kel-
logg 8 (Au); Lindheimer 1072 (Me, Me, Me), 1973 (Me, Me, Me).
Culberson Co.: Hitchcock & S Stanford 6782 (Po). Dallas Co.:

160 PET Oho Gea Vol. 2, no» 4

E. Brainerd sen. [Dallas, March 28, 1908] (Vt); Lundell &
Lundell 11415 [Flant. Exsicc. Gray. 1275] (Al, Au, H, I, Ka,
N, Pl, St, We); Reverchon sen. [Curtiss 1962*] (Cm, I, Vt).
DeWitt Co.: M. Riedel sen. [7-18-41] (Au). Hastland Co.:
Hodge Oak School 18 (Au). Ellis Co.: Cory 39243 (Au). Fannin
Co.: McGart 2032 (Au). Gillespie Co.: G. Jermy 162 (Ka);
Mainland & Barkley 14522 (Au). Gonzales Co.: CO. C. Albers
35006 (Au). Grayson Co.: C. S. Sheldon s.n. [Denison, June
13, 1891] (Al). Harris Co.: E. Hall 435, in part (Pr).
Kimble Co.: Straudtmann sen. [Aug. 19, 1941] (Au). La Salle
Co.: Mauermann 12 (Au). Lynn Co.: Rose-Innes & Moon 1052
(Au). Nueces Co.: Tharp sen. [11-9-39] (Au). Parker Co.:
Tracy 7999 (Om, Vt). Sutton Co.: Cory 39625 (Au). Tarrant
Co.: F. C. Gates 19134 (Ka); H. Hapeman s.n. [Fort Worth,
May 'S4J) (Hp); Ruth 92 (Ll), 107 (Cm, Ka), sen. [Fort Worth,

June 19, 1909] (Po). Taylor Co.: Tolstead 6927 (Au); Trac
8000 (Cm). Travis Co.: C. C. Albers 33021 (Au), 34011 (Au,
Au, Au); Birge 2957 (Au); Cohn & ee 13252 (Au); Herb.
Hort. Bot. Gothenb. sen. [Austin, June 6, 1903] (Go); R. He

Painter 6 (Ka); R. Be Payton 41 (Au); Ripperton & Barkley
14542a (Au, N); Smith sen. [Austin, 5/1/1935] (St); Straudt-
mann s.n. [July 17, 1940] (Au); Tharp 44090 (Au), sen. [Aus-
tin, 4712/35] (St), sen. [austin, 5/5/35] (st); Tharp & War-
nock 46084 (Au); Thompson & Hamilton 3451 (Se); Warnock 107
(Au), 46033 (Au); H. H. York sen. [3-18-08] (Au); York &
Wolf sen. [Sept. 29, TO8] (Au). Washington Co.: Brackett 253
(Au), sen. [Apr. 1928] (Au, Au). County undetermined: Capt.
Bolton. sen. [April, 1895] (Ka); Lindheimer 232 (Ka).

VERBENA BONARIENSIS L.

Galveston Co.: Mrs. A. F. Nelson sen. [5-5-42] (Au). Har-
ris Co.: Boon 60 (Au); G. L. Fisher 54094 (Au). Tyler Co.:
C. C. Albers 39011 (Au).

VERBENA BRACTEATA Leg. & Rodr.

Collin Co.: Timmons 743 (Au). Culberson Co.: Tharp 43253
(Au, N). Dallas Co.: F. C. Gates 20972 (Ka). El Paso Co.:
Fringle sen. [El Paso, 11 June 1885] (Vt). Garza Co.: Thar
sene [7/9/41] (Au, N). Hunt Co.: Legget sen. [Loneoak, 7/15/
1927] (Au); Tharp 2929 (Au). Lubbock Co.: Demaree 7562 (H).
ar Co.: Ruth 109 (Cm, Ka). Taylor Co.: Tracy 8001 (Cm,
Vt).

VERBENA CAMERONENSIS L. I. Davis

Cameron Co.: L. I. Davis sen. [Southmost, Spring '41]
(Au),:s.n. [Southmost, March 22, 1942] (Ml); Scology Class
Univ. Texas sen. [3.1.30] (Au); G. L. Fisher 41051 (A)

VERBENA CANADENSIS (L.) Britton

aie

1946 Moldenke, Supplementary Notes 161

Chambers Co.: Tharp 36006 (Au). Galveston Co.: Mrs. Ae Fe
Nelson sen. [3-20-42] (Au). Gregg Co.: C. Le York sen. [3-20
-38] (Au, Au). Harris Co.: G. L. Fisher s.n. [Houston, Apr.
3, 1913] (H, Hp), sen. [Houston, Apr. 22, 1923] (Hp), sen.
(Spring, May 4, 1924] (H), sen. [Mar. 17, 1930] (1); E. Hall
435, in part (Pr). Newton Co.: C. C. Albers 39009 (Au). Folk
Co.: Girvin 2000 (Au), sen. [March 15, 1940] (Au); Rose-
Innes & Warnock 765 (Au); Tharp sen. [3/15/41] (Au, Au).

ZRBENA CANESCENS var. ROEMSRIANA (Scheele) Ferry
Bee Co.: J. S. Williams 39378 (Au). Bexar Co.: Metz 76

[aug. 21] (1), 76 [Auge 24] (1), 881 (I); Texas Agr. &xp.

Sta. Herb. Exchange 3496 (Hp). Brewster Co.: Tharp sen. [6/
19/31] (N), sen. 1075736 ] (Pl); Warnock sen. [May 3, 1937]
(Au). Brooks Co.: Ferkins & Hall 2637, in part (Po). Brown
Co.: Reverchon sen. [Curtiss 1961] (Om, Ka). Comal Co.: OC.
C. Albers 38004 (Au); Lindheimer 294 (Ka), 1074 (Me, Me).
Hidalgo Co.: Mrs. &. J. Walker 30 (Au, N), 34 (Au, Au), 49
(Au, Au), sen. [Feb. 9, 1942] (Au, Au, N); M. Le Walker 66
(Au). Fecos Co.: Tharp 43-800, in part (Au). Starr Co.: ac-
ology Class Univ. Texas sen. [2.28.30] (Au). Travis Co.: C.
C. Albers 32016 (Au), 54009 (Au); Cohn & Barkley 15148 (Au);
R. H. Painter 249 (Au), 392 (Ky); Tharp 44099 (Au), sen.
[7-18-40] (Au, Au), Dt ore oda (aE Warnock 86 (Au); Me
S. Young sen. [10/10/13] (Au), sen. [4/29/14] (Au), sen.
th/viet (Au), sen. [4/5/18] (Au). Williemson Co.: Bodin s.
n. [Georgetown, Dec. 22, 1889] (Ka): Wolcott 117 (Au).

VERBENA CILIATA Benth.

The following notes from my friend, V. L. Cory, are of
particular interest. In speaking of his no. 35568, named by
me as V. ciliata, he says: "This is the white-flowered verb-
ena growing in the Ft. Stockton, Sanderson, and Marathon
country, and which has been determined before for me as V.
pumila (to which I objected) and as V. racemosa. Of the
three I favor V. ciliata. In the FLORA OF TEXAS I note that
you do not give this plant as having a white corolla -- al-
most everything but white; whereas our plant almost always
has a white corolla. In age there is a suggestion of purp-
lish coloration. This species grows in abundance in highly
calcareous soils, is of prostrate growth, and usually makes
rather a limited growth. It may have a connection with V.
racemosa or with V. ciliata, if it is not V. ciliata itself.
This number was of one plant only and was from a-draw about
two miles west of the Glass Mountains."

Brewster Co.: Warnock T.66, in part (Au), W.625 (Au).

Cameron Co.: R. Runyon sen. [Brownsville, 1930] (Hp).

VERBENA CILIATA var. LONGIDENTATA Perry

162 PHYTOLOGIA Vol. 2, no. 4

Bexar Co.: Barkley & Parsons 8 (Au). Cameron Co.: Le Ie
Davis sen. [Palm Grove, Summer '41] (Au). Nueces Coe: Ae Ae
Heller 1385, in part (N, Pl, Se); Tharp sen. [11-9-32] (st).

VERBENA CILIATA var. PUBERA (Greene) Perry
Jeff Davis Co.: Tracy & Zarle 162 (Cm--isotype, Vt--iso-
type ).

VERBENA CLOVERI Moldenke

Mr. Le I. Davis, to whom we are indebted for so much of
our accurate field knowledge of members of this genus, re-
ports that V. Cloveri is found in Kennedy County. He also
states that "our friend Thelma Walker doubts that a lavender
flowered V. Cloveri could be considered a valid variety sin-
ce she says all of the flowers are slightly on the lavender
side of blue*and she thinks every shade of intergrade is
found. She has ea much better chance to study that species...
as her farm is literally covered with the plants in the
spring." .

Brooks Co.: Painter & Barkley 14315 (Au). Frio Co.:
Painter, Lucas, & Barkley 14230 (Au). Kleberg Cow: Je Fe
Sinclair sene [Kingsville, Spring, 1940] (Au).

VERBENA DELTICOLA Small

The Scology Class Univ. Texas sen. [4.1.30] cited by me
in the "Flora of Texas" as this species is actially V. cam-
eronensis. I am deeply grateful to Mr. L. I. Davis for
pointing out to me this and several other mis-identifica-
tions corrected in this paper. He states that "there must be.
a thousand pink-flowering plants of V. delticola to one of
the lavender form. It seems to be a mutation that is likely
to happen anywhere though and once it occurs is likely to
spread locally. Although this color form is sometimes found
mixed up with masses of V. ciliata var. longidentata plants,
it is just as likely to be found where there are no V. cil-
ijata varieties for miles around."

Cameron Co.: Le I. Davis sen. [Olmito, Dec. '41] (Au); Re
Runyon 327 (Au). Hidalgo Co.: Clover 566 (Me).

VERBENA ELEGANS var. ASFZRATA Perry
Hidalgo Co.: Mrs. E. J. Walker 32 (N),-s.n. [Feb. 9,
1942] (Au). ‘ =a

VERBENA HALEI Small

Smith records the common name "European vervain" for this
species from Texas.

Anderson Co.: K. E. Smith s.n. [Palestine, 4/21/35] (St).
Bell Co.: Cohn T.1l (Au); Cohn & Barkley T.44 (Au). Bexar
Co.: Headley sen. [March 24, 1907] (I); Metz 62 (I), 75 (1),

1946 Moldenke, Supplementary Notes’ 163

3240 (Au); E. D. Schulz 766 (I); Texas Agr. Exp. Sta. Herb.
Exchange 3495 (Hp). Brazos Co.: Chenault sen. [April 12,
1937] ; ); Lake sen. (May, 1890] (Pr). Brewster Co.: Warnock
(Au). Burnet Co.: Rose-Innes & Warnock 798 (Au). Cameron
Le Ie Davis Seno {Palm Grove, Summer ‘Diy (Au ); R. Run-
on s.n. [Brownsville, 1930] (Hp). Cherokee Co.: F. Ae
Barkley 13585 (Au). Comal Co.: Lindheimer 537 (Ka), 1076
(Me, Me, Me). Culberson Co.: U. T. Waterfall 4496 (N). Den-
ton Co.: McCart 2006 (Au, St). De Witt Co.: M. Riedel sen.
[9-26-41] (Au). Duval Co.: Croft 119 (Ga). Galveston Co.: R.
L. Crockett sen. (Apr. 22, 1944] (Au); G. L. Fisher s.n.
[Galveston, June 6, 1920] (H); Tracy 7533 (Om). Gonzales Cos:
C. G. Ward 288 (St). Gregg Co.: C. L. York sen. [Fall, 1937]
(Au), sen. [4-2-38] (Au). Grimes Co.: Harding 579 (St); T.
V. Weaver 1038 (Ml). Harris Co.: L. Anderson s.n. [Sept.-_
Oct. 1936] (Au); Boon 62 (Au); G. L. Fisher sen. [Houston,
Apr. 23, 1914] (Hp), sen. [Apr. 9, 1931] (St); EB. Hall 432
(Pr); Rose-Innes & Warnock 673 (Au). Hidalgo Co.: Painter &

Barkley 14428 (Au, N). Jackson Co.: Tharp & Barkley 13Al14

Au, Ml, N, N). Kleberg Co.: J. F. Sinclair sen. [Kingsville
Summer, 1940] (Au). Leon Co.: E. C. Smith sen. [Oakwood,
5-4-1942] (Fe). Montague Co.: McCart 1 (Au). Nueces Co.:
Warnock 20999 (Au). Pecos Co.: Warnock 46166 (Au). Polk Co.:
Girvin sen. [April 12, 1940] (Au, Au); Tharp sen. [4-12-42]
(Au). Reeves Co.: U. T. Waterfall 4388 (N). Tarrant Co.: Fe
C. Gates 19133 (Ka); Ruth 84 (Au), 93 (L1), 108 (Om, Ka), 6.
n. [Fort Worth, June 5, 1909] (Po). Taylor Coe: Tracy 7996
(Om, N, Vt). Travis Co.: C. C. Albers 33016 (Au); Armer 5385
(Au); Cohn & Barkley 13253 (Au); Tharp 1529 (Au), sen. [Aus-
tin, 4723729] (Au, Pl), s.n. [Austin, 5/2735] (Au, St), sen.
{Austin, 5/9/35] (Au, St); Warnock 45-5 (Au), 46104 (Au); Me
S. Young 77 (Au), sen- [4/5/18] (Au). Val Verde Co.: Me Ee
Jones 26229 (1). Waller Co.: Dooley 2 (Au, N). Washington |
Co.: C. C. Albers 53017, im part (Au); Brackett 253 (March
29, 1938] (Au), sen. [April 21, 1939] (Au). Webb Co.: Perk-

ins & Hall 2627, in part. (Po).

VERBENA MACDOUGALII Heller
Culberson Co.: Grassl 175 (I).

VERBENA MATTHESII Turcz.
Fayette Co.: Matthes 13 [Macbride photos 34343] (Kr--
photo of isotype, N--photo of isotype).

VERBENA NEOMEXICANA (A. Gray) Small

The Sixth Grade Brownsville 34, s.n. [Nov. 1934], and
Tharp 1201 cited by me in the "Flora of Texas" as this spe-
cies prove, upon more careful examination, to be V. Runyoni
Moldenke. I am grateful to L. I. Davis for calling my atten-

164 PB TOs LiOeGrigk Vol. 2, no. 4
tion to these mis-~identifications.

VERBENA NEOMEXICANA var. HIRTELLA Ferry

The Ecology Class Univ. Tex. sen. [2.28.30], Re H. Paint-
er 249, and Tharp sen (6/19731] cited by me in the "Flora
of Texas" as this variety prove, upon re-examination, to be
V. canescens var. Roemeriana.
~ Brewster Co.: L. T. Murray sen. [May 22, 1928] (Tt ewe
J. Palmer 34065 (N--isotype); Rose-Innes & Moon 1200 (Au);

Rose-Innes & Warnock 537 (Au); Tharp sen. [Wilson Rench]

(Au); Warnock 20436, in part (Au), sen. [May 3, 1937] (Au).
Hudspeth Co.: U. T. Waterfall 5143 (N). Presidio Co.:

Hinckley 1971 (Au). Travis Co.: C. L. York 46035 (Au, N).

VERBENA NEOMEXICANA vare XYLOPODA Ferry
The Nelson & Nelson specimen cited below is very anomal-
ous. Its fruiting-calyxes and fruit are much larger than
normal. It may prove to be a new variety or even species.
Terrell or Webb Co.: Nelson & Nelson 5138 (Au). Victoria
ri Owens 3125 (Au). Webb Co.: Perkins & Hall 2627, in part
(Po).

VERBENA PERENNIS Wooton

Brewster Co.: Ge L. Fisher sen. [Alpine, Aug. 24, 1932]
(Hp); Warnock 287 (Au), 21090 (Au), 21205 (Au), 21279 (Au),
21827 (Al). Culberson Co.: U. T. Waterfall 3795 (N), 4510
(Au, N), 5209 (N).

VERBENA PLICATA Greene

Baylor Co.: Bridge sen. [near Seymour, 6/16] (Cn). Bexar
Co.: Metz 557 (1), 2156 (Se). Brewster Co.: Cory 43929 (Au);
Rose-Innes & Moon 1169 (Au); Warnock 20436, in part (Au),
20437 (Au), 21230 (Au), sen. [May 3, 1937] (Au). Qulberson
Co.: U. T. Waterfall 4171 (N), 5172 (Au, N). Ector Co.:
Tharp sen. [7710741 | (Au). Frio Co.: Lucas, Painter, & Bark-
ley 14227 (Au). Hidalgo Co.: Mrs. E. Je Walker son. [2/9/42]
Te: Au). Hudspeth Co.: Tharp 43-804 (Au); U. T. Waterfall
5348 (N). Kendall Co.: Metz 170 (I). Kleberg Cow: J. Fe Sin-
clair sen. [Kingsville, Spring 1940] (Au). Midland Co.: Cory
42034 (Au). Mitchell Co.: Cory 48041 (Au). Pecos Cos: Tharp
q 850, in part (Au); Warnock 46122 (Au, N). Taylor Co.:
Tolstead 7103 [Herb. Texas Agr. Exp. Sta. 41986] (Au). Tra-
vis Co.: Armer 5380 (Au). Val Verde Co.: Cory 39090 (N).
Ward Co.: Tracy & Earle 30 (Cm--isotype).

VERBENA PLICATA var. DEGENERI Moldenke

Fecos Co.: Tharp 43-799 (Au), 43-801 (Au), 43-802 (Au),
4 5-80 y) (Au ) e

1946 Moldenke, Supplementary Notes 165

VERBENA PUMILA Rydb.

Bell Co.: Wolff 1373 (Hp). Bexar Co.: Lindheimer 434
(Ka); Metz 152 (I), 524 (1, 1), 3246 (Pl); Fatoni sen. [San
Antonio, Marzi 1914] (Me, Me). Brewster Co.: Sperry = (W).
Cameron Co.: He C. Hanson 322 (Ka). Coke Co.: Cory 37143
(N). Comal Co.: Lindheimer 1075 (Me, Me, Me, Ok). Dallas
Co.: J. Reverchon sen. [Curtiss 1963**] (Go). Edwards Co.:
Cory 37068 (Au), 37070 (N); Me E. Jones 26228 (I). Frio
Co.: Fainter, Lucas, & Barkley 14201 (Au), 14213 (Au). Hays
Co.: Friesner 10401 (Bt). Jeff Davis Co.: Tracy & Earle 178
(Om). Kleberg Co.: Je F. Sinclair sen. [Kingsville, Spring,
1940] (Au). Llano Co.: G. L. Fisher sen. [Biyffton, Apr. 20,
1931] (Bt); Rose-Innes & Warnock 793 (Au). Reeves Co.: Tracy
& Barle 106 (Cm). Schleicher Co.: Cory 34444 (N). Tarrant
Co.: Ruth 110 (Cm). Taylor Co.: Tolstead 7071 (Au, Au). Tom
Green Co.: Cory 40913 (N). Travis Co.: C. C. Albers 34006

(Au); Armer sen. [Austin, 4-2-29] (Fl); BE. Hall wu esr
Tharp 1362 (Au), 1364 (Au), 44116 (Au); Warnock 20623 (Au),
Toes (Au); M. Se Young sen. [2728/14] (Au, N). Williamson
Co.: Wolcott 121 (Au). County undetermined: Turpin sen. (Au,
Au); M. S. Young sen. [west of IAGN Ry.] (Au).

ERBENA QUADRANGULATA Heller

An additional Texas reference is A. Me Te Davis, A study
of Boscaje de la Falma in Cameron County, Texas, and of Sab-
al texana (thesis), p. 62. August, 1942.

Bexar Coe: Metz 755 (Hp, 1). Cameron Co.: A. M. Davis 3s.
n. [Olmito, Summer '41] (Au). Hidalgo Co.: Mrs. 5. J. Walker
TT (Au, N). Nueces Co.: A. A. Heller 1385, in part (Fl),
1388 (Pl--isotype, Se--isotype). Wharton Co.: J. Ks. Small as.
n. [April 12, 1925] (H, We). Zavala Co.: Cory 43814 (Au).

VERBENA RACHMOSA Eggert

Brewster Co.: Cory 31653 (N), 43930 (Au); Rose-Innes &
Warnock 527 (Au), 51436 (Au); Warnock 418 (Au), 20105 (Au).
Glasscock Co.: Cory 42070 (Au). Jeff Davis Co.: Tracy &
®erle 106a (Om). Pecos Co.: He R. Reed 34064 (N); Tharp
43-805 (Au), 43-806 (Au), 43-807 (Au), 43-808 (Au); Warnock
7. Au, Au), 46136 (Au, N). Reeves Co.: Nelson & Nelson
4685 (Au, Ka), 4995 (Au).

VERBENA RIGIDA Spreng.

Galveston Co.: Mrs. A. F. Nelson s.n. [4/20/42] (Au); E.
C. Smith s.n. [Dickinson Bayou, 5-2-1942] (Fc). Harris Co.t
C. C. Albers 35004 (Au); G. L. Fisher sen. [Houston, Sept.
14, 1913] (Hp), sen. [Houston, Apr. 11, 1930] (Bt), sen.
[Sept. 10, 1932] (I). Trevis Co.: Tharp sen. [Austin, 5/2/
a (Bt, St). County undetermined: C. CO. Albers 39008 (Au,
Au).

166 PHY TOLO@ 2h Vol. 2, no. 4

VERBENA RUNYONI Moldenke
An additionel Texan paterdics is A. M. T. Davis, A study

of Boscaje de la Falma in Cameron County, Texas, and of Sab-

al texana (thesis), p. 62. August, 1942. Mr. L. I. Davis,
who-has done much fine work on the verbenas of Texas and
Mexico in the field, reports that "the nutlets of V. Runyoni
are not slightly arched down the back as in Ve neomexicana
and they are plainly longitudinally striate from top to bot-
tom" and the inflorescence is glandular-viscid before dry-
ing. He says "I think we had better assume that neither V.
xutha nor V. neomexicana occurs in Cameron County" and that
all s specimens so named hitherto are really V. Runyoni. "The
first part of this month [August] V. Runyoni — was blooming
everywhere hereabouts. Just below San Benito there was 6
field acres in extent where there was ea plant about every
tro feet. Here according to the soil and moisture supply one
could easily find every stage of V. Runyoni from the slender
one foot high plant to the giant four and a half foot, spice
stemmed: plant. But all Mer ay same seeds and pee
Cameron Co.: Cory 36 (N); Mrs. F. Cottrell 8745 (Au);
tos = Davis sen. re cero Mey, 1942] (Au); Lundell & Lun-
dell 10753 [Plant. Exsicc. Gray. 1274]. (al,-au; Heol ake
Fl, St, We); Rose & Russell 24238 (W); Re Runyon 3178 [Herb.
Texas Agr. Exp. Dp. Sta. 43663] (Au), 4187 (Au, is Sixth Grade
Brownsville 34 (Au), sene [Nov. 193 LJ] (Au); Tharp 1201 (Au).

VERBENA RUNYONI f. ROSIFLORA L. I. Davis

Additional Texan references are A. M. T. Davis, A study
of Boscaje de la Palma in Cameron County, Texas, and of Sab-
al texana (thesis), p. 62. August, 1942, and Moldenke, Known
GSPEr Distrib. Verbenac. Suppl. 1: 2, nom. nud. November 15
1943.

set Co.: Le I. Davis sen. [Southmost, Mey, 1942] (Au
--type).

VERBENA SCABRA Vahl

Bexar Co.: Lindheimer 618 (Ka), 1077 (Me, Me, Me, Ok,
Up); Metz 782 (I). Gonzales Co.: Tharp & Barkley 13850 (Au).
Kerr Co.: G. Le Fisher sn. (Kerrville, Aug. 27, 1932] (Bt).
Kimble Co.: | openings [Aug. 19, 1941] (Au). ate
Co.: B. J. Palmer 8557 y. Real Co.: Cory 39708 (Au), 39709
(N), 42774 (Au). Travis Co.: C. CG. Albers 40004 (Au, Au, Au,
Au, Au, u, Au); F. A. Barkley 13366 (Au); Strandtmann sene
[Aug. 1, 1940] (Au). Val Verde Co.: Cory 38068 (Au, Au),
38069 (N). Walker Co.: C. C. Albers 35010 ay. County unde-
termined: Lindheimer s.n. T1850] (Ka).

VERBENA STRICTA Vent.
Cook Co.: Strandtmann sen. [July 26, 1941] (Au). Tarrant

~
—= "

1946 Moldenke, Supplementary Notes 167

Go.: Ruth 162 (L1), sen. [Fort Worth, Aug. 27, 1909] (Fo).

VERBENA TENUISECTA Briq.

Galveston Co.: Mra. A. F. Nelson sen. 1 Reis (Au).
Jasper Co.: Rose-Innes & Warnock 21818 (Au); Tharp sen. [4/
13/41] (Au, N). Jefferson Co.: C. C. CG. Albers SoCs (Au, Au),
sen. [8/29/34] (Au). Nacogdoches Co.: - Biggar sen. [August 6,
1944] (Au). Polk Co.: Girvin sen. [March 15, 1 1940] (Au, N);
Tharp sen- [5/14/42] (Au).

VERBENA TUMIDULA Ferry
Edwards Co.: Cory 38940 (N). Uvalde Co.: Cory 44509 (Au).

VERBENA URTICIFOLIA L.
Tarrant Co.: Ruth 504 (Cm).

VERBENA WRIGHTII A. Gray
Brewster Co.: Nelson & Nelson 5025 (Ka); Rose-Innes &

Warnock 586 (Au); Warnock 20022 (Au), T. T.66, in part. (Au),
7.203, in part (Au). Culberson Co.: Hitchcock & Stanford

6782 (Pl, Se). Jeff Davis Co.: Hinckley sen. [H. 0. Canyon,
July 27, 1937] (N). Fecos Co.: G. Le Fisher | gene [July 20,
1936] (Se); Warnock C.802 (Au). Presidio Co.: Hinkle Bene
[July 9, LSAT] (Au). Reeves Co.: Nelson & Nelson 4985 (Au,
Ka), 5014 (Au); U. T. Waterfall 4386 2 N). Ward CO ge sere &.
Earle 61 (Ca, we eee

VERBENA XUTHA Lehm.

The Mrs. P. Cottrell 8743 cited by me in the "Flora of
Texas" as Verbena xutha proves, upon re-examination, actu-
ally to be Verbena Runyoni. I am indebted to my friend, L.
I. Davis, for calling my attention to this mis-identifica-
tion.

Aransas Co.: Cory 45879 (Au). De Witt Co.: M. Riedel s.
n. [7-4-41] (Au). Freestone Co.: Harding 399 (St). Galves-
ton Co.: G. L. Fisher sen. [San Leon, July 7, 1929] (Bt);

Mrs. A. F. Nelson sen. [11-2-41] (Au). Grimes Co.: T. Ve

te we S

Weaver 1039 9 (Ml, N). Harris Co.: Boon 61 (Au), 20001 (Au,
foe G. Le tei enes sen. [Houston, May 18, 1914] (Hp); Lind-
heimer s.n. [Hougton, 1843] (Pr). Jackson Co.: Tharp sen.
[8730741] (Au), sen. [Sept. 27, 1941] (Au). Jefferson Co.:
C. C. Albers 34005 (Au, Au); Tharp sen. [9/12/37] (Au, Au).
Leon Co.: EB. C. Smith sen. (Oakwood, 5-12-1942] (Fc). Lib-
erty Co.: Harding gz 172 (St). circ Co.: Le Morris 42
(Au). Travis Co.: F. A. Barkle 5 (Au); R. H. Painter 16
(Ka); Strandtmann sen. (July By + 1940] (Au); Thar 668 (Au),
sen. (8-7-40] (Au). Victoria Co.: Tharp sen. [McFaddin
Beach, 9-11-37] (Pl). Walker Co.: Albers & Warnock 45136
(Au, N). Waller Co.: H. B. Parks 2432 (Au). Washington Co.:

168 PHY 70-406 24 Vol. 2, no. 4

C. C. Albers 33013 (Au), 33017, in part (Au); Brackett 253
(Au), gen. [Apr. 21, 1939] (Au). Wharton Co.: 5. J. Palmer
6622 (W). County undetermined: Lindheimer sen. [Fasc. IV,

1849] (Ka).

VITEX AGNUS-CASTUS L.

Maverick Co.: C. C. Albers 368005 (Au). Real Co.: Cory
34773 (N)- ebita Co.: Ruth 993 (Ka, St). Travis Co.: Ey Ae
Barkley 13081 (Al); Harpin, Waldorf, & Barkley 13081 (Au);
Herb. Univ. Texas sen. rere 8/14/19] (Au). Cultivated:
CG. C. Albers 32019 (Au), 41003 (Au); I. Shiller 870 (Au).

VITEX AGNUS-CASTUS var. ALBA West.
Travis Co.: McKee & Wesley 3896 (Au).

VERBENA .XUTHA Lehm.
Washington Co.: E. Hall 434 (Pr).

September, 1946

CONTENTS

genus Hane I SES Rey ote

- Published by H. A. Gleason and Harold N. Moldenke
The New York Botanical Garden,
Bronx Park, New York 58, N. Y.

NEW yx
BOTANIC
GARDE

NOTES ON THE AVICULARIA. ITI

J. F. Brenckle

_An opportunity to observe the development of exserted
achenes presented itself in the fall of 1941 when weather
conditions were unusually favorable for their production.
Cloudy and damp days with occasional rain were frequent in
September, and during the first half of October there were
eleven days without sunshine. No killing frost occurred un+
til about October 20 so that growth continued up to that
date. Four native species were observed and all agree essen-
tially in the manner of production, but the readiness with
_ which exserted achenes were produced and the extent of
_ growth were quite different in each species. All species are
_ of the fall-fruiting type, that is, flowers and fruits are
_ very sparingly produced in spring and summer but a heavy
crop of seed comes in fall. Of the four species studied,
Polygonum exsertum Small most readily and abundantly produec-
ed exserted achenes. The plants grew near brackish water on
saturated soil, crowded among tall grasses and weeds. The
pond is near Mellette, South Dakota, and is fed by the over-
flow of artesian wells which supply the village. About the
middle of September an abundant crop of fruit and flowers
was present in various stages of development. Lowest on the
inflorescence were many ripe achenes, quite normal and cov-
ered by the perianth, ‘smooth, chestnut-brown, rather narrow,
1.1--1.4 mm. wide and 2.1--2.4 mm. long. Above these were
slightly larger achenes, smooth, chestnut-brown, about 1.5
mm. wide and 2.5--3.0 mm. long, mostly covered by the peri-
anth or sometimes slightly exserted. Above these again were
some immature, olivaceous, exserted achenes, and finally
some blossoms. The riper achenes were quite deciduous and
easily fell away during handling and pressing. It was evi-
dent that exserted achenes do not develop from fully ripe or
almost ripe fruit but grow from very young fruit or from
blossoms while continuously under the influence of a moist
atmosphere. Following a light frost or stormy weather the
_ normal and intermediate forms fell away and the plants had
_ only papery, olivaceous, exserted achenes. The proportion of
_ normal to intermediate and dilated forms depends upon weath-
@r conditions between summer and fall. A prolonged dry sum-
mer delays flowering and if followed at once by a moist fall
the intermediate and exserted forms will predominate. A sum-
mer gradually merging into fall causpgs early flowering, and
normal achenes are produced abundantly.
Polygonum ranosiesinum Mich when growing somewhat re-
169

OP ee ee ee eee

|; =.

ie Say, Ce Peg ty
aiding : oo; on

170 PHYTOLOGIA Vol. 2, no. 5

moved from standing water usually produces an abundant crop
of normal achenes. These plants may then dry up and die.
Other plants standing near water may survive to late fall
when a crop of exserted achenes is produced. Specimens of
these from which the leaves and normal achenes have fallen
are usually classified as P. exsertum.

Polygonum prolificum and its related forms only tardily
exhibit somewhat exserted achenes. Introduced species, if
and when they survive to October, usually show modified a-
chenes which are exserted with a pointed apex. In October P.
aviculare has elongated achenes with smooth, unstriated
faces which become rounded and partly or entirely lose the
triangular shape. The adjective “dimorphic” has been applied
to these variously shaped achenes.

Polygonum commixtum Greene. With the species so far
mentioned above the entire plant is involved, as well as the
entire population in an area. Dr. Greene set up the species
P. commixtum with the exserted and deformed achenes as the
chief character. Careful examination of many specimens dis-
closes that often the whole plant is not involved, but only
a twig, a branch, or some larger part which has become
thickened and condensed and on which the misshapen achenes
are formed. The normal parts of the plant have the usual
normal achenes. It may be noticed that the parts involved
may have been injured, as by trampling. Dr. Greene's specie:
is evidently P. Austinae with exserted and deformed achenes
of this character. In various herbaria specimens with this
deformity have been observed in the following species: P.
Austinae, P. Douglasii, P. Engelmannii, P. sawatchense, and
P. montanum.

The fully developed exserted achene presents an enlarg-
ed embryo loosely surrounded by a paper-thin, olivaceous
pericarp which contains more or less air or gas. The tenden-
cy to produce exserted achenes is common in the Avicularia
but is not present in the related Persicaria. That this uni-
que character. should serve some useful purpose seems almost
certain. The Avicularia do not grow on submerged land nor do
they long survive accidental inundation, although they are
abundant on newly emerged and marginal lands. Contrarily,
the Persicaria do survive inundation, and some species
thrive best in shallow water. On our semi-arid glaciated
prairie are many shallow lake-beds, round or elongated in
shape, interrupted water channels which act as catch-basins.
Some of them dry up regularly during the simmer, but others
may retain water for years. Our native species of Avicularia
are abundant along the shorelines of these basins. They are
among the pioneers which first occupy the bottoms of newly
dried sloughs and lakes. The uncertain waterlevel during
changes from dry to wet cycles, or the reverse, often leaves

1946 Brenckle, Notes on Avicularia 171

these bottoms with bare ground where neither a meadow nor a
slough flora can become established. A sudden filling of

- such a basin will drown standing vegetation and survival may
well depend upon buoyant seed floated to a higher shoreline.
We have here a logical reason for the development and use-
fulness of the exserted achene. Other facts corroborate this
theory. The time of their production, at the beginning of a
wet season, coincides with the most likely time for floods.
Examination of the distribution of colonies is often very
suggestive of waterlevel, windrow deposits. Tufted stems of
old dead colonies may be entirely ignored as likely location
points for the present year's colonies, the new being locat-
ed higher or lower on these shorelines.

The Avicularia have become obligatory land inhabitants
and have extended their range to higher and drier locations.
That the trend has been away from water rather than toward
it is denonstrated by the acquisition of protective ¢harac-
ters in xerophytic situations which are readily discarded
when the plents again occupy wet locations. One such charac-
ter is the accrescent perianth which grows with the achene
and fully envelopes it when mature. During dry weather this
perianth exceeds and clasps the achene, becomes veined and
wrinkled, and is more or less carinate. When wet conditions
prevail with production of exserted achenes the perianth re-
mains shorter and the sections are spreading and smooth.
Reticulations, striations, and other markings on the achenes
are probably due to shrinkage of the surface to reduce evap-
oration. Species of Avicularia on the coastal areas of the
Maritime Provinces normally have large smooth exserted
achenes surrounded by a spreading perianth. The atmosphere
moist from fog, spray, and marshland provides proper con- -
ditions for this type of achene.

SUMMARY :- The exserted achene' serves usefully in the
dispersion of seed during periods of inundation and aids in
its survival. It is produced only by species of Avicularia
under certain’ seasonal and weather conditions. Exserted
achenes have little or no diagnostic value in differentiat-
ing species.

ADDITIONAL NOTES ON THE GENUS PETREA. I
Harold Ne Moldenke
Since the publication of my monograph of thie group in

Fedde's Repertorium Specierum Novarum 43: 1--48 & 161--221
(1938) two hundred and seventy additional specimens have

172 PurtoL 0 @ rs Vol. 2, no. 5

come to my hands and a considerable amount of new informa-
tion. The new specimens, cited below, are deposited in the
herbaria indicated by the following symbols: Ar = United
States National Arboretum, Washington; Ba = Bailey Hortori-
um, Cornell University, Ithaca; Ca = University of Californ-
ia, Berkeley; Cm = Carnegie Museum, Pittsburgh; Dp = DePauw
University, Greencastle, Indiana; Du = Dudley Herbarium,
Stanford University; E = Missouri Botanical Garden, St.
Louis; F = Chicago Natural History Museum, Chicago; Fl = Un-
iversity of Florida, Gainesville; Fs = Forrest Shreve Herb-
arium, Tucson, Arizona; Gt = Botanische Anstalten, Gdétting-
en; Ha = Colegio de la Salle, Vedado, Havana; Hp = H, Hape-
man Herbarium, Minden, Nebraska; I = Langlois Herbarium,
Catholic University of America, Washington; Io = Iowa State
College, Ames; It = Cornell University, Ithaca; Ja = Museu
Nacional, Rio de Janeiro; Kr = B. A. Krukoff Herbarium, New
York Botanical Garden, New York City; La = University of
California at Los Angeles, Los Angeles; Lu = Botanisk Muse-
um, University of Lund, Lund 3 Me = Instituto de Biologia,
Universidad Nacional de México, Mexico City; Mi = University
of Michigan, Ann Arbor; N = Britton Herbarium, New York Bot-
anical Garden, New York City; Oa = Oakes Ames Economie Herb-
arium, Botanical Museum of Harvard University, Cambridge; Po
= Pomona College, Claremont, California; S = Naturhistoriska
Riksmuseet, Stockholm; Sp = Instituto de Botanica, Sao
Paulo; U = Jenman Herbarium, Botanic Gardens, Georgetown,
British Guiana; Ur = University of Illinois, Urbana; Vt = U-
niversity of Vermont, Burlington; W = United States National
Herbarium, Smithsonian Institution, Washington; and Z = He
N. Moldenke Herbarium, Watchung, New Jersey. ;

PETREA Houst.

Synonymy: Fetraea B. Juss. apud Hook. f. & Jacks., Ind.
Kew. 2: 477, in syn. 1895. -- Fetraea Jacq. ex piste ——
4: 70. 1844; Junell, Symb. Bot. Upsal. 4: 43. 1934.

Fet race .L. apud Benth. in Benth. & Hook. f., Gen. Pl. 2

(2): 1149. 1876. -- Peraea L. ex Nees, Flora 4 (1): 300,
ia: 1821; Moldenke, Prelim. Alph. List Invalid Names 33,
in syn. 1940. -- Fetroea L. ex Lam., Encycl. Méth. Bot.

Ill. 3: pl. 539, sphalm. 1797; Neumann, Ann. Fl. Pom. 1837--
1838: 254--255, sphalm. 1838; Moldenke, Prelim. Alph. ‘Liat
Invalid Names 35, in syn. 1940. -- Fetraca Jacq, ex Hoehne,
Resem. Hist. Sece. Bot. Inst. Biol. S.~Paulo 153, sphalm.
1937; Moldenke, Prelim. Alph. List Invalid Names 33 & 34, in
syn. 1940. -- Fehoia L. ex Moldenke in Fedde, Repert. 431 ad

in. syn. 1938; Moldenke, Prelim. Alph. List Invalid Names 33,

in syn. 1940, -- Petrie... ex Moldenke, Prelim. Alph. List
Invalid Names 35, in syn. 1940. -- Pitrea L. ex Moldenke in
Fedde, Repert. 43: 2, in syn. 1938. ~~

Se eee

1946 Moldenke, Notes on Petrea ee BS

References: L., Gen. Pl., ed. 1, 347. 1737; Adans., Fame
fl. 2: 12 & 200. 1763; Necker, Elem. Bot. 1: 362--389. 1790;
Lame, Encycl. Meth. Bot. Ill. 3: pl. 539. 1797; Nees, Flora
4 (1): 300. 18213; Ann. Sci. Nat. Paris, sor. 1, 1: 457.
1824; Neumann, Ann. Fl. Pom. 1837--1838: 254--255. 1838;
Wittstein, Handworterb., ed. 2, 675. 1856; Turez., Bull.
Soc. Nat. Imp. Mosc. 35 (2): 328. 1862; Turcz., Bull. Soc.
Nat. Imp. Mosc. 36 (2): 212. 1863; Benth. in Benth. & Hook.
f., Gen. Pl. 2 (2): 1149. 1876; Ulrich, Internat. Worterb.
Pflanzennamen. 1875; Britten & Boulger, Biogr. Ind. British
& Irish Botanists 135. 1893; Hook. f. & Jacks., Ind. Kew. 2:
477. 1895; Gerth van Wijk, Dict. Plant Names 971. 1911;
Junell, Symb. Bot. Upsal. 4: 43. 1934; Hoehne, Resenha Higt.
Comm. Viges. Anniv. Secc. Bot. 153 & 163. 1937; Standl.,
Field Mus. Publ. Bot. 18: 1011. 1938; Moldenke in Fedde,
Repert. 43: 1--48 & 161--221. 1938; Moldenke, Prelim. Alph.
List Invalid Names 33--35. 1940; Moldenke, Suppl. Alph.

List Invalid Names 6 & 12. 1941; Moldenke, Known Geogr. Dis-'
trib. Verbenac. 17, 20--35, 38, 40--41, 62, 64--65, 73--75,
& 97. 1942; Moldenke, Alph. List Invalid Names 34--36. 1942;
Niemeyer & Stellfeld, Arquiv. Mus. Parana. 3: 8. 1943; Phyt-
ge 2: 108. 1945; B. FP. Reko, Bol. Soc. Bot. Mex. 4: 35.
1946.

Junell in the work cited above gives a detailed discuss-
ion of the gynoecium morphology of the genus, based on P.
Wolubilis. As a result of his findings, he removes the genus
Timotocia from Briquet's Tribe Petraeeae and places it with
Ghinia in his "subtribe" Casselieae. Necker records the com-
mon name "petrée".

In the list of Excluded Species given on page 209 of my
monograph the following corrections are to be made: Fetraea

zanguebarica Gay, Petrea zanguebarica J. Gay, and Fetrea

zanquebarica J. Gay are all synonyms of Dicerocaryum
zanguebarium (Lour.) Merr. of the Pedaliaceae.

PETREA AMAZONICA Moldenke

Le Cointe, in "A Amazonia Brasileira III, Arvores e
Plantas Uteis", p. 470 (1934), under the mis-identification
"Petraea volubilis Jacq." lists the common name “viuvinha"
for this species.

Additional citations: BRAZIL: Amazonas: Krukoff 4930 (N--
photo of isotype, S--photo of isotype, Z--photo of isotype).

PETREA ARBOREA H-BoK.

Synonymy: Petraea arborea H.BeK. apud Pittier, Supl.
Plant. Usual. Venez. 55. 1949. -- Petrea Vincentiana Turcz.
ex Moldenke, Prelim. Alph. List Invalid Names 35, in syn.
1942. -- Petraea aborea H.B.K., in herb. -- Petrea arbirea
H.B.Ke, in herb. '

174 PHYTOLOGIA Vol. 2, no. 5

References: Urb., Symb. Ant. 3: 47--48. 1902; R. O. Will-
iams, Guide Royal Bot. Gard. Trinidad 6 & 11. 1927; Freeman
& Williams, Useful Pl. Trinidad 126. 1928; Stapf, Ind. Lond.
5: 39. 1931; Martyn, Ind. Phan. Jenman Herb. 464, mss. 1937;
Fhelps, Bull. Garden Club Amer., ser. 6, 2: 11. 1937; Pitt-
ier, Supl. Plant. Usual. Venez. 55. 1939.

Freeman and Williams in the reference cited above state
that the species may be propagated by seed or by layering.
They report that the corollas are violet in color, the sep-
als being of a lighter shade than the corolla, and that the
sepals change to a dully ashy-gray after the corollas have
dropped. Pittier in the reference cited above reports the
common name "tostadito" from Venezuela, Delgado records the
same name, and yet lions reports "tosatido". Broadway descri-
bes the species as a “small tree or shrub" in Trinidad. The
name "bejuco de Caballo" recorded by Phelps in the reference
cited above does not properly apply to this species, but to
F. volubilis.

‘The Haught 4046 specimen cited below is anomalous and may
prove not to be this species. It is described as a shrub 2 :
me tall, the leaves very harsh to touch, and the inflores-
cence showy and purple. It was collected on dry hillsides at
an altitude of 600 m. The leaf-texture is more like that of
P. rugosa than that of P. arborea. The label is inscribed
"Dept. Caldas", but E. P. Killip asserts that it was actual-
ly collected in Magdalena.

Additional citations: TRINIDAD: W. E. Broadway S27l (F, N°
--photo, S--phota, Z--photo), 7568 <P, teh, fae (May | 22, :
1911] (Du--120566, F). COLOMBIA: Magdalena: Haught 4046 (N),
VENEZUELA: Aragua: Delgado 135 (F); EB. G. Holt 323 (Cm), 327
(Om); Ll. Williams 10035 (F, F). Carabobo: Saer dHeguert
a, (N). Federal District: Delgado 101 (F, N); Fittier 13573

BE); Ll. Williems 10061 (F). CULTIVATED: Trinidad: Bailey &
Bailey sen. [March 9, 1921] (N--photo, Z--photo).

PETREA ARBOREA var. BROADWAYI Moldenke
Additional citations: CULTIVATED: Trinidad: W. E. Broad-
way sen. [1908] (N--photo of type, Z--photo of type).

ee eee ee ee ee ee ee. ee

PETREA ASPERA Turcz.

References: Pittier, Supl. Plant. Usual. Venez. 55 [as
"Petraea aspera" ]. 1939; Moldenke, Known Geogr. Distrib.
vesienas "2 » 31--33, 38, 73, & 97~ 1942; Phytologia 2: 108.
1945.

The, Seemann 594 cited by me on page 203 of my monograph
as from "Frovince undetermined" in Panama, is actually from
the Canal Zone; the Sonntag 11 cited on the same page as ’
from "Province undetermined" in Colombia is probably from |

fagdalena, but may possibly be from Bolivar (as there is a

“eo

1946 Moldenke, Notes on Petrea “175

"“Volador" in each of these departments). The species is des-
cribed by Williams as a vine growing into the crown of trees
and shrubs, with a deep-blue calyx and purplish-blue corol-
la, inhabiting thickets in "tierra caliente" or "tierra sub-
templada", ascending to an altitude of 730 m. Maggs describ-
es it as a "woody climber with long trailing inflorescences
of a bluebell color" growing at the edges of creeks and a-
long forest trails. Killip calls it a slender tree, 4--6 m.
tall, with drooping inflorescences, the calyx-tube green,
its lobes blue, the corolla-lobes purplish-blue, inhabiting
the edges of woods. Haught reports it to be a high-growing
liana, 20 ms tall, growing along rivers at 50 m. altitude,
with harsh leaves and showy blue inflorescences. The common
name "tostadito" is recorded by Pittier and by Williams
from Venezuela. It has been collected in anthesis in March,
May, and June, and in fruit in March. The Macbride photo-
graph of the type collection is erroneously labeled “Brazil?
- Additional citations: PANAMA: Canal Zone: Maggs II.48 (F,
N). Colon: G. P. Cooper 234 (F). COLOMBIA: Antioquia: Haught

_ 4569 (N). VENEZUELA: Aragua: Ll. Williams 10077 (F), 10160

(F). Bolfvar: Ll. Williams 11475 (F, F, F, N). Carabobo:
Funck & Schlim 507 [Macbride photos 34293] (F--photo of iso-
type, Kr--photo of isotype, lu--isotype, N--photo of iso-
type); Whetzel & Miller s.n. [Herb. Estac. Exper. Agric.

746] (W). Yaracuy: Killip 37068.(W). BRAZIL: State undeterm-

ined: Glaziou sen. (N--photo, Z--photo).

PETREA ATROCOERULEA Moldenke x
Additional citations: COLOMBIA: Antioquia: Kalbreyer 1634
(N--photo of type, Z--photo of type).

PETREA BLANCHETIANA Schau.

The Sellow specimen cited below was erroneously determin-
ed as "Petraea volubilis L." by the collector.

Illustrations: Mart., Fl. Bras. 9: pl. 45. 1851; Correa,
Diccion. Pl. Uteis Brasil 1: 502. 1926.

Additional citations: BRAZIL: State undetermined: Sellow
180 (Vt).

PETREA BRACTEATA Steud. 3
Synonymy: “"Petraea macrostachya Benth." sensu Pulle,
Enum. Pl. Surinam. 402. 1906 [not Petrea macrostachya Benth.,
1839]. -- "Petraea volubilis Jacq." sensu Fulle, Mum. Pl.
Surinam. 403. 1906 [not Petrea volubilis Le, 1753]. --
"Petraea racemosa Nees & Mart." sensu Pulle, mnum. Pl. Suri-
nam. 403. 1906 [not. Petrea racemosa Nees, 1821]. -- "Petraea
arborea H.B.K." sensu Pulle, Enum. Pl. Surinam. 403. 1906

| [not Petrea arborea H.B.Ke, 1818]. -- “Petraea rugosa H.B.K:'
etrea

—

sensu Fulle, Enum. Pl. Surinam. 403. 1906 [not rugosa

176 PHYTOLOGIA Vol. 2, no. 5

H-BeKe, 1818]. ;

References: Benth., Ann. Nat. Hist. 2: 448. 1839; Pulle,
Enum. Pl. Surinam. 402--403. 1906; Martyn, Ind. Phan. Jenman
Herb. 464--465, mss. 1937; Moldenke, Prelim. Alph. List In-
valid Names 34--35. 1940; Moldenke, Alph. List Invalid Names
34--35. 1942; Moldenke, Know Geogr. Distrib. Verbenac. 33,
38, & 97. 1942; Fhytologia 2: 108. 1945.

Smith describés the species as having the "inflorescence-
stalk, calyx, and corolla rich blue", growing in dense for-
est at edge of isolated savannas. Ducke says it has violet
flowers and grows on non-inundated land. Ducke 1144 was err-
oneously determined by him as P. Martiana, while S72 was er-
roneously determined by me and cited in Castanea 10: 42
(1945) as P. rugosa.

Illustrations: Miquel, Stirp. Surinam. [Nat. Verh. Holl.
Maatsch. Wet. Haarlem, ser. 2, 7:] pl. 42 [as "P. macro-
stachya"]. 1850; Glifck, Blatt- u. Bliftenmorphol. Stud. 382
[as "P. Schomburgkiana"]. 1919.

Additional citations: BRITISH GUIANA: E. H. Graham 254
(Cm); A. S. Hitchcock 17249 (N--photo, Z--photo); Jenman
3882 (U), 5955 (U); Maguire & Fanshawe 22885 (N); M. Re
Schomburgk 108 [Macbride photos 17574] (Kr--photo, N--photo,
ephote F. 175 (F); As Ce Smith 2626 (F, N). SURINAM: Host-
mann 39 (Macbride photos 34294] (F--photo of isotype, Kr--
ph~to of isotype, N--photo of isotype); Hostmann & Kappler
39 [Macbride photos 22776] (Kr--photo of isotype); Kegel
1179 (Gt, Gt), 1180 (Gt); Maguire 24831 (N); Nolte sen.
(Gt); Wullschligel 411 (Gt), 1587 Tay BRAZIL: Amazonas?
Ducke 872 (N), 1133 (N). LOCALITY OF COLLECTION UNDESIGNAT-
ED: Herb. De Candolle s.n. [Macbride photos 7875] (Kr--
photo).

PETREA BREVICALYX Ducxe

Synonymy: Petraca Kuhlmannii Moldenke, Prelim. Alph. List
Invalid Names 34, in syn. 1940.

References: Ducke, Archivos Jard. Bot. Rio Janeiro 6: 87
{as "Petraea brevicalyx" ]. 1933; Moldenke, Prelim. Alph.
List Invalid Names 34. 1940; Moldenke, Alph. List Invalid
Names 34. 1942; Moldenke, Known Geogr. Distrib. Verbenac. 38
& 97. 1942.

Ducke describes the species as a low, weak, scandent
shrub, with a dark-violet calyx and violet corolla. It has
been collected in anthesis also in January.

Additional citations: BRAZIL: Amazonas: Ducke 140 (F, N;
S), sen. (Herb. Rio de Janeiro 22544] (N--cotype).

PETREA COLOMBIANA Moldenks
One of the specimens of Lopez R. cited below includes a
photograph of the species growing in situ.

.
|

‘
wz .
a 7. eS. ee ee ee

'
eae ee

1946 . Moldenke, Notes on Petrea 177

Additional citations: COLOMBIA: Santander Sur: Lopez R.
sen. (Bucaramango, 12/19/1918] (Ar, Ar). CULTIVATED: Colomb-
fa: Killip & Smith 19067 (N--photo of type, S--photo of
type, Z--photo of ty pe )e

PETREA DUCKEI Moldenke

References: Moldenke, FPhytologia 1: 469--470. 1940; Mol-
denke, Known Geogr. Distrib. Verbenac. 38 & 97. 1942.

Scandent shrub; branches slender, grayish, minutely pub-
erulent, glabrescent in age, somewhat lenticellate; nodes
not annulate; principal internodes 1.2--11.5'cm. long;
leaves decussate-opposite; petioles stout, 8--12 mm. long,
minutely puberulent, flattened above; blades firmly charta-
ceous, stiff, rather uniformly gray-green on both surfaces,
somewhat shiny above, elliptic, 10.5--16 cm. long, 4.3--8
em. wide, abruptly acute or short-acuminate at apex (the
very point often obtuse), entire, acute or rounded at base,
very minutely puberulent and scabrellous on both surfaces,
glabrescent and merely punctate in age, the immature blades
very thin-membranous and nigrescent in drying; midrib rather
stout at base, rapidly diminishing in size as the apex is
approached, prominent on both surfaces; secondaries very
slender, 9--15 per side, prominulous above, sharply promin- .
ent beneath; vein and veinlet reticulation abundant, promin-
ulous on both surfaces, the tertiaries sharply prominent be-
neath; inflorescence axillary, racemiform, 14--25 om. long,
rather loosely many-flowered; rachis slender, minutely pub-
srulent; pedicels slender, 2--5 mm. long, elongate to 9 m.
in fruit, minutely puberulent; calyx lilac, subtended by 1--
5 foliaceous prophylla, which are thin-membranous, elliptic,
venose, 5--6 mm. long, 3--4 mm. wide, sharply acute or at+
tenuate-acuminate at apex; corolla violet; fruiting-calyx
indurated, its tube 6--7 mm. long, 5--7 m. wide at apex,
very minutely puberulent, its lobes greatly enlarged, broad-
ly elliptic, to about 15 mm. long and 12 mm. wide, pinnately
venose, very minutely and obscurely puberulent, abruptly a-
cute or obtuse at apex.

The type of this species was collected by Adolfo Ducke
(Herb. Jard. Bot. Rio de Janeiro 22542] -- in whose honor it
is named -- on inundated shores at Parana de Anavilhana, on
the lower Rio Negro, Amazonas, Brazil, on July 24, 1939. The
species is obviously very closely related to P. insignis.

Citations: BRAZIL: Amazonas: Ducke 688 (N), | Sone com
Rio de Janeiro 22542] (N--type).

-PETREA GLANDULOSA Pittier

References: J. A. Clark, Card Index, issue 116. January
10, 1928; Pittier, Supl. Plant. Usual. Venez. 55 [as "Pet-
Yraea glandulosa"). 1939; Moldenke, Known .Geogr. Distrib.

178 PHYTO LOG LA Vol. 2, noe 5

Verbenac. 32 & 97. 1942; Moldenke, Fhytologia 2: 108. 1945.
Pittier, in the reference cited above, records the common
name "“penitente" from Venezuela.

PETREA INSIGNIS Schau.

References: Le Cointe, A Amazonia Brasileira III, Arvores
e Plantas Uteis, 169 fas "Petraea insignis" ]. 1934; Moldenke,
Prelim. Alph. List Invalid Names 35. 1800; Moldenke, Known

Geogr. Distrib. Verbenac. 38, 73, & fe 1942; Moldenke, Alph.

List Invalid Names 35. 1942; Le Cointe, O Estado do Para 248
& 251 [as "Petraea insignis"]. 1945.

Le Cointe in the referencescited above records the common
names "flor de S. Miguel", "flor de S. Miguel", "viuvinha",
and "flor de folha grande" ;,and states that the species is
cultivated in parks and gardens in Parad. Spruce records the
common name "flor de Espirito Santo" on his no. 1354.

Additional citations: BRAZIL: Amazonas: Schwacke 304
(Herb. Rio de Janeiro 32217a] (Ja); Spruce sen. [in vicini-
bus Barra, Dece--Mart. 1850--51] (F, Lu). Para: Martius sen.
[Pard; Macbride photos 20348] (ne-sphatoe of type).

PETREA KOHAUTIANA Presl

References: Britton, Fl. Bermuda 320. 1918; Re O. Will- .
iams, Guide Royal Bot. Gard. Trinidad 15 [as "P. volubilis"}
1927; Freeman & Williams, Useful Pl. Trinidad 126--127 [es
"P. volubilis"]. 1928; Moldenke, Prelim. Alph. List Invalid
Names 354 & 35. 1940; Moldenke, Alph. List Invalid Names 35.
1942; Moldenke, Known Geogr. Distrib. Verbenac. 28, 29, 73,
& 97. 1942; Phytologia 2: 108. 1945.

Britton, in the reference. cited above, records the common
name "tree petrea", while Freeman and Williams record “white
petrea" and "bridal wreath", but it seems probable that
these last two names apply more strictly to var. alba. Duss
records the common name "liane rude". Hodge states. that the
species grows along roadsides on Dominica. It has been col-
lected in fruit in August. The Harvey Herbarium specimen
cited below bears a very interesting label reading, in a
very old hand: "Anonimos scandens ramulis asperrimis a lim-
arum instar lignum rodentibus plum.-lin. G.748. didinamia
angiospermia monop. Chelone lin. Digitalis virginiana etc.
Pluk. Species. ou liron licti, liane a feuilles rude bone
pour Amer. l'argent an ou Citiragouli, Malpighiae Species;
planta a fleurs en rosette." Delrisse records the common
names "liane rude" and “fleur St. Jean."

Additional citations: HISPANIOLA: Haiti: pede et
(Du--166367). GUADELOUPE: Delrisse s.ne [1844] (p art
Duchassaing sen. [1852] (Du). DOMINICA: Hodge 870. (), 3651
(N). MARTINIQUE: Duss 1979 (F); Kohaut sen. [Sieber, ae
Mart. 157, in part] (N--pheto of isotype, Z-~photo of iso-

re

_——~
ies A

1946 Moldenke, Notes on Fetrea 179

type), Sen. Faféber: Fl. Mixta 374] (Lu). GRENADA: W. Ee
Broadway 8-! sen. (Presbytery, June 6, 1906] (F). CULTIVATED:
St. Croix: L. A. Ricksecker 329 9 (F).

PETREA KOHAUTIANA var. ALBA (Freeman & Williams) Moldenke

Synonymy: Petrea volubilis var. alba Freeman & Williams,
Useful Pl. Trinidad 127. 1928. -- Petrea Kohautiana var.
gnomala Moldenke in Fedde, Repert. 43: 31. 1938. -- Fetraea
alba Hort. ex Moldenke in Fedde, Borat, 43: 31, in syn.
1938.

References: Re O. Williams, Guide Royal Bot. Gard. Trini-
dad 15. 1927; Freeman & Williams, Useful Pl. Trinidad 126--
127. 1928; Moldenke, Geogr. Distrib. Verbenac. 49, 1939;
Moldenke, Alph. List Common & Vern. Names 6. 1939; Lilloa 4:
309. 1939; Moldenke, Prelim. Alph. List Invalid Names 34 &
35. 1940; Moldenke, Alph. List Invalid Names 34 & 35. 1942;
Moldenke, Known Geogr. Distrib. Verbenac . 73 & 97~ 1942;
Phytologia 2: 108. 1945.

Freeman and Williams record the common names “white pet-
rea" and "bridal wreath". Broadway, as long ago as 1908, de-
scribed the plant on his collection labels as "Petraea
volubilis (white variety)", and stated that it has "branches
pendent, sometimes climbing."

Additional citations: CULTIVATED: Trinidad: W. E. Broad-

way 3197 (S--photo), sen. [St. Ann's, 1908] (La).

PETREA LONGIFOLIA Moldenke

Synonymy: Petraea longifolia Moldenke, Suppl. List Inval-
id Names 6, in syn. 1941.

References: Moldenke, Alph. List Invalid ‘ames 34. 1942;
Moldenke, Known Geogr. Distrib. Verbenac. 75 & 97. 1942.

~Tais species is very close to P. maynensis Huber and may
prove to be conspecifie with it.

Additional citations: LOCALITY OF COLLECTION UNDESIGNAT-
ED: Herb. Mus. Paris son. (F--fragment of isotype).

PETREA MACROSTACHYA Benth.
Synonymy: FPetraea guianensis Cham. ex Moldenke, Prelim.

_Alph. List Invalid Names 34, in syn. 1940.

References: Martyn, Ind. Phan. Jenman Herb. 465, mss.
1937; Moldenke, Prelim. Alph. List Invalid Names 3h. 1940;
Moldenke, Known Geogr. Distrib. Verbenac. 33, 38 & 97.
1942; Moldenke, Alph. List Invalid Names 34 & 35. 1942;
Phytologia 2: 108, 1945.

Ducke describes the flowers as violet, while Smith re-
ports the calyx "rich blue; corolla rich violet" or "calyx
pale blue, marked with green" and states that the species
grows in dense forests at altitudes of from 150 to 400 m. It
has been collected in anthesis in January, March, and April,

180 PB Yi? . Ob O9G- Ik Vol. 2, now 5

and in fruit in March, April, and September. The Macbride
photograph no. 34292 » cited below, is erroneously labeled
"Funck & Schlim 150 ". The Hostmann sen- distributed as this
species, from Surinam; ig actually Tripleris surinamensis
Cham. in the Polygonaceae, as is also the Schweinitz sene
from the same country. The illustration in Miquel, Stirp.
Surinam. [Nat. Verh. Holl. Maatsch. Wet. Haarlem, ser. 2, 7:]
pl. 42 (1850) is not P. macrostachya, but is P. bracteata
Steud.

Additional citations: BRITISH GUIANA: M. Re Schomburgk
158 [Macbride photos 34292] Sica of isotype, Kr--photo
of of isotype); Ae Ce. Smith 2148 (N), 3401 (F, N)» BRAZIL:
Pard: Ducke sene [Herb. Rio de Janeiro 14294] (N).

PETREA MARTIANA Schau.

“References: Le Cointe, A Amazonia Brasileira III, Arvores
e Plantas Uteis, 470 [as "Petraea martiana" ]. 1934; Molden-
ke, Prelim. Alph. List Invalid Nemes 34° & 35. 1940; Molden-
ke, Known Geogr. Distrib. Verbenac. 35, 38, & 97. 1942; Mol-
denke, Alph. List Invalid Names 34 & 35. 1942; Le Cointe, 0
Estado do Para 251 [as "Petraea martiana"]. 1945.

The species has been collected in anthesis in June and
October, and in fruit in February and October. Ducke descri-
bes the flowers as violet and states that the species grows
along the margins of woods.« Le Cointe says it is cultivated
th parks and gardens in Para, and records the common name

"vyiuvinha"

Additional citations: PERU: Loreto: W. Fox 96 (F). BRAZIL:

Pard: Ducke s.n. (Herb. Rio de Janeiro 14291] (N); Sampaio
5092 (Herb. Rio de Janeiro 19128, in part] ES

PETREA MAYNENSIS Huber

References: Moldenke, Prelim. Alph. List Invalid Names
35. 1940; Moldenke, Alph. List Invalid Names 35. 1942; Mol-
denke, Known Geogr. Distrib. Verbenac. 31, 35, 38, 40, 73, &
97. 1942.

Ducke reports the calyx as almost white and the corolla
violet, while Kuhlmann describes the calyx as blue and the
corolla violet, but white at the center. It has been col-
lected in anthesis also in January and February. It is poss-
ible that P. longifolia may be conspecific with this species

Additional citations: COLOMBIA: Putumayo: Klug 1894 (s).
PERU: Loreto: Huber 1489 (N--photo of fragment, N--photo of
type, Z--photo of fragment, Z--photo of type). BRAZIL: Ama-
zonas: Ducke sen. [Herb. Rio de Janeiro 35657] (N). Para:
Huber 3676 (N--photo, Z--photo). BOLIVIA: El Beni: J. G.
Kuhlmann sen. (Herb. Rio de Janeiro 22541] (N). La Faz: Kru-
koff 10729 (N), 10735 (N). Santa Cruz: Steinbach 3470 (Herb.
Inst. Miguel Lillo 38010] (N, N). CULTIVATED: Brazil: Ducke

ae se se ee

(1946 Moldenke, Notes on Petrea 181
sen. [Para; Herb. Rio de Janeiro 22543] (N).

PETREA NITIDULA Moldernke

Additional citations: BRAZIL: Amazonas: Spruce 2536 (Lu),
2926 [Macbride photos 34290] (F--isotype, F--photo of iso-
type, Kr--photo of isotype, N--photo of isotype).

PETREA PERUVIANA Moldenke
Williams reports the vernacular name "sanango sacha" for
this species and records its blooming and fruiting in May
and June.
Additional citations: PERU: Loreto: Klug 170 (F), 637 (F
--isotype); Ll. Williams 690 (F), 8106 (F).

PETREA PERUVIANA var. ACUMINATA Moldenke

Williams describes this plant as a "forest creeper",
while Miss Mexia says it is a "vine climbing medium trees, .
altitude 110 m., frequent in cut-over woods, flowers purple,
February."

Additional citations: PERU: Loreto: Mexia 6498 (F, I);
Ll. Williams 647 (F).

PETREA PUBESCENS Turcz.

References: Pittier, Supl. Plant. Usual. Venez. 55 [as
"Petraea pubescens" ]. 1939; Moldenke, Alph. List Common &
Vernac. Names 25. 1939; Moldenke, Known Geogr. Distrib.
Verbenac. 31, 32, 34, 38, 73, & 97. 1942; Phytologia 2:
108. 1945.

Perez Arbeldez and Cuatrecasas describe the flowers of
this. species as lilac. It has been collected in fruit in
January, April, and August. The leaves on the Garcfa Barr-
iga specimen cited below are anomalous in being very thin-
textured. Ule 9722 is labeled "Seringal Auristella, Feru",
but seems actually to have been collected in Acre Terri-
tory, Brazil. The Macbride photograph 34291, cited below,
is erroneously labeled "Brazil".

- Additional citations: COLOMBIA: Cundinamarca: Pérez Ar-
beldez & Quatrecasas 6577 (N, W). Magdalena: H. H. Smith
1521 (Ca, Om, Vt). Meta: Garcfa Barriga gene THerb. Nace
Colomb. 5206] (W). Santander Norte: Quatrecaseas & Garcfa
Barriga 10173 (W). VENEZUELA: Mérida: Funck & Schlim 1504
[Macbride photos 34291] (F--photo of isotype; Kr--photo of
isotype, Lu--isotype, N--photo of isotype). BRAZIL: Acre
Territory: Ule 9722 (N).

PETREA PUBESCENS var. KLUGII Moldenke
Additional citations:, PERU: San Mart{n: Klug 4155 (E--
isotype, I--isotype, S--isotype).

182 Po XY PT Och sO. Gs S Vole 2, noe 5

PETREA RACEMOSA Nees

References: Mart., Fl. Brase 9: ple 45 [as "P. subserra+
ta"]. 1851; Boeq., Rev. Verbenac. pl. 20. 1861--1863; Baill-
on, Hist. Pl. 11: 80, figs. 78--81. 1891; Briq. in Engl. &
Prantl, Nat. Pflanzenfam. 4 (3a): 157. 1895; Crevost & Pate-
lot, Bull. Econ. Indo-chine 37: 1288 [as "P. subserrata"].
1934; Moldenke, Annot. List 108. 1939; Hoehne, Kuhlmann, &
Handro, O Jard. Bot. S&o Paulo 578 [as "Petraea subserrata" ]
1941; Moldenke, Prelim. Alph. List Invalid Names 34 & 35.
1940; Moldenke, Alph. List Common & Vernac. Names 12, 25,
31, & 53. 1939; Moldenke, Known Geogr. Distrib. Verbenac.
38, 41, 74, & 97. 1942. 1942; Moldenke, Alph. List Invalid
Names 34 & 35. 1942; Kuhlmann, Instit. de Botan. Observ.
Ger. Contrib. 5: 20 & VI [as "Fetraea racemosa" ]. 1942; Sam-
paio & Peckolt, Arquiv. Mus. Nac. Rio de Janeiro 37: 375 [as
"P, volubilis Vell." and "P. subserrata"]. 1943; Stellfeld,
Tribuna Farmaceutica [Vellozoa] 12: 55, 62, & 102 [as "PB.
volubilis Vell.", "P. subserrata", and "P. sub-serrata
Chame"]J. 1944; Phytologia 2: 108. 1945.

Hoehne, Kuhlmann, and Handro, in the reference cited a-
bove, record the common names "flor de S. Miguel" and
"viuvinha", Kuhlmann records “flor de Sao Miguel" and "flor
de viuva", and Stellfeld lists "coroa de viuva", "grinalda
de viuva", "touca de viuva", and "flor de S&o Miguel".
"Purple wreath" is recorded on the Ames Herbarium specimen.

Additional illustrations: Mart., Fl. Bras. 9: pl. 45 [as
"P. subserrata"]. 1851; Bocq., Rev. Verbenac. pl. 20. 1861--
1863; Briq. in Engl. & Prantl, Nat. Pflanzenfam. 4 (3a):
157. 1895; Crevost & Pételot, Bull. Econ. Indo-chine 37:
1288 [as "P. subserrata"]. 1934.

Additional citations: BRAZIL: Bahia: Wied-Neuwied s.n.
[Belmonte] (E--photo of type). Minas Geraes: G. Gardner 5127
(F); Heringer 63 [Herb. Inst. Biol. S. Paulo 39039] (Sp),
-gen. [Herb. Est. Exp. de Café; Herb. Dept. Bot. Est. Se
Par lo 44607] (N); Mello Barreto 3267 (Herb. Jard. Bot. Bello
Horizonte 931] (F), 268 [Herb. Jard. Bot. Bello Horizonte
11244] (F); Mosen 645 (Lu); Widgren s.n. (1845; Herb. Monac.
1645; Herb. Rio de Yaneiro 31718] (Lu, Lu, N). Parandé: Dusén
15874 (Lu), sen. [Jaguariahyva, 27.11.1914] (La, Mi). Rio de
Janeiro: Hagendorf sen. [cotype coll. of Petraea subserrata
Cham.] (S--photo); Herb. Rio de Janeiro 31719 (N); Martius
sen. [1823; Macbride photos 7874] (Kr--photo). Santa Cathar-
ina: Herb. Rio de Janeiro 31775 (N). S&o Paulo: Amaral 3
{[Herb. Inst. Biol. S. Palo 34704] (Sp); Santoro sen. (Herb.
Inst. Agron. Est. S. Palo 678] (Ba); Swentorzecxy 6 [Herb.
Dept. Bot. Est. S. Paulo 41835] (N); Zagatto sen. [Herb.
Inst. Agron. Est. S. Paulo 3096; Herb. Dept. Bot. Est. 3S.
Paulo 44503] (Sp). State undetermined: P. Clausen sen.
[1840] (Du--166368); Sellow sen. [Macbride photos 175/53 co-

dots ie. ae a eae) ee

i.

Pee er Pay were

;

1946 Moldenke, Notes on Petrea 183

type coll. of Petraea subserrata Cham.] (Kr--photo, S--photo,
Vt); Wied-Neuwied s.n. [Brasilia] (Lu). CULTIVATED: Brazil:
Piere sene [Rio de Yaneiro; Herb. Rio de Yaneiro 31520] (N).
Missouri: Herb. Ames sen. [Ste Louis] (0a). New York: Pe Je
Connolly sen. [New York Bot. Gard. Cult. Plants 27675] (N);
Hartling sen. [New York Bot. Gard. Cult. Plants 8885] (Ur),
gen. [New York Bot. Gard. Cult. Plants 11813] (Ur); H. Ne
Moldenke 4644 (N), 10409 (N).

PETREA RUGOSA H.BKe

.The species is reported by Haught as growing fully expos-
ed to the sun on dry sand ridges, with very. showy inflores-
cences of bright-blue flowers. Popenoe describes it as "a
rare and handsome ornamental slender shrub", blooming in
November. Ducke describes it as a woody vine, with violet
flowers, growing in non-inundated woods. Dryander reports
the flowers as "lilac" in color and reports the common name>
"“chaparilla". Daniel gives the common name "pluma de reina"
and states that the inflorescences are blue. The species has
been misidentified as P. arborea by some workers.

Additional citations: COLOMBIA: Antioquia: Daniel 2665
(W--1857567). Caldas: Haught 2103 (N). £1 Valle: Dryander
2303 (W). Tolima: Hartweg 1359 (Lu). VENEZUELA: Federal Dis-
trict: Bonpland sen. [Macbride photos 39477] (F--photo of
type, Kr--photo of type). BRAZIL: Amazonas: Ducke 872 (W).
CULTIVATED: Honduras: Yuncker 4705 (Dp). Colombia: Popenoe

1203 (Ar).

PETREA SCABERRIMA Moldenke
Additional citations: COLOMBIA: Cundinamarca: Purdie sen.
{Santa Fé de Bogota] (N--photo of type, Z--photo of type).

PETREA VOLUBILIS L.

Synonymy: Petraea arborea (Kunth) Smith & Wiles in
Forbes, Wand. Nat. East. Arch. 2: 78--79 & 514. 1885; H. J.
Lam, Verbenac. Mal. Arche, addenda. 1919. -- Fetrea mexicana
Schiede ex Moldenke, Prelim. Alph. List Invalid Names 35, in
syn. 1940. -- Petraea volubulis Merr., Plant Life Pacif. 161
_& 274, sphalm. 1945. -- Petrea uolubilis Sesse & Moc., in
herb.

References: Jacq., Select. Stirp. Amer. Hist. pl. 114.
1763; Jacq., Select. Stirp. Amer. Hist. Picta, pl. 173. 17&;
Lodd., Bot. Cab. 8: pl. 736. 1823; Vell., Fl. Flum. 6: pl.
59. 1827; Géel, Sert. Bot. 3: cl. 14. 1832; Knowles & Westc.,
Floral Cab. 3: pl. 108. 1840; Journ. Hort., ser. 3, 7! 53.~
1883; Forbes, Wand. Nat. East. Arch. 2: 78--79 & 514. 1885;
_Baillon, Hist. Pl. 11: 80, figs. 78--81. 1891; Bois, Dict.
Hort. 944. 1893--1899; G. W. Oliver & W. Miller in L. H.
Bailey, Cycl. Amer. Hort. 4: 1284. 1901; Millsp. & Loes. in

2) ae
ae
¢.

184 PHY’? 0.L-0-@:5, 4 Vole 2, noe 5 q

Engl., Bot. Jahrb. 36: Beibl. 80: 26. 1905; Gard. Chron.,
sere 3, 39: 24 & 25, fig. 15. 1906; Journ. Hort., ser. 3, Sk
390. 1907; Gard. Chron., ser. 3, 51: 287. 1912; Le. H. Bail-
ey, Stand. Cycl. Hort. 5: 2562. 1916; Rehnelt, Gartenwelt
28: 367s figs. 1924; Jordahn, Gard. Chron. Amer. 30: 171.
1926; Nessel, Gartenfl. 75: 321--322, fig. 1926; Junell,
Symbe. Bot. Upsal. 4: 43 & 45 [as "Fetraea volubilis
Jacqe"]. 1934; Catologo Quinta Perez Estr. San Pedro Sula
30. 1935; Phelps, Bull. Garden Club Amer., ser. 6, 2: ll.
1937; Moldenke in Fedde, Repert. 43: 1--48 & 161--221. 1938;
Svensk. Bot. Tidsk. 32: 231. 1938; Standl., Field Mus. Publ.
Bot. 18: 1012. 1938; Moldenke, Alph. List Common & Vernac.
Names 355 45:65 19 85. 99125195 VIG, 25 255 26, eee
30, & 41. 1939; Moldenke, Suppl. List Common & Vernac. Names
24. 1940;:Moldenke, Prelim. Alph. List Invalid Nemes 34 &
35. 1940; Calderén & Standl., Fl. Salvador., ed. 2, 238 [as
"p, arborea" ]. 1941; Ingtit. de Botan. Observ. Ger. Contrib.
2: 65 & 5: VI [as "Petraea volubilis L."]. 1942; Moldenke, 4
Alph. List Invalid Names 54--36. 1942; Moldenke, Known Geogr.
Distrib. Verbenac. 17, 20--27, 29, 62, 64, 65, 74, & 97. :
1942; He Fe Macmillan, Trop. Planting & Gard., ed. 5, 122.
1945; E. D. Merr., Plant Life Pacif. 161 & 274. 1945; Mol-
denke, Phytologia 2: 108. 1945; Le Cointe, O Estado do Para
251 {as "Petraea volubilis Jacq."]. 1945; New York World
telegram for April 5, 196, pe 17. 1946.

The common name "queen's-wreath" is recorded for this
species by A. Ce Jordahn in the reference cited above, who
describes the plant as "one of Florida's lovely vines." The
name "bejuca de caballo” recorded by Fhelps in the reference
cited above for P. arborea actually applies to P. volubilis
instead. The names "choreque" and "purple wreath" are record-
ed by Standley (above), who reports the species as frequent
in thickets and dry forests of the Facific "tierra calien-
te" of Costa Rica. Gerth van Wijk lists "purple wreath",
"liane rude", and "liane de St.-Jean". The Mayan name
"“yoxop-d3imin" is recorded in Phytologia 2: 108. Grey and
Hubbard in List Plants Bot. Gard. Atkins Inst. 157 (1933)
record the name “queen's wreath" from Cuba. The vernacular
designation "chaparro"” is listed in Catologo de la Quinta
Perez Estrada San Fedro Sula’ (above) from Honduras. scneers
ing to the lundells the species is called "piocha viejo" in

Yucatan, where it is a common woody vine to 2 m. tall in the ©

advanced deciduous forest and among second growth. Steggerda

‘records the Mayan name "yoch opp tzimin", while Millspaugh

and Loesener in the reference cited above record “opp-tziminl'

and “purple-wreath". From Cuba comes the name "flor de
papel", recorded by Leon. Le Cointe says (above) that in
Para it is cultiveted in parks and gardens and called
"viuvinha".

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BOTANIC:
‘ RDED

~ October, 1946 ie No. 6

CONTENTS

J. VAN MELLE, The Junipers commonly included
| in SUMED OVUR: CIATUOMSES: 50 s50. kon <, cots Sescat sa Hhesewes ea veahen settee
NRT OH OAL oy a. Nhs AY WUED ih REAR Etc yeyGEOR ATO A bo

_N. MOoLDENKE, Additional notes on the genus

MANOA ET Cr A Oe USN ITO) OMe aie pea

Published by H. A. Gleason and Harold N. ‘ra tain!
The New York Botanical Garden,
Bronx Park, New York 58, N. Y.

Price of this number, 50 cents; per volume, $5.00 in advance,

Vol. 2, No. 5, was issued September 20th, 1946

P. J. van Melle

_ The materials enumerated here, their relationships and
distributions, are treated more amply ina manuscript, "Re-
view of Juniperus chinensis", now in the press.

The present paper is intended to satisfy the requirements
of formal publication of the new species, varieties and com-
binations contained in it. Additional Latin diagnoses have
been rendered in cases where available diagnoses seemed in-
adequate.

: 1. Juniperus chinensis L. Mant. 127 (1767), 519 (1771)5 non
. Roxb. Fl. Ind. Or. III, 840 (1832); non Miquel Fl.
Jap. 12, 58 (1670); tb<7126,-217 Figs. ts 25 4
(1844); not, or only in part, of authors since about
1650. e
J. flagelliformis Loud. Ency. Trees II, 1090 (1842).
J. Reevesiana Hort. ex Endl. Syn. Conif. 31 (1847) sub
Je cernua.:
- chinensis foemina Gord. Pinet. 115 (1858).
chinensis sylvestris Hort.
chinensis oblonga Hort.

ae ee eee eee

[S[>|5

Ret} ). Juniperus chinensis f. aurea (Young, var.) stat. nov.
J. chinensis var. aurea Young ex Anon. in Gdrs. Chron.

mitt, 1195 (1G72).°

1 (2). Juniperus chinensis f. WILSON'S WSSPING (Clarke, var.)
stat. nov.
J. chinensis Wilson's Weeping W. B. Clarke & Co., San
José, Cale, Catalog, "Garden Aristocrats" (1934).

2. Juniperus sphaerica Lindl. in Paxt. Flw. Gdn. I, 58
(1350) ex Farlat. in DC. Prodr. XVI, pt. 2, 488

(1865).
f J. chinensis 2 Smithii Loud. Arbor., Frut. IV, 2505
(1838); non Hort.

J. chinensis mas Gord. Pinet. 115 (1858).

2a. Juniperus sphaerice var. pseudo-mas var. nov.
J. chinensis columnaris viridis, J. chinensis viridis
of U. S. nurseries.
A speciei typo differt: Habitu angustiore pyramidali
vel anguste columnari; ramis perseveranter adscen-
dentibus. Floribus obscuriore monoicis, i.e., planta

185

186 Pay °C: t Oe Fe Vol. 2, noe 6

tardiore fructificante. Galbulis plerumque plusminus-
ve bilobatis apice plusminusve applanato saepe trans-
verse constricto. In formis borealibus habitu angusto,
statu juvenili protracto.

Type: Herb. van Melle, No. 93 - cultivated.

Habitat: Out of Chekiang Province northward into Hopeh
Province.

on (1): Juniperus sphaerica var. pseudo-mas f. columnaris
(U. S. Dept. Agri.) comb. nov.
Je chinensis var. columnaris U. S. Dept. Agri., Bureau
~ Plant Ind., Ninth Ann. List New Intro., No. 18577
(1920—21).

2b. Juniperus sphaerica var. dioica var. nov.
J. chinensis Beissn. in Nuov. Giorn. Bot. Ital. N. S.—

IV, 183—91, No. 10 (1897).

A speciei typo differt: Habitu anguste pyranidali vel
late pyramidali vel late ovoidei; ramis in formis an-
gustioribus (Sinam orientalem versus) perseveranter
adscendentibus, in formis latioris (Sinam occidental-
em versus) saepe ultimo patentibus vel plusminusve
undulatis. Floribus dioicis. Galbulis maturis biloba-
tis, apice applanato transverse constricto.

Type: Arnold Arbor., Giraldi, 1897, Shensi.

Specimens. Arnold Arbor.: Purdom No. 3005, southern
Shensi; Meyer No. 1712, Sianfu, Shensi.

Habitat: Qut of Hopen Province westward, along the
slopes of, and to the north of, the Tsinling Range,
into Shensi Province; projecting into Inner hiongolia.

2c. Juniperus sphaerica var. neaboriensis (Veitch) comb.
nOVe

J. neaboriensis Veitch Man. Conif. 277 (1881); non
Laws. ex Gord. Pinet. 96 (1858) sub J. macrocarpa;
non Fitschen Handb. Nadelh. 604 (1930) sub J. chinen-
sis.

Habitu anguste pyramidali; ramis adscendentibus. Foliis
acicularibus remotis brevibus plerumque late patenti-
bus. Floribus dioicis. Galbulis ut in var. dioica.

Type: Ue. Se Dept. Agri., Bureau Plant Indus., Meyer No.
2014, Sianfu, Shensi.

Habitat: Apparently more or less local, in southern
Shensi Frovince.

2d. Juniperus spheerica var. pendula (Franch.) comb. nov.

Je chinensis var. pendula Franch. in Nouv. Arch. Mus.
Hist. Nat. Paris Ser. 2, VII, 101 (1884); non
Beissn.; non Gaujard ex Morel in Rev. Hortic. 349—50

-

1946 van Melle, Junipers 187

(1889). Non J. sphaerica var. pendula Lav. Arbor. Se-
grez. 290 (1877) - nomen.

Habitu late plusminusve laxe pyramidali, ramis ultimo
patentibus undulatisve vel pendulis, certe ad apices;
rariore, ut in hortorum plantis, valde et irregulari-
ter pendulis. Ramulis pendulis. Floribus dioicis.
Galbulis regulariter obovoideis vel ovoideis. Statu
juvenili non protracto.

Habitat: Type locality (David), southern Shensi. Min
Shan Mountains, Lower Tebbu country.

2e. Juniperus spheerica var. Keteleeri (Hort.) comb. nov.
Je chinensis Keteleeri Hort. gall. ex Beissn. in Mit-
- theil. Deutsch. Den. Ges. 140 (1910).
J. sinensis var. Keteleeri Venema in Jaarbk, Nederl.
Den. Vereen. 108—21 (1938). :
Habitu anguste pyramidali; ramis adscendentibus. Foliis
squamiformibus apice acuto. Floribus dioicis. Galbul-
is magnis regulariter globosis. Statu juvenili non
protracto. Planta mascula in hortis ignota.

Type: N. Y. Bot. Gard., Henry No. 6576.

Type locality: Mountains of northern Hupeh Province.

5. Juniperus Sheppardii (Veitch, var.) sp. nov.
J. sphaerica glauca Fort. ex Gord. Pinet. 39 (1858).

J. sphaerica Sheppardii Veitch Man. Conif. 290 (1881).

Arbor axibus pluribus principalibus hebitu juvenili
regulariter, adulto plusminusve irregulariter laxe
lateque pyramidali; ramis adscendentibus, certe ad a-
pices. Statu juvenili non protracto. Aspectu comae
juvenilis ex griseo argenteo-griseo, adultae ex opace
griseo glaucescenti-viridi, rariore clare viridi. In
statu juvenili ramulis ultimo ad apices ramorum plus-
minusve congestis, in statu adulto secundum ramos
confertis. Surculis adultis terminalibus nutantibus.
In partibus mature adultis ramulis ultimis plerumque
gracilibus filiformibus saepe longissimis. Floribus
monoicis vel dioicis. Galbulorum maturorum "peduncul-
is" erectis patentibusve vel plusminusve arcuatis non
nutantibus. Galbulis maturis plerumque regulariter
obovoideis vel ovoideis rariore turbinetis, ad 10 mm.
latis. Seminibus 2—6, plerumque 2—4.

Type: Herb. van Melle, No. 272 (monoecious) - cultivat-
ed.

Specimens. Arnold Arbor.: S. Chen No. 3383, No. 4169,
Chekiang; Herb. Univ. Anhwei No. 2675, Fukien;
Hongkg, Herb., Dunn ixped. No. 3507, Fukien; Chiao No.
14059, Chekiang. -- Bailey Hortorium: R. ©. Ching No.
1601, No. 2018, Chekiang. -- U. S. Dept. Agri., Bur-

188 Poy TO: LAO: Gets Vol. 2, n6e10

eau Plant Indus., S. P. I.: Access. No. 1583, Kiang-
su; Re Ce Ching No. 1601, Chekiang.

Habitat: Chekiang Frovince, northward into Kiangsu;
southward into Fukien, Kiangsi. Anhwei.

3a. Juniperus Sheppardii var. torulosa (Zastw.) comb. nov.

J. chinensis var. torulosa Zastw. in Bay Cities Garden
Monthly II No. 4 (1933).

J. chinensis kaizuka Hort. jap.

A speciei typo differt: Axi principali plerumque uno
interdum diviso. Habitu angustiori. Statu juvenili
dense conico-fastigiati aspectu griseo-viridi. Sur-
culis adultis terminalibus non nutantibus plusminusve
suculentibus. Foliis squamiformibus intense viridibus
sed..in formis multis pruina grisea obtectis. Floribus
dioicis, Galbulis maturis forma valde variis: apice
contracto vel applanato, basi decurrenti vel trunca-
to; plerumaue approximate cylindricis vel quadrangu-,
laribus interdum regulariter obovoideis; longitudine’
latitudinem excedentibus; plerumque violaceo-pruinos-
is. Seminibus magnitudine valde variis.

Specimens. Arnold Arbor.: Wilson 10, 10, '14, Mt. Yaku-
shima; Wilson No. 8201, Senzu, Idzu Feninsule, Honsm.
-- Ne Y. Bot. Gard.: C. Wright, U. S. North-Fecific
axp. (1853—56), Shimoda, Idzu Feninsula; Maximowicz,
Iter Sec. 1862, Yokohama - cultivated; Maximowicz,
1863, Nagasaki.

Habitat: Southern parts of Korea. Japan, islands south
of Honshu; pernaps wild on Idzu Peninsula, Honshu.

3a (1). Juniperus Sheppardii var. torulosa f. aureo-variega-

ta (Hort.) comb. nov.

J. chinensis eureo-variegata of some U. S. nurseries.

3a (2). J. Sheppardii var. torulosa f. albo-notata nom. nov.
- J. chinensis albo-variegata of some, not all, Usa.
nurseries; non Veitch.

3b. Juniperus Sheppardii var. pyramidalis (Carr.) comb. nov.
Jd. chinensis Sieb. & Zucc. Fl. Jap. II, tt. 126, 127

figs. 1, 2, 4 (1844), text, Miquel, 58 (1870).

J. japonica Carriere Traité Gén. Conif. 33 (1855), pro

parte: "Arbrisseau dressé".

Je japonica var. pyramidalis Carr. 1. c., Hd. 2, Jl
(1867). |

J. chinensis pyramidalis Hort. ex Beissn. Handb. Nad-
elh. 120 (1591).

J. excelsa stricta of U. S. and Huropean nurseries.

Arbor plerumque axibus pluribus principalibus. A speci-

— ———

1946 van Melle, Junipers 189

ei typo differt: Statu juvenili valde protracto saepe
in senectutem retento, a adulto valde dissimili; juv-
enilis habitu primo dense conico-fastiziati ut in
var. torulosa sed denique late columnares valde regu-
lares, aspectu ex griseo viridi-griseo; ramulis ulti-
mo in thyrsis densissimis erectis ad apices ramorum
congestis. Habitu adulto laxe lato ultimo valde irr-
egulari pyramidsli vel diffuse arborescenti; ramis
late adscendentibus, ramulis valde irreguleriter dis-
positis; ramorum surculis terminalibus non nutanti-
bus. Floribus dioicis. Galbulis maturis plerumque ad
apicem plusminusve applanatis, regulariter vel irreg-
ulsriter 2—4-lobatis, plerumque latitudine longitud-
inem excedentibus.

Specimens. Herb. van Melle: No. 160, 160 A, B (juven-
ile); No. 55 (early adolescent); No. 38 (advencedly
adolescent); Nos. 88, 88 A, 162, 162 A, B, C, D, =
(early adult); No. 305, 305 A (fruiting).

Habitat: Conjecturally, central and eastern Honshu,
north to between 38° and 39° latitude. Frobably Kor-
ea, to 39° latitude.

4, x Juniperus media (J. Sabina x sphaerica) hybr. nov.

Frutices plerumque habitu J. Sabinam similis certe in
planterum juvenis. Foliis acicularibus interdum di-
morpnis: his brevibus textura mollibus ut in J. Sab-
ina, illis longioris rizidioribus ut in Js spnaerica.
Floribus dioicis. "Fedunculis" zalbulorum maturorum
semper nutentibus. Galbulis meturis plusminusve dis-
tincte bilobatis.

4a, x Juniperus media var. arbuscula nom. nov.
See iniana pendulis viridis Gord. Pinet. Suppl.
1862 ).

J. spheerica var. pendula Lavallée Arbor. Segrez. 290
(1877) - nomen. f

J+ virginiana Smithii pendula Hort. ex Beissn. Handb.

~ Nadelhe 125 (1891).

Je chinensis Smithii Slavin in Rept. Conif. Conf. R. H.
S. 103, (1932).

Frutex ramis primariis e trunco brevi adscendentibus,
primo fruticem erectum habitu laxo formantibus; pler-
umque habitu ultimo densiori ad apices ramorum plus-
minusve patenti. Statu juvenili non protracto. Plant-
arum juvenum foliis grate fragrantibus, squamiformi-
bus clare viridibus textura delicatula; plantarum
senescentium plusminusve opace viridibus textura dur-
iora. Flanta mascula ignota. Galbulis bilobatis ad 10
mne latis "pedunculos" graciles cernuos terminantib-

190 PHYTOLOGIA Vole 2, no. 6
us. Seminibus ad 6, ad 6.5 mm. longis plusminusve
tenuibus.

Type: Herb. van Melle, No. 283, cultivated in N. Y.
Bot. Gard.

Habitat unknown.

4b. x Juniperus media var. Pfitzeriana (Beissn.) comb. nov.

J. japonica pendula Morren & A. de Vos, Index Bibliogr.
in Bull. Fed. Société Hortic. Belg. 32 (1877) -
nomen.

J. chinensis pendula C. de Vos Handb. Boomen, Heest.
(1885); Beissn. System. Zintheil. (1887) - nomen;
Gaujard ex Morel in Rev. Hortic. 349 (1889); Fitsch-
en, Handb. Nadelh. 606 (1930); non var. pendula
Franchet.

J. chinensis Pfitzeriana Beissn. in Mittheil. Deutsch.

~ Den. Ges. 102 (1899).

Frutex odore J. Sabinae. Ramis primariis. basi oblisue
divergentibus, arcuato patentibus; ramis ramulisque
lateralibus ad apices arcuato-pendulis. Foliis squam-
iformibus ad apicem acutis, ex griseo-viridis ultimo
(in senectute ) opace viridibus. Galbulis maturis ad
10 mm. latis "“pedunculos" graciles cernuos terminant-
ibus. Seminibus 2—4, ad 5 m. longis quem in J.
sphaerica tenuioribus. Planta femina in hortis ignota.

Type: U. S. Nat. Herb., No. 1245121, Ching No. 52.

Habitat: Ho Lan Shan Mountains, Inner Mongolia.

4b (1). x Juniperus media var. Pfitzeriana f. aurea (Hort.)
combe NOVe
Je chinensis aurea pendula Beissn. Handb. Nadelh. 120
(1891).
J. chinensis Pfitzeriana aurea D. Hill Nurs. Co., Cata-

log (1938).
fe (ede = Juniperus media var. Pfitzeriana f. compacta

(Hort. ) comb. nov.
J. chinensis Ffitzeriana compacta Bobbink & Atkins,

~ Catalog (undated).
4b (5). x ee media var. Pfitzeriana f. Armstrongii

(Bailey) comb. nov.

Armstrong Spreading Juniper Armstr. Nurs., Catalog.
(1932).
J, chinensis Armstrongii Bailey Hortus Suppl. (1935).

4b (4). x Juniperus media var. Pfitzeriana f. glauca (Hort.)
comb. nov.
Silver-blue Juniperus U. S. Plant Patent No. 422 (1940).

1946 van Melle, Junipers 191
J. chinensis Pfitzeriana glauca Hort.

4c. x Juniperus media var. globosa (Hornibr.) combe nove

Je virginalis globosa Hort. jap. - pro parte.

Je chinensis nana Hort.

Je japonica globosa, J. japonica nana of Boskoop dis-
tribution.

J. japonica Bandai-sugi Hort. jap.

Js chinensis var. globosa Hornibr. Dwarf and Slow-gr.
Conif. 62 (1923).

Frutex humilis odore J. Sabinae. In plantis juvenis
ramis primariis oblique divergentibus apice non pend-
ulo. Ramulis non pendulis. Stature rare 1 m. superan-
te; ultimo plusminusve 2.5 m. latus. Foliis squami-
formibus e pallide praeclare viridibus, apicem obtuso
vel acuto. Foliorum ramulorum textura delicetula.
Galbulorum maturorum "pedunculis" quam in J. Sabina
brevioribus. Galbulis ad 5 mm. latis, latitudine lon-
gitudinem excedentibus, forma variis: apice rotundato
vel plusminusve applanato; valde irregulariter gibbo-
sis vel obscure bilobatis; vix pruinosis. Seminibus
plerumque 2—4, saepe 1, ad 2.5 mm. longis. Flanta
mascula ignota.

Type: Herb. van “elle, No. 260, cultivated in Arnold
Arbor. as J. chinensis var. plumosa.

Habitat unknown.

Ac (1). x Juniperus media var. globosa f. aureo-globosa

(Rehd. var.) comb. nov.

J. chinensis procumbens aurea Hort. ex Beissn. Handb.
Nadelh. 120 (1891) — pro parte.

J. japonica globosa aurea of Boskoop distribution.
Je chinensis var. aureo-globosa Rehd. Manual (1923).

ce chinensis globosa f. aurea Hornibr. Dwf. & Slow-gr.
Conif. 63 (1923).

J. japonica Bandsi-sugi aurea Hort. jap.

4d. x Juniperus media var. plumosa (Hornibr.) comb. nov.

J» chinensis procumbens Sndl. ex Beissn. Handb. Nadelh.
120 (1891), and of Boskoop distribution; non Zndlich-
er.

J. virginalis globosa Hort. jap. - pro parte.

J. chinensis var. plumosa Hornibr., 1. c. 66 (1923).

Je japonica of Boskoop distribution, and of many nur-
series today; non Carr.; not clearly of any author.

J. chinensis L.Ching No. 53 ex E. H. Walker, Flants
Coll. Re C. Ching, in Contrib. U. S. Nat. Herb.
XXVIII, pt. 4, 594 (1941); non L.

J. virginalis D. Hill Nurs. Co., Catalog (1942).

192 PH YT Ob OG. & Vol. 2, nide-e

Frutex nanus odore J. Sabinae. In plantis juvenis ramis
primariis oblique diverzgentibus non arcuatis. Ramis
rigidibus; ramis ramulisque ad apices non pendulis.
Statura ad 1.5 x 1.5 m. Surculis adultis terminalibus
plusminusve suculentibus; ramulis lateralibus brevi-
bus plusminusve rigidibus' saepe apice plusminusve ro-
tundato convexo. Foliis squamiformibus, apice obtusi-
usculo vel rotundato, saepe ad dorsum valde convexis;
aspectu plerumque opace viridi. Flanta femina ignota.

Type: Arnold Arbor., Re C. Ching No. 53

Habitat: Ho Lan Shan Mountains, Inner Mongolia.

4d (1). x Juniperus media var. plumosa f. albo-variegata

(Hort. ) comb. nov.

Je chinensis procumbens albo-variegata Hort. ex Beissn.
Ls Gace. (1891).

Je chinensis var. decumbens albo-veriegatea Hornibre. le
Ga 66 (1923).

Je chinensis var. plumosa albo-veriegata Hornibr. ts 66
ys eee 106 (1938),

J. japonica albo-variegata of Boskoop distribution.

4d (2). x Juniperus media var. plumosa f. aurate nom. nov.
J. chinensis procumbens aurea Hort. ex Beissne le Ge
120 (1891) - pro parte.
Je japonica aurea of Boskoop distribution.
J. chinensis var. plumosa aurea Hornibr. 1. c. 66

~ (1923).

ha (3). =x Juniperus media var. plumosa f. aureo-variegata

(Hort.) comb. nov.

J. chinensis procumbens aureo-variegata Beissn. Lia

Set (eo).

J. chinensis var. decumbens sureo-variegata Hornibr. le
c. 66 (1923).

J. chinensis var. plumosa aureo-variegata Hornibre. leC.
Ed..2, 205 (1938),

J. japonica aureo-variegata, J. chinensis procumbens
aureo-veriegata of Boskoop distribution.

Sa. Juniperus davurica Pallas var. Parsonsii (Hornibr.)
comb. nOv.e
J. chinensis var. japonica Lav. ex Slavin in Rept.
Conif. Conf. R. He S. 102 (1932) ?; non Lavallée; non
ex Hornibr. 1. ce. 100 No. I (1938); non Vilmorin ex
Wils. Conif. Tax. Japan 85 (1916); non Vilmorin.
chinensis var. Parsonsii Homibr. l. ce. 96 (1938).
squamata Farsonsii Bailey Hortus Sec. (1940).
squamata of some U. S. nurseries; non Lamb.

a al

1946 ven Melle, Junipers 193

J. squamata prostrata of some U. S. nurseries; non
Hornibr.

Frutex humilis denique cumulum depressum regulariter’
rotundatum formans; gratissime fragrans. Ramis prim-
ariis inxplantis juvenis horizontesliter patentibus
sed non ad humum adpressis nec decumbentibus. Ramis
velidis valde rigidibus, cortice laeve cinnamomeo
lamelloso, ramulos longitudine gradatos subbilaterale
distributione emittentibus. Foliis acicularibus oppo-
sitis vel ternatis; squamiformibus glaucescenti-
viridibus vel opace viridibus, apice acutiusculo vel
obtuso. Ramorum surculis terminalibus plusminusve
suculentibus non nutantibus nec adscendentibus. Ramu-
lis ultimis adultis gracilibus filiformibus saepe
longissimis. Floribus dioicis. Galbulorum maturorum
"nedunculis" nutantibus. Galbulis maturis depresso
globosis vel 3—4-gibbosis saepe approximate quadran-
guleribus usque ad maturitatem valde coerulei pruino-
sis, ad 10 mm. latis. Seminibus vulgo ad 6, ad 5 mm.
longis.-

‘Type: Herb. van Melle, No. 107, containing material
from two plants - one male, one female.

Habitat: Conjecturally, central Korea; Japan.

combe nNOVe
Je chinensis L. var. japonica Lav. f. alba Rehd. in
Journ. Arnold Arbor. VI, 202 (1925) ? ,
Je chinensis var. expansa variegata Hornibr. le Ce 9
(1938). -
Je squamata var. albo-variegata and var. variegata Bai-
ley Hortus Sec. (1940), and of some U. S. nurseries.
Je squama ta argenteo-variegata of some U. S. nurseries

5a (1). Juniperus davurica var. Farsonsii f. variegata
(Hort. )

6. Juniperus Sargentii (Henry, var.) Takeda ex Nakai Fl.
Jap. Kor. in Tokyo Bot. Mag. XLIV, S11 (1930); ex
Koidzumi in Tokyo Bot. Mag. XXXIII, 204 (1919) - no-
men; non Sasaki, List, Plants Formosa 53 (1926) - no-
men (fide Masamune in Mem. Fac. Science & Agri, Tai-
hoku Imper. Univer. XI, Dec. 1934, p. 131); doubtful-
ly of many Jeépanese floras. i

de chinensis var. procumbens Nakai Veget. Quelpt. Isl.
13 (1914); Takeda Fl. Isl. Shikotan in Journ. Linn.
Soc. XLII, 486 (1914); Miyabe & Miyeke Fl. Seghalin
593 (1915); non Endl.; non Beissn.; not Sabina chin-
ensis var. procumbens Antoine Cupress. Gatt. (1857).

J. davurica Nakai in Tokyo Bot. Mag. XXX, 2 (O17):
and of several other Japanese authors, in part; non
Pallas.

194 PH Y-20470.6-78 Vol. 2, Hos a

Frutex humilis odore terebinthaceo, denique cumulum de-
pressum plusminusve irreguleriter rotundatum formans,
Ramis primariis prostratis cortice cinnamomeo, ramu-
los longitudine gradatos subbilaterale distributione
adscendentes vel suberectos apicibus patentibus emit-
tentibus. Foliis acicularibus brevibus non rigidis;
squamiformibus apice obtusiusculo vel rotundato, ex
opace viridibus griseo-viridibus. Ramulis ultimis ead-
ultis vix longis vix filiformibus. Floribus dioicis.
Galbulorum maturorum "pedunculis" nutantibus. Galbul-
is meturis forme variis, saepe approximate globosis
apice plusminusve applanato, plusminusve obscure bi-
lobatis; ante maturitatem ex viridibus griseo-viridi-
bus, saepe ad epicem brunneo-violaceis, vix pruinos-
is; plerumque ad & mn. latis. Seminibus seepe ad 5
(in plantis tsushimensis, leg. Wilson, ad 8).

Habitat: Mainly coastal and insular, Korea, Japan, Sak- *
halin.

7. Juniperus procumbens Sieb. & Zucc. Fl. Jap. Ih). tae
fig. 3 (1844), text, Miquel, 59 (1870); non Sargt. in
Gdn. & Forest X, 421 (1897).

J. chinensis var. procumbens Endl. Synopsis Conif. 20

~ (1847); non Beissn.; non Nakai Vegt. Quelpt. Isl.
(1914); non Takeda Fl. Isl. Shikotan (1914); non Mi-
yabe & Miyake Fl. Saghalin (1915); non Sabina chinen-
sis var. procumbens Antoine Cupress. Gatt. (1857).

J. japonica Hort. ex Carr. Traité Gén. Conif. 33
(1855), in part; ex Héring Manuel Plantes IV, 314
(1857), in part; non Sargt. in Gdn. & Forest X, 421
(1897).

Js recurva var. squamata Masters in Bull. Herb. Boisse
VI, 274 (1898); Matsumura Index Plants Jap. 11
(1905); non Parlatore.

J. chinensis var. japonica Vilmorin Hort. Vilmor. 58
(1906); non ex Wilson, Conif., Tax. Japan 85 (1916).

7a. Juniperus procumbens f. nana (Hornibr., var.) stat. nov.
Js japonica nana D. Hill Nursery Co., Catalogs up to
19 os :
J. procumbens var. nana Grootendorst ex Hornibr. Dwarf
& Slow-gr. Conif. 122 (1938).

Incompletely Known:

Juniperus chinensis var. Luptonii Hort.
Of this juniper I have seen cultivated plants up to 10

feet high, but never normally developed ones; nor
have I seen fruit of it. I have not been able to i-

ie

1946 van Melle, Junipers iw

dentify it with herbarium material collected in the
wild. I diagnose it tentatively as one of the inland,
dioecious developments of J. sphaerica.
Juniperus chinensis ver. arenaria Wilson ex Rehd. & Wils.,
Plants Coll. J. Fe Rock, in Journ. Arnold Arbor. IX,
20 (1928).
From the description cited, and from specimens eat the
Arnold Arboretum I cannot diagnose this juniper. I
- gee, however, no sufficient grounde for regarding it
as a variety of J. chinensis or J. sphaerica or J.

She ppardii.

Juniperus chinensis var. tsukusiensis Masamune Frelim. Rept.
Veget. Isl. Yakushima 39 (1929) - nomen; in Journ.
Soc. Trop. Agri. (Taihoku Imper. Univ.) II, 152
(1930).

I have seen no specimens of this juniper. The descrip-
tion rendered by Masamune seems to me not to contain
sufficient grounds for its identification as a varie-
ty of J. chinensis.

i —_— « _-_ — -_- —_—_ ll —_ _ -_- sl = ~_

ADDITIONAL NOTES ON THE GENUS FSTREA. II

Harold N. Moldenke

PETREA VOLUBILIS L.

Calderon and Standley list for this species the common
names "flor de Jesus", “lengua de vaca", and "adolfina".
Conzatti and Sanchez report that in Mexico it is called
"yerba del soltero". The New York World Telegram article
cited in. the previous installment of these notes calls the
plant "petrea".

Detailed descriptions of the floral morphology are given
by Junell in the reference cited in the previous install-
ment, and other morphological notes will be found in
Svensk. Bot. Tidsk. 32: 231. Macmillan describes the plant
as blooming twice a year, and the calyx as bright helio-
trope in color, "persisting long after the violet corollas
have fallen." Popenoe describes the species as a half-
climbing shrub, 15 feet tall, with blue flowers, "a rather
common cultivated ornamental of the Cauca valley, Colombia}
and "occasional in gardens, rare" in Florida. The lundells
describe the calyx as "bluish-purple, pale" or "“bluish-
lavender" and-the corolla as "purplish". It has been col-

196 PWY F000 Gor Vol. 2, noe 6

lected in anthesis also in September. Hinton says that it
grows in the sun in the mixed forests of Guerrero. Standley
describes it as a "large woody vine" or a "small vine, with
blue-purple flowers, in dry thickets" in Guatemala, and C.
L. Wilson reports it as a “liana common in the virgin for-
ests" of the same country. LeSueur collected it on dunes in
Tamaulipas; Steyermark as a shrub 10 feet tall on moist
banks in Jalapa, Guatemala; Pringle found it "running over
rocks on limestone ledges" in San Luis Potosi. Fruit has
been collected in September. The beauty of the plant is well
indicated by the statement in the New York World Telegram
article previous!y referred to, where it is reported that
"Mrs. Paine [a director of the Garden Club of America] glows
with enthusiasm as she describes more of Costa Rica's lush
plants and flowers, such as 'petrea', a kind of blue vine

undetermined", Mexico, is actually from either Osxaca or
Veracruz, according to information received by me recently
from Professor Martinez; Liebmann 112835 is from Veracruz;
Liebmann 11285 is probably from Veracruz; Schiede & Deppe se
ne [Artopaz, Mart. 29] is probably from Actopan, Veracruz;
and Schiede & Deppe sen. [Malpayo.de Hautingo, April 29] is
probably from Huetengo, Hidalgo. The Sturrock s.n. [Finca
Mulgaba, 1916] cited on page 43 as from "Province undeter-
mined", Cuba, is actually from Havana.

Paxton in the reference cited in the previous installment
of these notes gives the conmon name "stapelia-flower petre-
a" for this species. The Buswell specimen cited below bears
no indication on its label that it was collected from culti-
vated material, but the collector writes me thet it actually
was. The Jamaican specimen cited may be from cultivated mat-
erial, but there is no indication on the label to this:ef-
fect. The synonymy given by Ch. Crevost and A. Fételot in
the reference cited is that of P. Kohautiana, but the illus-
tration is definitely P. volubilis. The Forbes reference is
extremely interesting in that it records the finding of this
plant in a wild state in Timor and Java. A quotation from
pages 78—79 is worth repeating here: "On one of the lower”
knolls I found perhaps the most interesting plant in my Jav-
an collection, a species of Petraea (P. arborea) growing en-
tirely wild in the forest. This genus.......is almost en-
tirely confined to the South American continent and it is of
extreme interest to find it, in this inexplicable way, crop-
ping up in a region so far removed from the centre of its
distribution. A species from the island of Timor occurs,
witnout history, in the collection in the British Museum

1946 Moldenke, Notes on Petrea 197

made by Mr. Robert Brown, but these are the only two examp-
les, so far as I am aware, hitherto collected uncultivated
in the Old World."
Additional illustrations: Rehnelt, Gartenwelt 28: 367,
figs. 1924; Nessel, Gartenfl. 75: 321—322, fig. 1926; Cre-
vost & Pételot, Bull. Econ. Indo-chine 37: 1289 [as "FP. sub-
serrata" ]. 1934; Junell, Symb. Bot. Upsal. 4: figs. 80—86.
1934; H. Fe Macmillan, Trop. Planting & Gard., ed. 5, 122.
1943.

Additional citations: CUBA: Oriente: Leon 17259 (Ha, N).
JAMAICA: Hatch sen. [August 8, 1932] (F1--105108). PUERTO
RICO: Otero 252 (N)- MEXICO: Coahuila: Artamanoff s.n. (F).
Guerrero: Hinton 14135 (N, N); EB. We Nelson 2318 (F); adw.
Palmer 395 (Ca, F, Me, Me, Me, Me). Michoacén: Emrick 2557

bis (F). Oaxaca: ee a 2100 (Me, Me), 5305 (N); Con-
zatti & Sanchez 3 342} (Me ); Conzatti, Reko, & Makrinius ~ 3001
(Me); Galeotti roe erie coll. of Fetraea ovata Mart. &
Gal.] (Br, N--photo, Z--photo). Puebla: F. Salazar sen.
{Huauchinango] (Me). San Luis Fotos{: fdw. Palmer 1064 (Io);
Pringle 5003 (Fs; Me, Me, Mi), 5003 1/2 (V vt), 8004 (cm, F,
Io, It, Me, Me, Po). Tamaulipas: Berlendier 136 [type Olle
of "Fetraea (volubilis?) mexicana Cham."] (Du--166365, N--
photo, Z--photo), 182 (Lu); LeSueur 542 (Au, F); Sdw. Falmer
279 (F), 317 (F). Veracruz: Galeotti 795 [type coll. of P.
mexicana H.B.K.] (Br); Matuda 1478 (Mi, N); Medellin 17
(Me); Orcutt ae (Du--155196); FPurpus 6354 (F), 13677 (Du--
184943); C. L. Smith 1017 (N, Vt). Yucatan: G. F. Gaumer 379
(F); Lundell & Lundell 7329 (Mi, N), 7413 (Mi, N); _Steggerda
27a (F). State undetermined: Haenke 1582 (N); Sessé, Mocifio,
Castillo, & Maldonado 2225 (F). GUATaMALA: Alta Verapaz: Te
L. Wilson 334 (F). Chimaltenango: J R. Johnston 1149 (F)._
El Feten: Aguilar Hidalgo 362 (I); H. H. Bartlett 12133 (F),
12559 (Ca, I); C. L. Lundell 3451 ones Izabal: Bur. Pl. Ind.
Sone sen. [1922] (Ar); He He V. Johnson 1265 (La, Ia). Jalapa: Stey-
ermark 32957 (F). Retalhuleu: P. C. Standley 88384 (N).
BRITISH HONDURAS: Chanek 143 (F); Gentle 2366 (Dp, Mi), sn.
[C. Le Lundell 4972] (Ca, Hp, I). COSTA RICA: Alajuela:
Brenes 13620 (F), 14290 (F), 20462 (F), 20468 (F); Orozco

16 (F). Guanacaste: C. W. Dodge 6474 (F); M. Valerio 513
(F). PHILIPPINZ ISLANDS: fags Me (. Ramos Sone {Herb. Fhil-
ipp. Bure Sci. 12194] (Ar). CULTIVAT=D: Bahamas: Degener
18788 (Ml, N). Colombia: Fopenoe 1214 (Ar). Cuba: Leon 41
(Ha); Popenoe 427 (Ar); Van Hermann 2674 (Fo). pee Rica:
M. Valerio 63 (F). Florida: Bur. Fl. Ind. S. P. 1. 36024
(Ar, Ar); Buswell s.n. [March 8, 8, 1939] ] (Bu); Mo lowry - West s.
n. [19 May 1932] (Fl); Popenoe 236 (Ar); Ricker (Any
Simmonds sen. [Bur. Pl. Ind. Se Pe I. 36024] rye gt

la: L. H. Bailey 579 (N); FP. 0. Standley ese (F); Steyer-
mark - 39880 (F). Hawaiian Islands: Degener 1/7851 (N); O. Be

198 PH YR OL Ors Vol. 2, now G

Johnson sene (Honolulu, 1897] (P1--22595, Se--14931, Se--
14932). Hispaniola: g. C. Leonard 10143 (Ca). Honduras: Yun-
cker 4524 (Dp). Mexico: Conzatti 5305 (Mi). Netherlands:
Herb. Hort. Clifford s.n. [Herb. Linnaeus G.78l, Sel] (E--

photo of isotype).

PITREA VOLUBILIS var. ALBIFLORA (Standl.) Moldenke

Synonymy: Fetrea volubilis f. albiflora (Standl.) Standl.,
Field Mus. Publ. Bot. 18: 1012. 1938.

References: Standl., Field Mus. Publ. Bot. 18: 1012.
1938; Moldenke, Prelim. Alph. List Invalid Names 34. 1940;
Moldenke, Alph. List Invalid Names 35. 1942; Moldenke, Known
Geogr. Distrib. Verbenac. 21 & 97. 1942; H. F. Macmillan,
Trop. Flanting & Gard., ed. 5, 122. 1943,

The type collection was orizinally identified and dis-
tributed as P. arborea H.B.K. Schipp states that the variety
grows in open forests. It has been collected in flower and
fruit in March.

FETREA VOLUBILIS var. FUBZSCENS Moldenke

Edwards describes the variety as inhabiting open mountain

forests; Pringle found it running over rocks on limestone

ledges, at an altitude of 400 feet. It has been collected
in sites in March. The Collector undesignated 611 and Lan-
kester sen. [El Rodeo] cited by me on page 46 of my mono-
graph as from "Province undetermined", Costa Rica, are both
actuelly from San José. The-variety nas been conta in the
one with "Petraea arborea H.B.K." Tne comnon name "jasmin
o'Coamecate azul" is recorded by Urbina.

Additional citations: MEXICO: Hidalgo: Urbina sen. (Me ).
Oaxaca: Seler & Seler 1777 (Du--283661). San Luis Potos{: &
Fringle 8004, in part (Vt). HONDURAS: Comayagua: J. Be Ed- M
wards P.586 oo P.601 (F). COSTA RICA: Province undetermin-
ed: Pittier s.n. (Herb. Instit. Physico-geogr. Nat. Costar-

ic. 16655] (Cm).

ADDITIONAL NOTES ON THE GENUS AMASONIA. I

Harold N. Moldenke

Since the publication of my monograph of-.this genus in
Fedde, Repert. Sp. Nov. 46: 193—228 (1939) twenty-nine add-
itional specimens and photographs of specimens have come to
my hands. This surprisingly small amount of material to
come in during seven years is a fair index of the paucity of

1946 Moldenke, Notes on Amasonia 199

herbarium specimens of this genus to be found in the world's
herbaria. The new material is deposited in the herbaria in-
dicated by the following symbols: F = Chicago Naturel Hist-
ory Museum, Chicago; Ja = Museu Necional, Rio de Janeiro; Je
= J. Cuatrecasas Herbarium, Cali, Colombia; Kr = Krukoff
Herbarium, New York Botanical Garden, New York; Mi = Univer-
sity of Michigan, Ann Arbor; N = Britton Herbarium, New York
Botanical Garden, New York; and W = United States National
Herbarium, Smithsonian Institution, Washington.

AMASONIA L. f.

References: A. Le Juss., Gen. Fl. 119—12%3. 1789; Neck.,
BZlem. Bot. 1: 362—389. 1790; Wittstein, Etymolog.-bot.
Handworterb. 34. 1852; Bentham in Benth. & Hook. f., Gen.
Pl. 2: 1147. 1876; Junell, Symb. Bot. Upsal. 4: 107. 1934;
Moldenke, Frelim. Alph. List Invalid Names 4—5 & 42. 1940;
Moldenke, Known Geogr. Distrib. Verbenac. 30, 32, 33, 36,
40, 71, & 86. 1942; Moldenke, Alph. List Invalid Names 4,
23, & 43. 1942; Prytologia 2: 91. 1945.

The zeneric name is erroneously accredited to Linné the
elder by Wittstein in the reference cited above. Three add-
itional common names for members of the genus are recorded:
"taligale", "“amasonée", and "duphysteme", the first by Juss-
ieu and the two latter by Necker in the references cited a-
bove.

AMASONIA ANGUSTIFOLIA Mart. & Schau.

References: Junell, Symb. Bot. Unsal. 4: 107. 1934; Mol-
denke, Frelim. Alph. List Invalid Names 42. 1940; Moldenke,
Known Geogr. Distrib. Verbenac. 36 & 86. 1942; Moldenke,
Alph. List Invalid Names 4 & 43. 1942.

Additional citstions: BRAZIL: Goyaz: G. Gardner 3411
(Herb. Monac. 924; Macbride photos 20345 & 28390] (F--photo
of isotype, Kr--photo of type, Kr--photo of isotype).

AMASONIA ARBOREA H.B.K.

The type collection of this species was gathered in wood-
ed places near Javita, on the banks of the Rfo Tuamini, Mis-
Ssiones del Orinoco, Venezuela. Pinkus describes the species
as a shrub to 3 feet tall. The corolla is described as yell-
ow or pale-yellow, the bracts as red or scarlet. It has been
collected in fruit in May and September. In Colombia it as-
cends to 240 m. Williams says that it inhabits clearings on
"terra firma", while Pinkus found it in rocky soil of thick
forests.

Additional citations: COLOMBIA: Vaupes: Quatrecasas 7084
(Jc). VENEZUELA: Amazonas: Cardona 166 (W); Ll. Williams
15175 (W). BRITISH GUIANA: A. S. Pinkus 2 (N).'

a oe

200 PP. Ek OL -Oie ee Vol. 2, noe 6 |
;

AMASONIA CALYCINA Hook. f.

References: J. D. Hooker in Curtis, Bot. Mag. 113: pl.
6915. 1887; Moldenke, Known Geogr. Distrib. Verbenac. 33,71,
& 86. 1942; Moldenke, Alph. List Invalid Names 43. 1942.

Hooker, in the reference cited above, states that this
species is ectually a native of British Guiana, but to date
-I have seen only cultivated material from botenical gardens
in Austria, Belgium, ingland, Italy, New York, and Trinidad.

AMASONIA CAMPSSTRIS (Aubl.) Moldenke |
References: Willd., Sp. Pl. 3: 394. 1800; Benth., Ann. ¥
Nat. Hist. 2: 450. 1838; Griseb., Fl. Brit. W. Ind. 50m ;
1861; Junell, Symb. Bot. Upsal. 4: 107 [as A. erecta]. 1934; ~~
“Moldenke, Frelim. Alph. List Invalid Names 4, 5, & 42. 1940;
Fulle, Fl. Suriname 4 (2): 283—284. 1940; Pittier, La Mesa j
de Guanipa 23 & 45 [as A. punicea]. 1942; Moldenke, Known )
Geogr. Distrib. Verbenac. 72, 33, 36, & 86. 1942; Moldenke,
Alph. List Invalid Names 4 & 43. 1942; Phytologia 2: 91.
Pons <6). ‘
Froes describes the species as a "low shrub". Monteiro da
Costa states that the flowers are red, but he certainly~-
means this to be a descrirtion of the bracts, not the flow-
ers. Bentham, in the reference cited above, says for "Re er-
ecta": "The corolla is said by Schomburgk and Vahl to be
red, by Aublet to be yellow". Here again the reference to
a "red" corollas must be an error for the bracts. Monteiro da
Costa says that the species innabits low land and reports
the vernaculer name "herva de picapao", while Drouet records
it as innabiting open woods. Pittier, in the reference cited
above, lists the species as a constituent of the "sabanas de
saetas" in Venezuela. Willdenow records the common names
"sufrechte Amasonie" and "rothe Amasonie". The label of Mac-
bride photo 22773, cited below, reads "Cen. America" in er-
ror. The plant of which this is a photograph was collected
on the island of Trinidad.

Illustrations: Junell, Symb. Bot. Upsal. 4: pl. 7, fig. 2
[as A. erecta]. 1934.

Additional citations: TRINIDAD: Ryan s.n. [Macbride
photos 22773; type coll. of A. punicea] (Kr--photo). BRITISH
GUIANA: A. C. Smith 2441 (F). BRAZIL: Bahia: Blanchet 3156
[Macbride photos 7887 & 30184; type coll. of A. velutina]
(F, F--photo, Kr--photo, Kr--photo). Ceara: Luetzelbur
26095 (F). Maranh&o: Froes 1862 (F, Mi), 11779 (N), 11790
(N). Para: Drouet 2125 5 (F); Monteiro da Costa osta 263 (F).

me oe

Te ae ea eS ee ee ee ee

A.

AMASONIA CAMPESTRIS var. SURINAMENSIS Moldenke

References: Moldenke, List Geogr. Distrib. Verbenac. 21,
nom. nud. 1939; Pulle, Fl. Suriname 4 (2): 282, 284, & 285.
1940 ;Moldenke,Known Geogr. Distrib. Verbenac. 33 & 86. 1942.

7 : i=

PHY! | |
PHY TOLOGIA
bee Designed to expedite botanical publication

ne April, 1947 No. 7

CONTENTS
H. A. GLEASON, The preservation of well-known
J. V. Monacuino, A new species of Daphnopsis

DimomMials: .........0reesesseesssensesserecsesvesseenssosonnessvaseersunnerser QO
PYOUAEPEMAR OT Ga so oid cursesaxo-saees Asia cess Vie bubs apple c aie

TAMIR eae ccederayttie pessvasbus eaters saseesbnasanetdpnedn QD

SINE idols sacks savidod pualdaterwsses caeivine aioe cava to oa

N. MOLpDENKE, Additional notes on the genus
ee i Amasontas ED ..lo.issoiacicn Rad ect Lexis gs

| . MoLDENKE, Additional notes on the genus
PIPER TTT Feith ta Bea US Saco eetacsobedearbon eee

a

Published by H. A. Gleason and Harold N. Moldenke
The New York Botanical Garden,
Bronx Park, New York 58, N. Y.

LIBRARY
NEW YORE
BOTANICAL

GARDEN

THE PRESERVATION OF WELL KNOWN BINOMIALS

He Ae Gleason

The International Code of Botanical Nomenclature, as re-
vised at Cambridge in 1930 and further amended at Amsterdam
in 1935, is now followed by all working taxonomists in Amer-
ica. Nevertheless, there are certain facts about the Code
and certain principles involved in its provisions which are
not always understood by botanists, especially by non-
taxonomists, and not always appreciated by the taxonomists
themselves.
One of these relates to the history of codes in general
but especially to the so-called Feris Code of 1867, since it
is the direct progenitor, in a figurative sense, of the mod-
ern code of 1935. The Faris Code was the first formulation
of nomenclatural principles and rules for which the claim of
internationality was made and to which adherence by all tax-
onomists was expected. In the Feris Code the principle of
_ priority was the leading feature, just as it remains today.
But those who care to study the code carefully and to in-
quire into the circumstances of that period which led to the
appointment of De Candolle to draft the code will at once
realize that absolute priority was not intended and that the
effect of absolute priority was probably not imagined. If
there was.a conflict in the general usage of names in tha
various countries of western Europe (America apparently re-
ceived little or no consideration), the choice of the con-
flicting names should depend on priority of publication,
other things being equal. De Candolle never insisted on in-
vestigation of the merits of all published names: those that
had alresdy been relegated to the nomenclatural waste-basket
were better left there undisturbed.

Other persons doubtless realized the potential danger in
a strict interpretation of the rules. Some readers will re-
member the presidential address of Le H. Bailey before the
American Society of Flant Taxonomists, in wnichn he told of
finding the Faris Code on the library shelves at Harvard and
his proposal to Asa Gray that he (Bailey) translate them into
English. To which Gray replied "Mr. Bailey, you will do no
such thing. Let sleeping dogs lie."

Yet Asa Gray followed the principle of priority. If there
* was a choice to be made between two or more names, it was
_ his prevailing prectice to adopt the oldest. And so far as I
» know, he did so without mentioning them as the justification
_ for his action.
z About twenty years elapsed before anyone aroused the
ype 32) 8

202 PHT O° L:00 Ie Vol. 2, noe 7

sleeping dog. Nathaniel Lord Britton, my former professor
and for many years my superior officer at the New York Bot-
anical Garden, whose botanical ability, measured by his ac-
complishments, stands second to none in the country, first
attempted to follow the provisions of the code beyond its
original intent. In the late eighties and nineties he, some-
times alone and sonetimes with assistants, hunted out hun-
dreds of forgotten or discarded specific epithets, combined
them with the valid generic names, and introduced the new
combinations to the botanical public.

Of course there was a storm of protest, although Britton
was right, according to the provisions of the current Inter-
national Code. But the gates were now open and the flood-
waters of nomenclature inundated the fields of taxonomy. Af-
ter fifty years of drainage, after forty years of damming by
nomina conservanda, those fields are still miry. Hardly an
issue of Rnodora appears in which e change of name of some
eastern American plant is not proposed, strictly in accord-
ance with the code, of courses. Some of these authors, who
now stand on technicalities of the code, might well remember
that their own predecessors were among the loudest in con-
demnation of Britton, who also was guided by similar techni-
calities in the code of his day.

Tne first attempt to restore nomenclature to some degree
of sanity came with the codes of 1905 and 1910. In them
there was no change from the early provision for the use of
the oldest valid specific epithet, which was the prime cause
of the trouble, but an attempt was made to reduce the effect
of this provision. The use of tautonyms was abolished; epi-
thets used in one category were not required to be transfer-
red to another category; a number of generic nomina conser-
vanda were adopted; a starting point later than 17535 was
fixed for certain groups. Each of these provisions tended to
restrict the damage caused by the discovery of unknown names
or the revival of forgotten ones. All of them have been con-
tinued in the code of 1955 and the number of nomina conser-
vanda has been increased.

Of late years a new dam has been opened, again to flood
taxonomy. Tnis is the problem of typification, not yet thor-
oughly controlled by the recent codes. The waters swirl
round and round between Quercus rubra and Quercus borealis;
between Euphorbia maculata and Euphorbia supina, leaving
marooned and helpless the poor botenist who uses names as
appellations for plants end not as botanical footballs.

Football players are invited to consider this: Nowhere in
the Code is there any requirement that botanists should la-
boriously investigate encyclopedias, books of travel, text-
books of horticulture, and similar works and attempt to ap-
ply the names which they may find therein. It does not re-

1947 Gleason, Well Known Binomials 203

quire that they find, investigate, typify, and apply every
published binomial. The code does require that they use the
oldest known legitimate epithet, not the oldest one as yet
unknown. If they insist on looking up hitherto unknown
nemes, then they should be consistent «and investigate all
encyclopedias, all books of travel, all textbooks of horti-
culture, all back volumes of the Congressional Record, all
printed literature in every language, and thereby be sure
thet they have really found the oldest name.

The current code of nomenclature is intended to achieve a
definite stated purpose; it is based on certain general
principles; the use of these principles to attain the goal
is implemented by ea long series of rules.

The purpose is the establishment of a stable nomencla-
ture. The rules do not distinguish between stability of the
past and stability of the future. On the contrary, the rules
clesrly intend to maintain the stability of the past and to
project it into the future. This is evidenced by the general
principle thet no one should change names except for serious
reasons, by the use of different dates of departure, by the
abolition of tautonyms, by the adoption of nomina conservan-
da, and (what may seem strange to some botanists) by the
homonym rule, which often permits the segregation of a genus
without the publication of ea new generic name.

A careful study of the opening clauses of the Code will

convince any impartial reader that the Code is intended to

effect stability just as far as possible by maintenance of
names and just as little as possible by change of names. The
definite rules which follow and which constitute the bulk of
the Code should therefore be used to justify maintenance.
Only when maintenance is impossible should they be used to

determine the nature of the necessary change.

Those who frequently turn to the pages of the Code for
guidance and others who follow the current literature of
taxonomy are fully aware that there are clauses of dubious
application among the rules, rules which ectually or seem-
ingly conflict, nomenclatural problems connected with typi-
fication and hybridization which are not fully met. In all
such cases, the rules should be interpreted to favor the
maintenance of & name rather than its change. There are no-
menclatural problems the settlement of which seems to depend
On mere quibbling. I should not hesitate to quibble about
the interpretation of a rule if by so doing I can preserve a
well known name; I should quibble in the opposite direction
with equal readiness if I can thereby preserve another name.
If I can find any rule which will lead to the preservation

_ of a name, I shall adopt it, although another rule may be

Zé

f)
;
os

1

found which would necessitate a replacement.
In general, if botanists will search as assiduously for

204 rua: G5 oo to Vol. 2, now 7

reasons to maintain a name as they do for reasons to change
one, a considerable number of well known names will be saved.

I now present five instances of well known plants with
names long established in the literature of botany, forest-
ry, or horticulture which have come under recent criticism.
In each case strange names or new combinations have been
suggested for them. While I doubt that any change of name
can "throw science into confusion," (International Code,
Art. 3, paragraph 1) these plants are so common or so impor-
tant that any change in their names should be avoided.

Parthenocissus vitacea.

It is only a half-century since the existence of two spe-
cies of Virginia Creeper in our flora was noted. Apparently
lazenby was the first American botanist who in 1888 and 1890
called attemtion to the two forms, while Knerr gave the sec-
ond one a varietal name in 1893. In doing so he emphasized
the lack of adhesive disks on the tendrils and mentioned a
few other agubordinate features. A year later Hitchcock ele-
vated Knerr's name to specific rank as Pearthenocissus vita-
cea, under which name it has frequently appeared in American
literature.

Recently Rehder has found another name, Vitis inserta
Kerner, six years older than Knerr's variety and seven years
older than Hitchcock's species. He accordingly transferred
it to Parthenocissus and the plant appears as P. inserta in
such widely used works as Rehder's Manual of Cultivated
Trees and Shrubs (1940) and Deam's Flora of Indiana (1940);
Fernald accepted it in Rnodora (43: 604. 1941), where he
misspelled it as incerta.

Now let us examine Kerner's original publication. It con-
sists of a figure and a bit of description. The figure
shows what might be a bit of rock or a piece of bark, prob-
ably the latter, with two stems running vertically across
it. Each stem has a palmateiy compound, 5-foliolate leaf,
one of them with a tendril opposite it; the petiole of a
third leaf is shown, also opposite a tendril. Anyone will
recognize it as a Virginia Creeper and Kerner verifies this
by referring to the plant as Vitis (Ampelopsis) inserta.
Bach tendril branches with four apices; each apex has found
& crevice in the bark and has there enlarged into an adhes-
ive disk. The drawing does not show the inflorescence,
which,is the best diagnostic character of the species, nor
can it well show whether the leef is dull or glossy. Kern-
er's description is not that of a taxonomist nor is there
any evidence that he wished or intended to describe a spe-
cies or propose a name, although this fact is in itself no
reason for neglecting his name. He is writing about the be-

1947 Gleason, Well Known Binomials 205

hevior of tendrils and nothing else. Rather than quote the
original German, I append an excellent trenslation by Oliver
(Kerner & Oliver, Netural History of Plents 1: 7Cl.):

"Bignonia capreoleta, end Vitis (Ampelopsis) inserta
(whose tendrils are represented in fig. l behave differ-
ently from the three tendril-plants just mentioned. Here the
curved tips of the tendrils, growing towerds the wall, seek
the crevices and crannies of stone or bark and actually
creep into them, or when only shallow grooves are to be
found in the substratum, bury themselves in them.***When es-
tablished in the chinks and crevices, the ends, which until
now have been hooked, swell out like a club or ball, and in
a short time thicken so much that they occupy the entire
crack."

This is all the descriptive matter; the remainder of the
paregreph deals in more detail with the adhesive properties
of the tip of the tendril.

Kerner's descrirtion is not that of e taxonomist nor is
there any evidence that he wisned or intended to describe a
species or propose & name, although this fact is in itself
no reason under the Code for neglecting his name. He is
writing about the behavior of tendrils and nothing else. The
only structural feature of the plant to which reference is
made is the tendrils end special emphasis is placed on the
production of termine] hold-fasts.

That is precisely the feature which is used by modern
botenists, including Rehder, to characterize FP. ouinquefol-
ia! F. vitaces is the plant almoet always without hold-
fasts, and yet Sehder wents to displace that well known neme
by the one of Kerner. Vitis inserta, inadequately and acci-
dentally although effectually published, is merely a synonym

of Pp quinouefolia.

Nelumbo lutea.

The American lotus has regularly been known by this name
since 1805, while the specific epithet for it dates beck to
1799. Recently Fernald has drawn attention to Nymphaea pen-
tapetala Walt., published in 1788, end has advoceted the new
name Nelumbo pentepetala (Walt.) Fern.

Fernald has stated the facts correctly. Walter thought he
had two species of Lotus. One of them he misidentified with
the Old World species under the name Nymphaea Nelumbo. The
other he regarded as undescribed; he gave it the specific
neme pentapetala and a brief description: "foliis peltatis
undique integris, celyce pentaphyllo, corolla magna pente-
petala alba, loculis pericarpii monospermis." Now the spe-
cies of Yelumbo heve numerous petals, not five. Walter's
plent was either a monstrosity or en aged flower from which

‘4

206 Pow Y/T-0°TO Gtk Vol. 2, now 7

the other petals had fallen. Fernald expressed regret at
changing a well known name, but based his regret solely on
the inappropriateness of the epithet. In this he was probab-
ly guided by Recommendation XIII: "The specific epithet
should *** give some indication of *** the characters *** of
the species," rather than by Article 15: "The purpose of
giving a name to a taxonomic group is not to indicate the
characters or history of the group, but to supply e means of
referring to it." He could have avoided all regret if he had
relied on Article 65: "A name or epithet of « taxonomic
group must be rejected when it is based on a monstrosity." I
regard Walter's name as covered by this rule end reject it
accordingly.

Acer saccharum.

It is generally accepted as a fundamental principle of
good nomenclature that the publication of a misprint does
not produce a legal plent-name. There are all sorts of mis-
prints which one may note in botenical works. Most of them
are obvious, but there are some supposed cases which have
been interpreted in two ways, as a misprint and as an inten-
tional act.

Recently the botanical public has been asked to substi-
tute Acer saccharophorum for A. saccherum 4s the name of our
familiar northern Sugar Maple. The circumstances have 4l-
ready been treated in great detail and exactness by Rousseau
(Contr. Inst. Bot. Univ. Montreal 35: 1--66. 1940.). He,
however, wished to prove his own opinion and naturally pre-
sented all the evidence which he could develop in favor of
it, while excusably slighting evidence to the contrary.
Since his work may not be easily available to some readers,
@ very brief statement of the pertinent fects may be in or-
der. Some of these facts are taken directly from Rousseau;
others from the same literature from which Rousseau drew his
evidence. No edditional facts are necessary for proper ap~
preisal. .

Feter Kealm, in his travels in America, soon learned to
know the Sugar Maple and collected specimens of it. Two of
these are still extant. One, which came into the possession
of Queen Louisa Ulrika, is Sugar Maple. A second, unfortun-
ately sent to Linnaeus, is Silver Maple.

Linnaeus described four species of American maples in
1753, A. saccharinum, A. rubrum, A. pensylvanicun, and A.
Negundo. Knowing whet Kalm had learned about the Sugar .
Maple, and unawere of the confusion of the actual specimens,
he supposed that Kalm's specimen represented that tree. He
eccordingly named it the “sugary maple”, or A. saccharinum,
end gave it one of his usual brief diagnoses. The desorip-

Ee

1947 Gleason, Well Known Binomials 207

tion fits the specimen precisely, and both plant and des-
cription leave no room for doubt that the name A. saccharin-_
um belongs to our Silver Maple. Succeeding botenists gener-
ally supposed, as Linnaeus had, that the neme epplied to the
Suger Maple and it was commonly used for that tree until
1889.

The next name given the Sugar Maple was Acer sacchatum by
Fhilip “iller. It is generally supposed that this is a genu-
ine misprint. Since there is no controversy, it needs no
discussion here. The third name was Acer saccharum Mershall;
‘6 few others were given later, but since they are pure syno-
nyms they also need no discussion.

In 1889 Britton and Sargent called attention to the mis-
application of A. saccharinum™ They proceeded to use that
name for the Silver Maple and*have been followed by almost
all botanists since. For the Sugar Maple Britton brought up
Marshall's neme, A. saccharum, which was soon generally 4s-
dopted and has been in common use by botanists and foresters
for more than half a century.

Tnree decades more pass by and Mackenzie, always alert
for an opportunity to meke trouble in nomenclature, reported
thet saccherum, as originally used by Marshall, was merely a
misprint for saccharinum. This drew mild protests from Sud-
worth and Sprague, and in general botanists continued to use
seccharum, even down to the last edition of Rehder's Trees
and Shrubs. Rehder is not particularly averse to a chenge of
neme; neither is Fernald, who also continued to use sacchar-
um, although recently he has added saccharophorum in etpicae
theses. Apparently neither was convinced by Mackenzie's ar-
gument. Rousseau, examining all pertinent literature and re-
porting it in meticulous detail, is convinced thet saccherum
is a misprint.

The only valid evidence must be taken directly from Mar-
shall's Arbustum Americanum in which the name appeared. Let
us put ourselves in Marshall's position, turning the calen-
dar back 160 years. Encouraged by Bartram, we begin to write
an account of the trees known to us in America. We have a
good field knowledge of meny of them. When we come to the
maples, we note with astonishment that we have six different
species, while the great Linnaeus himself had only four.
Well, we shall do the best we can with them. Here is one de-
scribed iby Linnaeus as Acer foliis compositis, floribus ra-
cemosis. This seems to fit our Box Elder, which is the only
maple we have with compound leaves, and we write its name in
our book as Acer Negundo. Correct.

Next we take up two of our plants, both small trees with
flowers in racemes, and now we are baffled. Among his four
species Linnaeus has only one which will fit, “Acer foliis
trilobis acuminatis serrulatis, floribus racemosis. The des-

208 FB PxF (Ob OG Ek Vol. 2, noe 7

cription fits both of ours equally. We weigh every word of
it and after due deliberation we finally apply the name to
-- to the wrong species. Our A. spicatum of modern times ap-
pears as A. pensylvenicum L., while to the true A. pensyl-
vanicum is given a new name, A. canadense.

Now we have two Linnean names left and three species
still before us. A. rubrum is characterized by Linnaeus with
foliis quinquelobis subdentetis subtus glaucis, pedunoulis
simplicissimis aggregatis. Two of ours, the Red Maple and
the Silver Maple, have a crowded inflorescence and leaves
paler beneath. Again we consider the question carefully,
note that A. rubrum has leaves "quinquelobis", and with some
hesitation use that name for the Red Meple. This time we are
correct, but we are not fully satisfied, for in our later
description of the Silver Maple we hedge by writing "This is
perhaps the Acer rubrum of Linnaeus."

There are still two species to be named and only one name
available, A. saccharinum. This name seems to apply, by its
meaning, to the Sugar Maple: did not Kalm tell us how sugar
wags made from it? But Linnaeus said the leaves were ‘quinque-
partito-palmatis acuminato-dentatis, and notning more, while
the leaves of our tree would be described as “quinque-lobatis.
Our Silver Maple has five-parted leaves, to be sure, but no
one in Pennsylvania makes sugar from it. Besides its leaves
are whitened beneath; why did not Linnaeus mention such a
conspicuous cnaracter. The whitened surface leads us to
doubt whether our Silver Maple may not be the Acer rubrun,
but we have already decided to use that name for our Red
Maple.

There is only one obvious solution, that we have two un-
named species. We proceed to describe our Silver Maple as A.
gleucum, appropriately referring to the color of the leaves.
Our Sugar Maple, with merely lobed leaves, is not the one
which Kalm knew and Linnaeus described. Ours is a second
species of Sugar Maple, and we name it by translating its
local name directly into Latin, Acer saccherum.

That is no misprint. It is only an honest attempt by
Humphrey Marshall to identify his plants according to the
brief available descriptions written by a foreign botanist.
It was an attempt correct in only two instances; an attempt
which resulted in a misidentification for A. pensylvanicum,
an attempt in which he failed to recognize in his own mater-
ial any plants which corresponded to A. saccharinum L. and
failed to find in literature any names which he felt he
could properly use for the Silver Maple, the Sugar Maple,
and the Moosewood. The total result was three supposedly
new species.

Rousseau adduces one other faot as alleged proof of a
misprint. Marshall's book was translated into French a few

1947 Gleason, Well Known Binomials 209

years later by Lezermes and in the translation we find A.
saccharum replaced by A. saccharinum. Rousseau believes this
change was the correction of a misprint. We can more easily
infer that the translator believed there was only one Sugar
Maple in America and that it was an error in botanical
judgement which required correction, not a misprint. Marsh-
all gracefully bowed to European opinion and permitted the
change. Such a correction, of course, can not void the val-
idity of an, earlier name.

Finally Rousseau states that A. saccharum, if not a mis-
print, becomes a nomen nudum, since there is no accurate
means of deciding whether Marehall described the Sugar Maple
or the Black Maple. The last clause of this sentence is un-
doubtedly true; the conclusion which he drew from it is er-
roneous. Rousseau implies by his statement that the name ap-
plies to one or the other of these maples. It might also ap-
ply to both, since both live in eastern Pennsylvania. If it
applies to the Sugar Maple, it becomes the valid name for
that species. If it applies to both species, it "must be re-
tained for one of them, or (if it has not been retained )
must be re-established" [Article 52]. Britton in 1889 con-
sidered that the name belonged to both species, and by nam-
ing the Black Maple A. saccharum var. nigrum he indicated
that the typical nomenclatural element of the name applied
only to the Sugar Maple. If the name applies only to the
Black Maple, it has priority over and displaces A. nigrum
Michx. (1803) but, since its application has been fixed by
Britton's action and perpetuated by many years of usage, the
burden of proof is upon those who might wish so to restrict
it. Such proof has never been presented and probably can
never be.

I therefore retain Acer saccharum as the valid name for
the Sugar Maple.

Lathyrus maritimus vs. Lathyrus japonicus.

When Fernald discussed these names in 1932, he professed
to regret that the International Code compelled the dis-
placement 6f such e well known name as Lathyrus maritimus
for such a well known plant as the Beach Fea.

The facts of the matter are simple and were well stated
by Fernald. The Beach Fea lives on both Atlantic and Facific
shores of Burasia and North America and also inland in suit-
able habitats. In epite of this broad distribution, it is
regularly regarded as a single species. It was described
from Europe by Linnaeus in 1753 as Pisum maritimum. It was
described from Japan by Willdenow in 1803 as Lathyrus japon-
icus. It was described from Massachusetts by Bigelow in 1824
as Lathyrus maritimus. It was described from Scandinavia by

210 PRY OLO 8 Ts Vol. 2, noe 7

Fries in 1834 as Lathyrus maritimus. It has received other
specific or subspecific epithets, none of which have any
bearing on the present problem.

The earliest specific epithet is of course maritimus; the
next is japonicus. Now here is the crux of the question. If
Bigelow transferred the Linnean name from Pisum to Lathyrus
in 1824 he then created a new and valid binomial, Lathyrus
maritimus (L.) Bigel., which must stand as the name of the
species. On the contrary, if Bigelow described a new species,
then the transfer of the Linnean epithet to Lathyrus by
Fries in 1834 merely created a homonym which is invalid un-
der the International Code. Being invalid, the next oldest
specific epithet must be used, which is japonicus.

Did Bigelow transfer an epithet, or did he describe a new
species? Fernald, apparently looking for a reason to change
@ name, says @ new species was described.

The essential purpose of the International Code is stated
in Article 4. It is to strive for fixity in nomenclature.
This purpose is implemented by the long series of rules and
recommendations which constitutes the bulk of the code. If
we are to strive for fixity of names, we must search the
rules for clauses which will permit us to maintain a well
known name. Fernald found clauses which permitted him to
change a name. Are there other clauses which will authorize
us to maintain the name? If Bigelow made a transfer, the
name will automatically be maintained. Did he make such a
transfer?

Some evidence on this point may be discovered by examin-
ing Bigelow's treatment of other species.

There are thirty species in his Florula Bostoniensis
which are treated differently from the others, in that the
usual diagnosis in English is preceded by a diagnosis in
Latin. Of these thirty, twenty-three include no statement of
synonyms of any kind, and are each preceded by an asterisk.
Each of them represents the first publication of a new bi-
nomial (in one inetance a trinomial) to designate what Bige-
low believed to be a new species (in one instance a varie-
ty). Not all of them stand today, most of them having been
previously described without Bigelow's knowledge or being .
otherwise untenable. The point is, that in describing e "new
species, he preceded the name by an asterisk and gave a Lat-
in diagnosis. Five of the thirty are preceded by an aster-
isk, have a Latin diagnosis, but include some mention of
synonyms. Bunias edentula is merely continued from its orig-
inal publication in the first edition; the synonym, Cakile
americana Nutt., is later than Bigelow's name. Galium Tor- Tor-

reyi is new here as a species; its synonym is a variety, —
here raised to specific rank. Prunus obovata is a new spe-
cies here, the synonym merely indicates that Pursh had con-

1947 Gleason, Well Known Binomials 211

fused it with P. serotina. Prunus littoralis is also new;
its synonym indicates that Micheux had confused it with P.
sphaerocarpa. The fifth, Actaea alba, is followed by an ex-
planatory note: "First published as a distinct species, in
my name, in Eaton's Manual of Botany, afterward by Mr. Elli-
ott under another name." The synonyms include Elliott's name
and two varietal names under which the plant was treated by
Michaux and Pursh. Considering these five with the preceding
twenty-three, we are at once led to the conclusion that ev-
ery species or specific name for which Bigelow was respons-
ible was so designated by an asterisk

There are still two left over which heave a Latin diagno-
sis but no asterisk. The first of these is Iris prismatica
Pursh, a plant "first described by me in the former edition
of this work under the name of I. gracilis. Two years after-
wards Mr. Fursh gave it the name of I. prismatica, which
name I am willing to adopt." The other is Lathyrus palus-
tris, under which he cites "Syn. Pisum maritimum. Pursh?"

In both cases the absence of an asterisk indicates a species
for which Bigelow is not responsible.

We can easily interpret Pisum maritimum as the basinyn,
and we shall do so if we are seriously interested in the
spirit of the International Code. It was not necessary to
cite the original author of the name (Linnaeus); there was
no other Pisum maritimum with which it could be confused.
Citation of authors is for "purposes of precision" [Code,
Sect. 7] and “in order that the date may be readily verifi-
ed" [Article 46]. Article 44 states that "the name of a spe-
cies *** is not validly published unless it is accompanied
*** by the citation of a previously and effectively publish-
ed description *** under another name.” The mention of
Pursh can be construed to cover this requirement. The Code
does not specifically require the mention of volume and
page.

The case is closely parallel to that of Hedysarum glutin-
osum Willd. (1802) and Desmodium glutinosum Wood (1845).
Both names apply to the same species. If Wood's name is a
transfer of Willdenow's oldest specific epithet, it becomes
the valid binomial for the species. If on the other hand it
is a description of a new species, its existence invali-
dates the later transfer of Willdenow's name to Desmodium by
Schindler (1926) and necessitates the revival of the next
oldest specific epithet, acuminatum Michx. (1803), in the
well known binomial Desmodium acuminetum (Michx.) DC. Miss
Schubert [Rhodore 44: 279] says: "Although it is true that
Wood cited neither authority nor synonyms his description
leaves no doubt as to his intention nor as to the identity
of the plant he was considering." Here she has done precise-
ly whet Fernald refused to do for the Beach Pea and done it

212 Fier.) Ot Oe 2s Vol. 2, now 7

probably with Fernald's knowledge and possibly with his ap-
proval. The adoption of opposite opinions for the two plants
has permitted them to recommend the abandonment of two well
known namese

And Fernald himself has done the same thing. In Rhodora
44: 424 he takes up the name Rhynchosia difformis (E£ll.) DC.
He says “Although DeCandolle failed to cite the synonym Ar-
cyphyllum difforme Ell., the diagnosis *** and the habitat
*** are so clearly derived from Elliott that the combination
should certainly be written Rhynchosie difformis (Ell.) DC."

In each of these three cases we admit the conspecificity
of the pkants involved and we know the source of the specif-
ic epithet used in the combination. Bigelow is the only one
who cites the name-bringing synonym; Bigelow also shows by
his typography that he did not regard his neme as designat-
ing @ new species, a change of name, or a replacement of an
untenable name. How else do valid names arise except by
transfer?

Following the spirit and intent of the Code, taking ad-
vantage of loopholes in Article 44, and imitating the prec-
edent of Schubert and Fernald, I shall maintain the well
known and long established name Lathyrus maritimus (L.)
Bigel. for the Beach Pea.

A NEW SPECIES OF DAPHNOPSIS FRO¥ ECUADOR

Joseph V. Monachino

DAPHNOPSIS ESPINOSAE Monechino, sp. nov.

Arbuscula; foliis ellipticis ca. 4--8 em. longis et 1.5--—
3 cm. latis glaberrimis; petiolis 3--4 mm. longis, 1.5 mn.
latis; inflorescentiis caulifloris 1.5--2 cm. longis; flori-
bus femineis 6--12 subumbellato-racemosis; calyce campanula-
to, ca. 2.5 mme longo, extus parce pubescente, lobis rotun-
datis ca. 1.5 mm. longis paullo letioribus intus pubescenti-
bus; staminodiis et petalorum rudimentis nullis; ovario gla-
bro; stylo 0.8 mm. longo; gtigmate capitato exserto; disco
crateriformi irregulariter lobato glabro.

Vegetative parts completely glabrous except for the cili-
ate bud-scales; petioles about 3 or 4 mm. long and 1.5 mm.
broad; leaf-blades glabrous on both surfaces from the begin-
ning, becoming chartaceous or subcoriaceous and shining a-
bove, elliptic, narrowed at both ends, obtuse or acute at a-
pex, 4--8 om. long and 1.5--3 cm. broad, the reticulation
prominulous; inflorescences cauliflorous, 1.5--2 cm. long,

ote

1947 Monachino, New Species of Daphnopsis 213

sparsely hispidulous; only female flowers seen, 6--12 in um=-
belloid racemes at the ends of short (6--13 mm. long) simple
peduncles; pedicels up to about 1.5 mm. long, articulate
near the apex; calyx campanulate, about 2.5 mm. long, glab-
rescent or sparsely pubescent outside, glabrous inside, the
calyx-lobes reflexed, rounded, about 1.5 mm. long and
slightly broader, pubescent on the inner surface and with a4
tuft of hairs at the apex; staminodes and rudimentary petals
none; ovary glebrous, about 1.5 mm. long; style 0.8 mm.
long; stigma capitate and densely papillose, exserted from
the calyx; disk conspicuous, crateriform, oblique, irregu-
larly lobed, glabrous.

Type: Reinaldo Espinosa 205, collected at Namanola, alt.
2400--2500 m., southern Loja, Ecuador, April 18, 1946, de-
posited in the Britton Herbarium at the New York Botanical
Garden. The type specimen consists of young leaves and
flowers. The following matured flowering specimen has also
been examined: Reinaldo Espinosa sen. [Herb. Krukoff 19848]
from the type locality, received in February, 1947.

Daphnopsis Espinosae has affinity with D. zamorensis
Domke, the type of which was collected at Zamora, Loja. D.
zamorensis, however, is described as having leaves about 186
to 27 om. long and 5.5 to 8 cme broad, petioles 1 to 1.5 cm.
long and 0.35 to 0.4 cm. broad, and inflorescences & om.
long. The much smaller leaf and inflorescence size of D.
Espinosae is an obvious means of distinguishing it from D.
zamorensis. From other species found in Ecuador and Feru --
D. loranthifolia, D. caribaea var. ecuadoriensis, D. carib-
aea Vare peruviensis, De Weberbaueri, and De Pavonii -- the
present species is easily distinguished by “ite entirely
glabrous leaves and by other characters.

-— sr 2 2s ese @F =e =f sF Ff 2s ee =e =e se = = - = =

NOTES ON NEW AND NOTEWORTHY PLANTS. I

Harold Ne Moldenke

Tne present paper is the first in a series of notes on
plants of various parte of the world, based in part on
field studies and in part on herbarium studies in the her-
barium of the New York Botanicel Garden and elsewhere.
Numerous new species, varieties, forms, and hybrids will
be described and several new names and combinations pro-
posed. Abbreviations used herein for the names of herbaria
in which cited specimens are deposited are in conformity
with my previous publications, but for the convenience of

214 PHYTOLOGIA Vol. 2, .m@elt

the readers of the present paper the ones herein used are as
follows: Al = New York State Museum, Albany; Be = Bailey
Hortorium, Ithaca; Be = Barnard College Herbarium, New York
Botanical Garden, New York City; Bt = Butler University,
Indianapolis; Bu = Buswell Herbarium, University of Miami,
Coral Gables; C = Columbia University Herbarium, New York
Botanical Garden, New York City; Cm = Carnegie Museum,
Fittsburgh; Dm = C. C. Deam Herbarium, Bluffton, Indiana;

Dp = DePauw University, Greencastle; Du = Dudley Herbariun,
Stanford University, California; Fe = Colorado Agricultural
& Mechanical College, Fort Collins; Fl = University of
Florida, Gainesville; Go = Botaniska Tradgard, Goteborg,
Sweden; H #» Duke University, Durham; Hp = H. Hapeman Herbar-
ium, Minden, Nebraska; Hs = Crispus Attucks High School,
Indianapolis; I = Langlois Herbarium, Catholic University of
America, Washington; Io = Iowa State College, Ames; It =
Cornell. University, Ithaca; Mi = University of Michigan, Ann
Arbor; N = Britton Herbarium, New York Botanical Garden, New
York City; Fl = State College of Washington, Pullman; Fo =
Pomona College, Claremont, California; St = Oklahoma Agri-
cultural & Mechanical College, Stillwater; T = Torrey Herb-
arium, New York Botanical Gerden, New York City; Ua = Utah
State Agricultural College, Logan; Up = University of Penn-
sylvania, Philadelphia; Ur = University of Illinois, Urbana;
Vt = University of Vermont, Burlington; and We = West Vir-
ginia University, Morgantown.

ABGIPHILA HOZHNEI var. PUYENSIS Moldenke, var. nov.

Haec varietas a forma typica speciei pilis ramorum ramu-
lorumque atrobrunneis rigide patentibus et pilis foliorum
non bulbosis recedit.

This variety differs from the typical form of the species
in the pubescence on its branches and branchlets being dark.
brownish and stiffly wide-spreading and that of the upper
leaf-surfaces not being bulbous-based.

The variety is described as a woody vine about 3 m. long,
with off-white flowers, and war collected by We Ce Steere
and W. H. Camp (no. 8283) at an altitude of 3000 feet in the
vicinity of Puyo, Parroquia Puyyv, Oriente, Ecuador, on May
12, 1944, and is deposited in the herbarium of the Chicago
Museum of Natural History.

ANASTRAPHIA RECURVA var. INTZGRIFOLIA Moldenke, var. nov.

Haec varistas a forma typica speciei foliis integris re-
cedit.

This variety differs from the typical form of the species
in having all its leaves entire-margined.

The type was collected by Julia Acufia Galé (no. 12780) at
Rfo Yagrumajes, Moa, Oriente, Cuba, on April 14, 1945, and

1947 Moldenke, New and Noteworthy Plants 215

is deposited in the herbarium of the Estacion Experimental
Agronomica at Santiago de las Vegas, Havana, Cuba.

CALPIDISCA LUNDII (A. DC.) Moldenke, comb. nov.
Utricularia Lindii A. DC., Prodr. 8: 14. 1844,

CALYPTRANTHES CAROLI var. LONGIPEDUNCULATA Moldenke, var.
nov.

Haec varietas a forma typica speciei pedunculis 1.5--3
cm. longis ebracteatis recedit.

This variety differs from the typical form of the species
in having peduncles 1.5 to 3 cm. long, without any leaf-like
bracts at its apex.

The type was collected by Brother Leon and Juan T. Roig
(no. 13544) at Loma Pelada, Cayajabos, Pinar del Rfo, Cuba,
on August 10, 1928, and is deposited in the Britton Herbar-
jum at the New York Botanical Garden.

CISSAMPSLOS LAXIFLORA Moldenke, sp. nov.

Fruticulus scandens; ramis gracilibus sulcato-striatis
adpresso-pilosis; laminis foliorum leviter chartaceis in
siccitate atrobrunneis vel nigrescentibus supra nitidis
non peltatis late ovatis acuminatis mucronatis, ad basin
truncatis vel subtruncatis, integris supra glabris subtus
minute adpresso-pilosulis.

Small vine; stems twining, slender, longitudinally sul-
cate-striate with many narrow striae, more or less appress-
ed-pilose; principal internodes 3.5--7 cm. long; leaf-blades
thin-chartaceous, dark-brown or nigrescent and shiny above
in drying, lighter beneath, not peltate, broadly ovate, 5--
8.5 cm. long, 3.5--7.5 cm. wide, acuminate at the apex, the
acumination attenuate into a mucro about 3 mm. long, trun-
cate or subtruncate at the base, entire, sometimes irregu-
larly angulate at the widest part, glabrous above, minutely
appressed-pilosulous beneath; principal veins 5, issuing
from the very base of the blade, along with the secondaries
and tertiaries slightly prominulous on both surfaces; stam-
inate inflorescence axillary, 1 or 2 per axil, pedunculeate,
paniculate-racemose; peduncles straight, slender, 1--1.5 cm.
long; rachis straight, erect, slender, 5--10 cm. long, ap-
pressed-pilose; branches filiform, usually less than 1 cm.
long, spreading, pilose; bracts absent or very minute; pist-
illate flowers: sepal 1, thick, ovate-elliptic, about 1.6
mm. long and equally wide if pressed flat, very convex on
the outer and concave on the inner surface, enfolding the
remainder of the flower, rounded at apex and base, glabrous
on both surfaces; petal 1, on the same side and in front of
the sepal, broadly obovate-orbicular, lighter textured than
the sepal and lighter in color, about 1 mm. long and wide,

216 PHYTOLOGIA Vol. 2, noe 7

rounded at apex and base, glabrous on both surfaces; pistil
1, about 1.2 mm. long, glabrate; style obsolete; stigmas 3,
about O.3 mm. long, spreading, acute; staminate flowers: se-
pale 4, membranous, elliptic, about 0.6 mm. long and O.4 mm.
wide, subacute at apex, narrowed at base, glabrous, very
fragile; petals 4, connate; stamens 4, connate; pistillate
inflorescence axillary, racemose, simple or paniculately
branched, one per axil; bracts foliaceous, orbicular-ovate,
5--15 mm. long, 3--11 mm. wide, long-mucronate at apex,
long-stalked at base; rachis slender, 12--18 om. long, bear-
ing the flowers in fascicles at intervals of 5--10 mm., the
branches (if any) few and wide-spreading; fruiting racemes
with 1--4 fruits in a cluster, their pedicels 10--15 mm.
long, the clusters 1--1.5 cm. apart, the rachis very slen-
der, the bracts persistent but only one subtending each
cluster of fruit and therefore widely separated; fruit pyri-
form, about 6 mm. long and 4 mm. wide, minutely pilosulous
or glabrate, nigrescent in drying, striate with several con-
centric tuberculate ridges.

The type of this species was collected by J. Murga Pires
and Ge Ae Black (no. 949) at Tabatinga, Amazonas, Brazil, on
November 30, 1945, and is deposited in the Britton Herbarium
at the New York Botanical Garden. The type is pistillate;
another pistillate collection from the same locality is no.
1072 and staminate collections are nos. 939 and 947, all
collected by the same collectors at the same locality. Klug
2322, from Loreto, Peru, is probably the same species and is
pistillate.

XCISTUS CULTORUM Moldenke, nom. nov.
Cistus villosus L. x C. laurifolius L. ex Rehd., Man.
Cult. Trees & Shrubs, ed. 2, 646. 1940.

DESFONTAINIA PULCHRA Moldenke, sp. nov.

Suffrutex debilis; caule ramisque flexilibus griseis gle-
bris, in statu juventute subtetragonis marginatisque; nodis
distincte annulatis; internodiis abbreviatis; petiolis 1--3
mn. longis glabris compressis late marginatis; laminis cori-
aceis nitidis oblanceolatis 1--2 cm. longis, 5--8 mm. latis
acutis saepe muticis, ad basin attenuatis, subintegris vel
2-denticulatis; floribus solitariis nutantibus.

Sprawling subshrub; stems and branches flexible, gray,
glabrous, the younger parts more or less subtetragonal and
margined, the outer bark readily peeling off; nodes dis-
tinctly, annulate; principal internodes abbreviated, 1--2.5
cme long; twigs numerous, short, leafy; leaves decussate-
opposite; petioles 1--35 mm. long, glabrous, broadly margin-
ed and flattened; blades coriaceous, deep-green and very
shiny above, pale-green or silvery beneath, oblanceolate,

1947 Moldenke, New and Noteworthy Plants 217

1--2 om. long, 5--8 mm. wide, acute and often muticous at
the apex, gradually attenuate to the base, subentire or with
two very small muticous teeth near the apex, the margins us-
ually revolute; midrib very slender, deeply impressed above,
slightly prominulent beneath, the short secondaries and ter-
tiaries impressed above, practically indiscernible beneath;
inflorescence terminating the short twigs; flowers solitary,
apparently nutant; pedicels about 1.5 cm. long, dull-green,
glabrous, ‘shiny; calyx dull-green, deeply 5-fid, the lobes
oblong-elliptic, 6--6.5 mm. long, 2--2.5 mm. wide, acute,
glabrous, shiny; corolla-tube cylindric-infundibular, scar-
let-crimson outside, pale-yellow within, about 2.5 om. long,
about 4 mm. wide at the base and 10 mm. wide at the apex,
glabrous; corolla-lobes rich-yellow, ovate-lingulate, about
6 mm. long and 8 mm. wide, rounded at the apex, venose,
glabrous; fruit globose, fleshy, about 5 mm. long and wide,
glabrous.

The type of this very distinct species was collected by
Julian A. Steyermark (no. 57344), sprawling over bluffs in
rich moist woods at the base of Paramo de Tama, 4--10 km.
above Betania, 2500--2895 m. altitude, Tachira, Venezuela,
on July 15, 1944, and is deposited in the herbarium of the
Chicego Natural History Museum (sheet no. 1205340). Its
smell oblanceolate entire or minutely 3-denticulate leaves
distinguish it eat once from all other known species of this
genus.

DESFONTAINIA STEYERMARKII Moldenke, sp. nov.

Frutex; ramis ramulisque gracilibus griseis glabris mar-
ginatis; nodis annulatis; petiolis glabris paullo marginat-
is; leaminis coriaceis ovato-elliptiris vel ellipticis acutis
muticis, ad basin longe cuneato-attenuatis, glabris non
conspicue marginatis 4--6-denticulatis; calyce profundo 5-
fido, lobis ovate-lanceolatis glabris; corolla 1.5--l.7 cm.
longa.

Shrub, about 4 feet tall; branches and branchlets slend-
er, gray, glabrous, the younger parts more or less subtet-
ragonal and margined, the bark readily peeling off from old-
er parts; nodes distinctly annulate; principal internodes
1.5--5 cm. long; leaves decussate-opposite, numerous, often
with very much abbreviated several-leaved twigs in their ax-
ila; petioles slender, 3--8 mm. long, glabrous, slightly
margined; blades coriaceous, rich-green above, pale-green
beneath, not shiny, ovate-elliptic or elliptic, acute and
muticous et apex, long-cuneate-attenuate to the base, with 2
or 5 irregular muticous teeth along each margin, glabrous,
not revolute or very slightly so on the very margins; midrib
slender, plane above, prominent beneath; secondaries very
slender, 3--5 per side, mostly rather obscure above or very

218 PHYTOLOGIA Vol. 2, now 7

slightly subimpressed in drying, conspicuous and prominulent —

beneath; veinlet reticulation mostly obscure above or very
slightly subimpressed in drying, only the largest portions
subprominulous beneath; inflorescence axillary or subtermin-
al, apparently erect, solitary; pedicels slender, 1.5--1.8
mm. long, glabrous; calyx deeply 5-fid, the lobes ovate-
tanceolate, 1--1.5 mm. long, acute, glabrous; corolla-tube
cylindric, orange-red, 1.5--1.7 cm. long, 2--4 mm. wide, ab-
ruptly ampliate to 6 mm. just below the limb, glabrate; cor-
olla-lobes elliptic-lingulate, pale-yellow, about 5 mm.
long and 3 mm. wide, subacute, venose, glabrate; style ca. 2
em. long, curved at apex, glabrous; fruit elliptic or sub-
globose, apiculate, about 9 mm. long and 8 mm. wide, glab-
rouge
The type of this species was collected by Julian A. Stey-

ermark (no. 54597), in whose honor it is named, on wooded
slopes along the Rfo Valladolid, between Quebrade Honda and
Tambo Valladolid, 2000--3000 m. altitude, Santiago-Zamora,
Ecuador, on October 12, 1943, and is deposited in the herb-
arium of the Chicago Natural History Museum (sheet no.
1205653). The species is obviously closely related to De

splendens H.B.K. and De spinosa Rufz & Pav., both of which
differ in their much more heavily leathery-coriaceous leaves
with long teeth and greatly revolute margins and their flow-
ers 2.5--4 cm. in length.

ERIOCAULON CONGENSE Moldenke, sp. nov.

Herba; foliis rosulatis numerosis crassis 10--15 cm. lon-
gis glabris, ad apicem cucullatis, ad basin amplietis et
pellucido-fenestratis; vaginis laxis 10 cm. longis strietis
obscure vel non fenestratis glabris, ad apicem bilobatis,
lobis ovetis 1 cm. longis subacutis; pedunculis solitariis
15 cm. longis 8-costatis glabris non tortis; capitulis glo-
bosis albis 1 cm. diametro; floribus trimeris.

Herb; stems much abbreviated; leaves rosulate, numerous,
thick-textured, bright-green, 1C--15 cm. long, 4--8 mm.
wide at the mid-point, ampliate and pellucid-fenestrate at
base, cucullate at the spex, glabrous on both surfaces;

sheaths loose, about 1C cm. long, striate, very obscurely or

not at all fenestrate, glabrous, 2-lobed at the apex, the
lobes ovate, about 1 cm. long, subacute, glabrous, erect;
peduncles solitary, about 15 cm. long, 8-costate, glabrous,
not twisted; heads globose, white, about 1 cm. in diameter;
involucral bractlets few, stramineous, ovate, about 4 mm.
long end 2.5 mm. wide, acute, glabrous; receptacle glabrous;
receptacular bractlets stramineous, oblong, about 4 mm. long
and 1.5 mm. wide, long-acuminate at the apex, glabrous;
staminate florets: sepals 5, black except at the base, sep-
arate except at the very base, oblanceolate, conduplicate-

1947 Moldenke, New and Noteworthy Plants 219

falcate, about 3 mm. long, about 1.4 mm. wide if flattened
out, acute at the apex, white-pilose on the back toward the
apex; petals 3, connate into an infundibular, white, glab-
rous tube about 1.5 mm. long, the terminal free portions
narrow-elliptic or oblanceolate, about 2 mm. long and 0.5
mm. wide, bearded toward the apex on the inner surface, with
a narrowly elongate black gland in the center neer the apex;
stamens 6; filaments adnate to the corolla, 3 attached to
the mid-point of the free portion of the petals, the other 3
attached to the sinuses between the petals, white, glabrous,
somewhat surpassing the petals; anthers black, elliptic, a-
bout 0.4 mm. long; pistillate florets: sepale 3, bleck, sep-
arate, elliptic-falcate, conduplicate, about 4 mm. long and
2 mm. wide if flattened out, glabrous, subacute, short-
pilosulous toward the apex on the outer surface; petals 3,
firm, erect, white, narrowly oblong, separate, 2.5 mm. long,
0.5--C.7 mm. wide, obtuse, densely white-pilose on the inner
surface on the upper half, with a narrowly elongate black
gland in the center below the apex within; style 2 mm. long,
glabrous; stigmas 3, erect, 2 mm. long; ovary subglobose, 2
mm. long and wide, 3-lobed, 3-sulcate, 3-celled, 45-ovulate.

The type of this species was collected by J. P. Chapin
(no. 404) in the Kikeri meadow at the western base of Mount
Mikeno, Kivu district, altitude 7200 feet, Belgian Congo, on
June 20, 1927, and is deposited in the Britton Herbarium at
the New York Botanical Garden. The collector describes it as
a “lily-like plant with small heads of white flowers."

ERIOCAULON ROCKII Moldenke, sp. nov.

Herba aquatica submersa; foliis caespitosis leviter mem-
branaceis erectis argute attenuatis ubique glabris; vaginis
gracilibus adpressis glabris non strietis non tortis, lemine
lanceolata erecta adpressa attenuata saepe bilobata vel fis-
fa; pedunculis gracillimis stramineis tricostatis paullo
tortis glebris; capitulis nigris ellipticis vel hemispheeri-
cis; floribus trimeris.

Submerged aquatic herb; stems obsolete; leaves tufted,
thin-membranous, erect, 1--2.5 cm. long, about 1 mm. wide at
the mid-point, sharply attenuate at the apex, glabrous
throughout; sheath slender, appressed, about equaling the
leavee, about 2 cm. long, not conspicuously striate, not
twisted, glabrous throughout, obliquely split at the apex,
the blade lanceolate, about 5 mm. long, erect, appressed,
attenuate, sometimes bilobed or again split to the base; pe-
duncles very slender, stramineous, 2.5--6.5 cm. long, 3-cos-
tate, slightly twisted, glabrous; heads black, elliptic or
hemispheric, 1--4 mm. wide; involucral bractlets black,
broadly elliptic or suborbicular, about 2 mm. long and 1.5
mm. wide, rounded at the apex, very concave on the inner and

220 PES T.O0L O:.G 7A Vol. 2, noe 7

convex on the outer surface, glabrous throughout, shiny; re-
ceptacle glabrous; receptacular bractlets black, oblanceola-
te, about 1.9 mm. long and 0.6 mm. wide, acute or shortly
subacuminate at the apex, glabrous throughout; staminate
florets: sepals 3, separate almost to the base, narrowly ob-
long, falcate, 1.5--1.7 mm. long, about 0.5 mm. wide, the
upper 2/3 black, hyaline at the base, acute, glabrous
throughout or very minutely ciliolate at the very apex; pet-
als united into a subhyaline tube 1.8--2 mm. long, no free
lobes seen; stamens 6 (sometimes only 57); Pilements wide-
spreading, white, about 0.4 mm. long; anthers not seen; pis-
tillate florets: sepals 3, separate, black, elliptic, navic-
ular, about 1.7 mm. long and 0.6 mm. wide, sharply acute at
the apex, glabrous throughout; petals 4, separate, narrowly
oblong or linear, gray, about 1.5 mm. long and 0.2 mm. wide,
acute et the apex, glabrous throughout, not glanduliferous;
style slender, about C.6 mm. long, glabrous; stigmas 3, fil-
iform, erect, 0.4--0.6 mm. long; ovary subglobose, about 0.5
mm. long and wide, glabrous, J-lobed, 3-celled, j-ovulate.
The type of this little species was collected by J. Fe
Rock (no. 10843) at Saba on the eastern slopes of Likiang
Snow Range, Yangtze watershed, Ylinnan, China, in 19235 or
1924, and is deposited in the Britton Herbarium at the New
York Botanical Garden. The material dissected was old, with
seeds fully ripe in the pistillate florets. The staminate
florets, therefore, were not observed at their best.

ERIOCAULON ROBINSONII Moldenke, sp. nov.

Herba parva; foliis caespitosis levibus non fenestratis
adscendentibus graminoideis multinerviis obtusis glabris;
-vaginis cylindraceis arcte adpressis vel laxiusculis stria-
tis leviter membranaceis vel subhyalinis, ad apicem oblique
fissis, lamina lanceolata saepe bilobata erecta; pedunculis
gracilibus 4-costatis tortis glabris griseis; capitulis
hemisphericis vel conicis dense villosis; floribus trimeris.

Dwarf herb; stems extremely abbreviated or obsolete;
leaves tufted, variable in width, thin-textured, not fenes-
trate, more or less ascending, grass-like, 1--4 cm. long,
1--4 mm. wide at the mid-point, apparently the earliest
longest and broadest and these often not present any more at
time of anthesis, many-nerved, rather blunt at the apex,
glebrous; sheath cylindric, closely appressed or rather
loose, 1--1.5 cme long, striate, thin-membranous or almost
subhyaline, usually shorter than the leaves, obliquely split
at the’apex, the blade lanceolate, 3--4 mm. long, often bi-
lobed or even split to the base, appressed or rather loose,
erect; peduncles slender, 1--8 cm. long, usually 2--3 cm.
long, 4-costate, twisted, glabrous, gray; heads hemispheric
or conic, 2--5 mm. in diameter; involucral bractlets light-

1947 Moldenke, New and Noteworthy Plants 221

gray, broadly elliptic, sometimes hyaline, lightly pigmented
toward the apex, about 2.5 mm. long and 1.9 mm. wide, obtuse
at the apex, glabrous, shiny, the upper margin often more or
less erose; receptacle long-villous; receptacular bractlets
broadly obovate, hyaline, cucullate, about 2 mm. long and 1
mm. wide, rounded at the apex, glabrous throughout; stamin-
ate florets: sepals 3, separate, hyaline, elliptic-oblanceo-
late, decidedly falcate, about 1.3 mm. long and 0.4 mm. wide
blunt at the apex, glabrous throughout; petals 3, united in-
to a slender tube about 1.7 mm. long, the free lobes very
short, about 0.2 mm. long, mucronate; stamens 6; anthers
brown; pistillate Plorets: sepals 5, separate, narrowly ob-
long-lanceolate, gray, about 1.5 mm. long and 0.2 mm. wide,
minutely bifid at the apex, long-pilose on the back; petals
3, separate, hyaline, nerrowly oblong or linear, about 1 mm.
long, long-pilose on the back, with a very narrow black
gland on the back near the apex, not bearded; style fili-
form, about 0.5 mm. long, glebrous; stigmas 3, erect, fili-
form, about 1 mm. long; ovary subglobose, about 0.4 mm. long
and wide, 3-lobed, J-sulcate, glabrous, J-celled, 3-ovulate.

The type of this species was collected by Charles Budd
Robinson (no. 1043) at Nha-trang and vicinity, Annam, French
Indo-china, between March 11 and 26, 1911, and is deposited
in the Britton Herbarium at the New York Botanical Garden.
In habit and general appearance this species greatly resem-
bles E. achiton Kérn. of India, but that differs in having
only 2 sepals in the staminate and pistillate florets and no
petals in the pistillate florets. It is a pleasure to name
this species in honor of the distinguished, though ill-
starred, collector to whom we owe so much of our knowledge
of the flora of the Philippines and other southeastern Asia-
tic areas.

ERIOCAULON YUNNANENSE Moldenke, sp. nov.

Herba; caule valde abbreviato; foliis paucis erectis
firmis graminoideis multistriatis supra villosis subtus pil-
osulis glabrescentibus subuletic plerumque conduplicatis non
fenestratis; vaginis cylindricis adpressis multistriatis
glabris, ad apicem fissis, lamina lanceolata erecta adpressa
attenuato-subulata glabra; pedunculis 1 vel 2 crassiusculis
S-costatis argute angulatis glabris; capitulis hemisphaeri-
cis albis; floribus trimeris.

Herb; stem greatly abbreviated, about 1 cm. long or less;
leaves few, basal, erect, firm-textured, grass-like, 55--60
cm. long, about 1 cm. wide at the mid-point, many-striate,
more or less villous on the upper surface and pilosulous on
the lower surface toward the base and when young, glabres-
cent in age, subulate-tipped, often more or less condupli-
cate, not plainly fenestrate; sheath cylindric, appressed to

222 PRY TOL 061s Vol. 2, now 7

the peduncles, shorter than the leaves, 25--41 om. long,
many-striate, hardly twisted, glabrous, obliquely split at
the apex, the blade lanceolate, erect, appressed, 3--6 cm.
long, attenuate-subulate at the apex, glabrous; peduncles 1
or 2 per plant, relatively stout, about S-costate with very
prominent and sharp angles, glabrous (or microscopically
puberulous within the sulcae); heads hemispheric, white, 10
--12 mm. in diameter; involucral bractlets broadly obovate
or suborbicular, about 3 mm. long and 2.1 mm. wide, rounded
or obtuse at the apex, usually membranous-margined at the a-
pex and toward the apex and often splitting there, brownish
toward the apex and in a median band to the base, villosu-
lous on the back; receptacle very densely long-villous even
between the involucral bractlets; receptacular bractlets
broadly obovate, about 3.4 mm. long and 2.1 mm. wide, dark-
brown toward the apex, lighter brown or stramineous toward
the base, abruptly acuminete-mucronate at the apex, densely
white-pubescent on the back from the widest part to the apex
and including the mucro with short antrorse hairs, not oth-
erwise bearded, glabrous toward the base; staminate florets
short-pedicellate: sepals 3, separate, navicular, broadly
obovate, cannot be flattened out, dark-brown except at the
base, about 2.9 mme long, each half about O.7 mme wide, ab-
ruptly short-acuminate or mucronate at the apex, densely
short-pubescent at the apex with white antrorse hairs, hard-
ly distinctly bearded; petals connate into a slender stram-
ineous tube about 1.7 mm. long, glabrous, the free lobes
lanceolate, about 0.4 mm. long, sharply attenuate-acute,
black-glanduliferous near the apex, pilose; stamens 6; pis-
tillate florets short-pedicellate: sepals 4, separate, nav-
icular, cannot be flattened out, elliptic, dark-brown on the
upper half, lighter at the apex and base, about 3.2 mm.
long, each half about 1 mm. wide, blunt or subacute at the
apex, short-pubescent at the apex with white antrorse hairs,
hardly distinctly bearded, otherwise glabrous; petals 3,
separate, subhyaline, linear-oblong, about 3 mm. long and
O.2 mm. wide, subacute at the apex, densely long-villous on
and near the margins at about the middle with hairs that al-
most reach the top of the petal, bearded at the apex, gland-
uliferous on the back just below the apex; style about 0.6
mm. long, glabrous; stigmas 3, erect, about 0.8 mm. long; o-
vary elliptic, about 1 mm. long, deeply 3-lobed and 3-
sulcate, glabrous, J-ovulate.

The type of this large and distinct species was collected
by Ae Henry (no. 12362) at Szemo, Ylinnan, China, and is de-
posited in the Britton Herbarium at the New York Botanical
Garden. In habit it reminds one of E. decangulare L. of the
southeastern United States.

1947 Moldenke, New and Noteworthy Plants 223

HELIETTA CUBENSIS Monachino & Moldenke, sp. nov.

Frutex (?); ramis gracilibus glabris suberosis; foliis 3-
foliolatis; petiolis gracilibus glebris pellucido-punctatis;
foliolis sessilibus leviter coriaceis oblanceolatis glabris
utrinque conspicue pellucido-punctatis, subtus glaucescenti-
bus, supra nitidis, ad apicem rotundatis, ad basin longe at-
tenuatis vel cuneatis, integris; inflorescentiis terminali-
bus paniculatis amplis ubique glabris et pellucido-punctatis
regulariter trifurcatis; floribus 4-meris.

Shrub (7); branches slender, glabrous, somewhat longitud-
inally fissured, suberose, and ridged in drying; leaves opp-
osite, trifoliolate; petioles slender, 2--6 cm. long, gla-
brous, conspicuously pellucid-punctate; leaflets sessile,
thin-coriaceous, oblanceolate, 3.5--12 cm. long, 1.5--4 cm.
wide, glabrous and conspicuously pellucid-punctate on both
surfaces, shiny above, glaucescent beneath, rounded at the
apex, long-attenuate or cuneate at the base, entire, the
margins slightly eubrevolute (in drying, at least); inflor-
escence terminal, paniculate, ample, about 12 cm. long and
to 14 cm. wide at the base, glabrous and pellucid-punctate
throughout, regularly trifurcate, the branches wide-spread-
ing; peduncles about 1.5 cm. long, glabrous, pellucid-punct-
ate; inflorescence-branches or flowers borne in pairs at
each node of the infllorescence, in cymose fashion, one on
each side of the axis, each subtended by a triangular-acute
glabrous scale-like bractlet about 1 mm. long; pedicels very
slender, about 2 mm. long, glabrous; flowers 4-merous; sep-
als 4, heavy, tough, suborbicular, about 1 mm. long and
wide, rounded at the apex, imbricate, scarious-margined,
glabrous on both surfaces except for the slightly erose-
ciliolate margins, very convex on the back, concave within;
petals 4, oblanceolate-lingulate, about 3 mm. long and 1.3
mm. wide, translucent-margined, rounded at the apex, only
slightly narrowed toward the base, glabrous, pellucid-punct-
ate; disk large, cupuliform, the erect rim about 0.78 mm.
high, irregularly undulate and scaly; stamens 4, inserted at
the base of the rim of the disk, among the scales; filaments
terete, about 1.5 mm. long, translucent, broadened at the
base, attenuate above the disk; anthers about 0.5 mm. long,
2-celled, apiculate at the apex, the 2 cells divergent at
the base; pistil solitary, central; style very short, blunt,
about 0.4 mm. long, terminated by a discoid stigma of the
same diameter as the style; ovary 4-celled; ovules 2 in each
cell, apically attached; fruit not seen.

The type of this species was collected by Brothers Clé-
ment, Chrysogono, and Alain [Clément 3971] at Mina Cayoguan,
Pta. Gorda, Oriente, Cuba, on July 21, 1944, and is deposit-
ed in the Britton Herbarium at the New York Botanical Gard-
en. The species is obviously related to H. glaucescens Urb.,

224 PR UE P'OrL Oo Gi*é Vol. 2, noe 7

the only other known West Indian species of the genus, which
differs in its shorter petioles, shorter and narrower leaf-
lets, very much smaller and narrower non-trifurcate inflor-
escences, puberulent inflorescence-branches, pedicels,
bractlets, and sepals, more triangular-ovate and acute sep-
als, and slightly larger anthers.

HYPERBAENA LONGIUSCULA var. CLEMENTIS Moldenke, var. nov.

Haec verietas a forma typica speciei foliis oblongo-ell-
ipticis 15--16.5 cm. longis, 5--6 cm. latis, ad apicem ob-
tusis vel rotundatis, ad basin non attenuatis, et costa sup-
ra argute elevatis recedit.

This variety differs from the typical form of the species
in having leaves with blades oblong-elliptic, 15--16.5 cm.
long, 5--6 cm. wide, obtuse or rounded at the apex and base,
not attenuate to the base, and the midrib very sharply ele-
vated on the upper surface from the base to the apex.

The type was collected by Augustin Clement Teteau, Broth-
er Clément (no. 3633) on the new way to Rfo Yegrumajes, east
of Moa, Oriente, Cuba, on May 17, 1944, and is deposited in
the Britton Herbarium at the New York Botanical Garden.

IPOMOEA BATATAS f. TRIFIDA Moldenke, f. nov.

Haec forma a forma typica speciei foliis profunde tripar-
titis recedit.

This form differs from the typical form of the species in
having all of its leaves uniformly deeply 3-parted or the
lateral lobes sometimes again bifurcate. The lobes are ob-
lanceolate, long-acuminate at the apex, and attenuate to the
base.

The type was collected by Reinaldo Hspinosa (no. 492) in
cultivated and irrigated land at La Fornia, alt. 1400 Mey
Loja, Ecuador, on June 5, 1946, and is deposited in the Brit
ton Herbarium at the New York Botanical Garden. It is called
“camote indio" by the natives.

IPOMOEA CARNEA f. ALBIFLORA Moldenke, f. nov.

Haec forma a forma typica speciei corollis albis recedit.

This form differs from the typical form of the species in
having white corollas.

The type was collected by Reinaldo Espinosa (no. 490) at
La Fornia, alt. about 1400 m., Loja. Ecuador, on June 5,
1946, and is depasited in the Britton Herbarium at the New
York Botanical Garden. The collector states that it was
growing among plants of the typical form.

IPOMOEA DUMETORUM f. ALBA Moldenke, f. nov.
Haec forma a forma typica speciei corollis albis recedit.
This form differs from the typical form of the species in

1947 _ Moldenke, New and Noteworthy Plants 225

heaving white corollas.

The type was collected by Reinaldo Espinosa (no. 215a) at
La Argelia, southern Loja, Ecuador, on April 25, 1946, and
is deposited in the Britton Herbarium at the New York Botan-
ical Garden.

LANTANA DEPRESSA Small (Fig. 1)

Literature: Small, Bull. N. Y. Bote Gard. 3: 436. 1905;
Small, Addisonia 3: 69--70, pl. 115. 1918; Moldenke, Annot.
List 108. 1939; Moldenke, Known Geogr. Distrib. Verbenac. 5
& 94, 1942.

This species used to be justly considered a rare and lit-
tle-known one, but so much splendid collecting done in Flor-
ida during the past 30 years has yielded so many collections
thet the species can no longer be regarded as anything but
well-known. The following is a list of some of the specimens
recently annotated by me in my monographic work on the
group:

FLORIDA: Dade Co.: Bailey & Bailey 6016 (Ba), 6217 (Ba),
6217a (Ba); N. Le Britton 156 (N), sen. [Miemis April 1,
1903] (Cm); Buewell son. [Mey 14, 1934] (Bu), sen. fApril
21, 1935] (Bu); Demaree 10208 (Bt, Du, Hp); Zlder 442 (H),
910 (H); Esselbaugh sen. [Frinceton, March 16, 1946] (Ur);
Harshberger sen. [August 15, 1911] (Up); Hawkins s.n. [Home-
stead, 9/16/27] (Fl, Fl); Lightfoot s.n. [Key Biscayne, Apr.
28, 1917] (Ba); Be McAllister 315a (H); He Ne Moldenke =
(Go, H, I, N, N, Up, Ur); O'Neill 1941 (I), 7133 (1), 713
(I); 7135 (T)5 ot Or 7138 (I), 7140 (cee gen. [Feters,
September 17, 1929] (1), sen. [Princeton, September 19,
1929] (I); Be H. Patterson sen. [Miami, Feb. 2, 1918] (Cm);
W. W. Rowlee sen. [Dec. 23, 1902] (It); J. Ke Small 2217
(N), 3645 (N), 7255 (N, N), 8793 (N)s Small & Carter 747 (N
--type), 2678 (N), sen. [Oct. 31st to Nov. 4th, 1903] (We);
Small & Mosier 5523 (N), 6367a (N); Small, Mosier, & Small
5667 (N), 6506 (N); Small & Nash 180 (N); Small & Small 4781
(Ny, 4808 (It), 4818 (Go, N), 6524 (Fl, Mi, N, Up)s Small &
Wilson 1826 (N), sen. [May 16, 1904] (H); Tidestrom 6987
(I); Welch 1536 (Dp); Je P. Young 195 (It), 204 (It). Saint
Lucie Co.: O'Neill 7145 (I).

Explanation of Figure 1: a, Habit, x 3/4; b, bractlet,

x 5; c, celyx, x 5; d, pistil, x 5; e, corolla split open
and flattened out, x 5.

LANTANA HISPIDA var. TERNATA Moldenke, var. nov.

Haec varietas a forma typica speciei foliis ternatis re-
cedit.

This variety differs from the typical form of the species
in having its leaves whorled in groups of three.

The type was collected by Gustavo Aguirre B. and B. P.

226 PHYTOLOGIA Vol. 2, now 7

Reko (no. 172) at Necaxa, Puebla, Mexico, in April, 1946,
and is deposited in the Britton Herbarium at the New York
Botanical Garden.

LIPPIA BRACTEOSA (Mart. & Gal.) Moldenke, comb. nov.
Lantana bracteosa Mart. & Gal., Bull. Acad. Roy. Brux.,
sér. 1, ll (2): 326. 1844,

LIPPIA LIBERIENSIS Moldenke, sp. nov.

Frutex vel arbor; ramis orscure tetragonis vel subtereti-
bus dense breviterque pubescentibus glabrescentibus; nodis
annulatis; foliis oppositis; petiolis dense breviterque pub-
escentibus, in statu senectute spareissime strigillosis vel
pilosulis; leminis firme membranaceis ovato-lanceolatis ad
apicem argute acutis serrulatis, ad basin acuminetis, in
statu juventute utrinque dense breviterque pubescentibus, in
statu senectute supra scabris et plusminusve strigosis, sub-
tus strigillosis pulverulentisque; inflorescentiis axillari-
bus capitatis nutantibus.

Shrub or tree; branches and branchlets obscurely tetrago-
neal or subterete, densely short-pubescent when very young,
glabrous in age, light-brownish; nodes annulate; principal
internodes 1.5--7 cm. long; leaves decussate-opposite; peti-
oles slender, 3--5 mm. long, densely short-pubescent when
very young, very sparsely strigillose or pilosulous when me-
ture, the hairs mostly in 2 bends on the axial surface;
blades firmly membranous, ovate-lanceolate, 2.5--725 cme
long, 1-5--3.5 cme wide, sharply acute at the apex, reguler-
ly serrulate from the base to the apex and more or less pro-
longed into the petiole, densely short-pubescent on both
surfaces when young, scabrous and more or less strigose a-
bove or subglabrous when mature, irregularly strigillose on
the venation beneath and usually more or less pulvenulent
beneath when mature; inflorescence axillary, capitate, nod-
ding, usually 2 per node near the apex of the branches or
branchlets; peduncles very slender, 2--2.5 cm. long, dense-
ly short-pubescent or puberulent; heads 1.3--1.6 cm. long,
1.7--2.4 cm. wide, many-flowered; bracts broadly ovate, the
lowest about 1.5 cm. long and almost 1 om. wide at the base,
subacuminate at apex, densely puberulent.

The type of this species was collected by Alberto Brenes
in the vicinity of Liberia, Guanacaste, Costa Rica, in 1910,
and is deposited in the Britton Herbarium at the New York
Botanical Garden.

MOZARTIA EMARGINATA Moldenke, sp. nove

Frutex vel arbor; remulis graeilibus griseis glabris;
foliis oppositis; petiolis gracilibus glabris; laminis levi-
ter coriaceis ellipticis vel oblanceolatis emarginatis, ad

227

1947

—=S

Cx

Liege

LAL

\

228 Pon YF Orbs Or Gils Vol. 2, noe 7

basin acutis vel acuminetis, utrinque glabris nitidisque non
punctatis integris.

Shrub or tree; branchlets and twigs slender, gray, glab-
rous; leaves decussate-opposite; petioles slender, 6--10 mm.
long, glabrous; blades thin-coriaceous, elliptic or oblance-
olate, 3.5--6.5 cm. long, 1./--3 om. wide, emarginate at the
apex, acute or acuminate at the base, glabrous and shiny on
both surfaces, not noticeably punctate, entire or slightly
wavy-margined, slightly subrevolute at the margins when dry;
midrib slender, impressed above, very strongly prominent be-
neath; secondaries very slender, about the same size and
strength as the tertiaries and veinlets, they, with the
veinlets, forming a dense conspicuous reticulum which is e-
qually and beautifully prominulous on both surfaces, a rath-
er indistinct collective vein uniting the secondaries near
the margins; inflorescence axillary, apparently few-flower-
ed; flowers not seen; fruiting peduncles slender, 1.5--2 om.
long, glabrous; fruiting-calyx incrassate, glabrous, persis-
tent, closely appressed to the base of the fruit, about 5
mm. in diameter (including the lobes), the lobes 4, ovate-
triangular, about 1.5 mm. long and 1 mm. wide at the base,
subacute at the apex; fruits 1--3 per peduncle, sessile on
the peduncle, hard, globose, about 6 mm. long and wide,
glabrous.

The type of this handsome species was collected by George
C. Bucher (no. 14253) at Moa, Oriente, Cuba, in July, 1939,
and is deposited in the Britton Herbarium at the New York
Botanical Garden.

PADUS CAPULI (Cav.) Moldenke, comb. nov.
Frunus Capuli Cav. ex Spreng., Syst. Vege 2: 477. 1825.

PAEFALANTHUS ESPINOSIANUS Moldenke, sp. nov.

Herba caespitosa; foliis numerosis firmis patentibus lan-
ceolato-attenuatis argute apiculatis utrinque plusminusve
pilosulis glabrescentibus strietis non fenestratis; vaginis
laxis glabris, ad apicem bilobatis, lobis ovatis erectis;
pedunculis solitariis gracilibus brevibus 3-costatis tortis
obscure pilosulis; capitulis obconico-hemisphaericis,.

Tufted herb; stems very much abbreviated, long-villous at
the apex, 1--2 cm. long; leaves numerous, bright-green,
firm, spreading, lenceolate-attenuate, 1--1.5 cm. long, 1.5
--2 mm. wide at the mid-point, sharply apiculate at the a-
pex, more or less scattered-pilose on both surfaces when
young, glabrescent in age, several-striate, not plainly fen-
estrate; sheath loose, glabrous, 1.3--1.5 cm. long, the bas-
al tubular portion 7--8 mm. long, the epical portion split
into 2 ovate, erect, dissimilar lobes almost 1 cm. long;
peduncles solitary at the apex of each stem, slender, almost

1947 Moldenke, New and Noteworthy Plants 229

obsolete or to 2.3 cm. long, 3-costate, twisted, obscurely
pilosulous, more persistently so beneath the head; heads ob-
conic-hemispheric, about 5 mm. in diameter; involucral
bractlets few, in 2 series, light-brown, lanceolate, about 5
mme long, about 1.5 mm. wide at the widest point, attenuate-
acute or subacuminate at the apex, glabrous and shiny
throughout, surpassing the florets, concave on the inner and
convex on the outer surface; receptacle long-villous; recep-
tacular bractlets narrowly oblong, about 2.6 mm. long, light
brown, darker toward the apex, navicular, about 0.4 mm.
wide, more or less appressed-villous on the back with ant-
rorse hairs, not bearded; staminate florets short-pedicell-
ate; sepals 3, separate practically to the base, dark-brown
on the upper half, oblong-oblanceolate, about 2.1 mm. long
and 0.5 mm. wide, obtuse at apex, more or less villous on
the back, with very much appressed antrorse hairs, bearded
at the apex; petals 5, united into a slender lightly stram-
ineous tube about 1./ mm. long, slightly ampliate at the a-
pex, the lobes erect, lanceolate-ovate, about 0.5 mm. long,
acuminate at apex, somewhat involute; stamens 3; filaments
filiform, very short, inserted at the base of the corolla-
lobes and opposite them; anthers not seen; pistillate flor-
ets short-pedicellate: pedicels about 0.6 mm. long; sepals
3, separate practically to the base, erect, brown, darkest
on the upper half, spatulate, about 2.1 mm. long, about 0.6
mm. wide at the widest part, acute at the apex, Long-vill«
ous with antrorse hairs on the back, usually more or less
bearded at the apex on the back; petals 3, separate to the
base, fitting snugly between the ovary-wings, lightly stram-
ineous, erect, oblanceolate, about 2.1 mm. long and 0.6 mm.
wide, acute or apiculate at apex, more or less villous on
the back, especially along the margins above the middle and
at the apex, not bearded, not glanduliferous; style about
0.8 mm. long, glabrous, terminated by 3 erect stigmas and 3
style-appendages which are all 0.6--0.8 mm. long; ovary ell-
iptic, deeply 3-lobed and Z-alate, ¢g glabrous, 3-celled.

The type of this species was collected by Julian A. Stey-
ermark (no. 54342) in dense tufts in moist places on a para-
mo et 11,200 feet elevation along the trail between FPailas
and £1 Pan, Santiago-Zamora, Ecuador, on September 10, 1943,
amd is deposited in the Britton Herbarium at the New York
Botanical Garden. The species resembles P. Karstenii Ruhl.
in habit. It is named in honor of Dr. Reinaldo Espinosa, who
is doing such noteworthy work on the flora of Scuador.

PAEPALANTHUS LOXENSIS Moldenke, sp. nov.

Herba caulescens; ramis gracilibus usque ad 10 cm. longis
brachiatis dense longeque villosis; foliis numerosissimis
firmis patentibus apiculatus utrinque glabris nitidis non

230 PHYTOLOGIA Vols 2, no. 7

fenestratis; vaginis brevibus profunde fissis, lobis lanceo-
latis acuminatis glabris erectis; pedunculis solitariis 3-
costatis paullo tortis ubique glabris; capitulis hemisphaer-
icis griseis vel stramineis.

Caulescent matted herb; stems slender, to 10 cm. or more
long, branched, densely long-villous, especially at the a-
pex, completely hidden by the abundant imbricately sheathing
leaf-bases except toward the base on older stems; leaves ab-
undant, rather firm, spreading, about 1 cm. long or less, a-
bout 1 mm. wide at the mid-point, apiculate, essentially
glabrous on both surfaces, not fenestrate; sheath hidden a-
mong the upper leaves, about 1.2 cm. long, deeply split to
below the middle, the 2 lobes equal, lanceolate, about 7 mm.
long, acuminate, glabrous, erect, but remote from the pedun-
cle; peduncles usually solitary at or near the tip of each
branch or stem, 4--5 cm. long, 3-costate, slightly twisted,
glabrous throughout; heads hemispheric, gray or stramineous,
hairy, 3--4 mm. in diameter; involucral bractlets elliptic,
very concave on the inner and convex on the outer surface,
stramineous or grayish, 5--4.5 mm. long, 1.5--1.7 mm. wide,
acute or slightly apiculate, more or less villous on the
back especially along the margins and at the apex with ant-
rorse hairs, usually somewhat short-bearded at the apex; re-
ceptacle densely long-villous; receptacular bractlets nar-
rowly spatulate, 1.7--1.9 mm. long, about 0.4 m. wide,
dark-brown toward the apex, hyaline at base, blunt at apex
and there densely bearded, otherwise glabrous, slightly nav-
icular; steminate florets: sepals 3, connate only at the
very base, oblanceolate, 1--1.3 mm. long, about O.4 mm.
wide, obtuse at apex, brown toward the apex, much lighter
toward the base, glabrous except for the densely bearded a-
pex; petals 3, united into an infundibular stramineous tube
about 0.8 mn. long, glabrous, the lobes lanceolate, erect,
about O.4 mm. long, not glanduliferous, glabrous; stamens 3,
inserted at the mouth of the corolla-tube; filaments about
O.3 mm. long, glabrous; pistillate florets: sepals 3, appar-
ently separate to the base, spatulate, dark-brown toward the
apex, much lighter or stramineous toward the base, about 1.5
mm. long and 0.6 mm. wide, rounded at apex, long-pilose on
the inner surface with antrorse hairs; petals 4, separate,
spatulate, hyaline, about 1.3 mm. long and 0.6 mm. wide,
long-pilose along the margins and toward the apex, not
bearded, not glanduliferous; style about O.4 mm. long, glab-
rous; ovary subglobose, deeply 3-lobed and-sulcate, glab-
rous, 3-celled; stigmas 3, about O.4 mm. long, erect; style-
appendages 4, about the same length as the stigmas and issu-
ing from the same level.

The type of this species was collected by Julian A. Stey-
ermerk (no. 54432 ), growing in dense mats on moist banks,

1947 Moldenke, New and Noteworthy Plants 231

between Tambo Cachiyacu, La mtrada, and Nudo de Sabanillas,
altitude 2500--3500 m., Loja, Ecuador, on October 7, 1943,
and is deposited in the Britton Herbarium at the New York
Botanical Garden. The species resembles P. Glaziovii Ruhl.,
but differs in the length of its peduncles and in floral
characters.

PAEPALANTHUS STEINBACHII Moldenke, sp. nov.

Herba acaulescens; foliis rosulatis leviter membranaceis
recurvo-adpressis linearibus fenestratis argute attenuatis
glabris; veginis arcte adpressis glabris, ad apicem oblique
fissis, lamina argute acuta; pedunculis numerosis gracilli-
mis 2-costatis tortis glabris; capitulis hemisphsericis
brunneis vel nigris.

Acaulescent herb; leaves basal, rosulate, thin-membran-
ous, appressed to the ground or ascending, shorter than the
peduncles, linear, 4--5 cm. long, about 1.5 mm. wide at the
middle, many-nerved, plainly fenestrate throughout, sharply
attenuate at the apex, glabrous throughout; sheaths closely
appressed to the peduncle, about 45 cm. long, glabrous, ob-
liquely split at the apex, the blade sharply acute; pedun-
cles numerous, about 20 per plant, very slender, 6--11 cm.
long, 2-costate, twisted, glabrous throughout, far surpass-
ing the leaves; heads hemispheric, brown or black, 3--4 mm.
in diameter; involucral bractlets lanceolate, hyaline or
grey, 1--1.5 mm. long, acute at the apex, glabrous on both
surfaces; receptacle apparently glabrous; receptacular
bractlets narrowly oblong, falcate, about 2 mm. long and
O.4 mm. wide, dark-brown or black on the upper half, the
lower half hyaline, acute at the apex, glabrous throughout,
not bearded; staminate florets: sepals 3, elliptic, about
1.3 mm. long and 0.2 mm. wide, black, united into a slender
tube at the base, acute, glabrous throughout; petals 3, a-
bout 1 mm. long, united into a hyaline tube, the free por-
tions 0.2--0.4 mm. long, hyaline, acute, glabrous, with a
black gland near the center on the back; stamens 3; anthers
globose, yellow, about 0.1 mm. long; pistillate florets:
sepals 3, narrow-elliptic or oblong, about 1.2 mm. long and
0.3 mm. wide, dark-brown or black except at the very base,
acute, glabrous throughout; petals 3, separate to the base,
falcate, narrowly oblong, about 1 mm. long and O.1 mm.
wide, brownish, glabrous throughout, not bearded, not
glanduliferous; style about 0.6 mm. long, glabrous; stigmas
3, about 0.4 mm. long; ovary oblong, about 0.3 mm. long and
0.2 mm. wide, brown, slightly granular, 3-celled, 3-ovulate

The type of this species was collected by Jose Steinbach
(no. 2669) at Campos de Terebinto, on the banks of a small
lake, Santa Cruz, Bolivia, on August 22, 1916, and is dep-

osited in the Britton Herbarium at the New York Botanical

232 FHYTOLOGIA Vol. 2, taa
Garden.

PAEPALANTHUS SUBSESSILIS Moldenke, sp. nov.

Herba valde pumila dense caespitosa; caulibus valde abb-
reviatis, ad apicem dense longeque villosis; foliis rosula-
tis lanceolatis 6--8 mm. longis, ca. 1 mm. latis, argute a-

piculatis multistriatis non fenestratis, in statu juventute
margine plusminusve longe giliatis recurvatis, in statu sen-
ectute glabrescentibus; inflorescentiis subsessilibus soli-
tariis; pedunculis 1.5--1.8 mm. longis antrorse adpresso-
pilosis, bracteis 4 foliaceis ad basin connatis glabris sub-
tentis.

Very dwarf densely despitose herb; stems very much abb-
reviated, densely long-villous at the apex, obscured by the
dense leaves; leaves rosulate, lanceolate, 6--8 mm. long, a-
bout 1 mm. wide at the mid-point, sharply apiculate at the
apex, greatly ampliate and sheathing the stem at the base,
many-striate, not fenestrate, membranous at the base, more
or less long~ciliate on the margins when young, completely
glabrous in age, recurved; inflorescence subsessile, soli-
tary at the epex of the stem; typical sheath absent; pedun-
cle absent or obsolete, 1.5--1.8 mm. long, antrorsely ap-
pressed-pilose, subtended by 4 leaf-like bracts which are
connate at the base into a short tube about 1 mm. long,
then bilabiate into 2 exactly similar wide-spreading lips
about 3.5 mm. long, each lip deeply bifid almost to the
base, each lobe ovate, about 2.5 mm. long and 1 mm. wide,
attenuate to an acute apex, ascending-spreading, curvate,
and appressed to the head, light-brown, glabrous throughout;
heads hemispheric, 3.5--4 mm. in diameter, gray-brown, vil-
lous; involucral bractlets few, in 1 or 2 series, broadly
ovate, very concave on the inner and convex on the outer
surface, brown, very dark at the apex, about 2,5 mm. long
and 1.8 mm. wide, abruptly acute at the apex, glabrous
throughout except for the very small tuft of short hairs on
the back at the very apex; receptacle very densely long-
villous; receptacular bractlets few, elliptic-oblanceolate,
about 1.7 mm. long and 0.4 mm. wide, dark-brown throughout,
rounded and subcucullate at the apex, densely bearded on
the back at the apex, otherwise glabrous; staminate florets:
sepals 3, separate to the base, dark-brown, elliptic, con-
cave on the inner and convex on the outer surface, about 1.3
mm. long and 0.5 mm. wide, densely bearded on the back at
the apex, otherwise subglabrous; petals 3, connate into a
slender hyaline tube about 0.6 mm. long, the free lobes lan-
ceolate, about 0.3 mm. long, hyaline, erect, blunt; stamens
3, inserted about 1/2 way down the corolla-tube; filaments
about 0.2 mm. long, glabrous; anthers white,oblong, about
0.4 mm. long, composed of two separate versatile thecae de-

1947 Moldenke, New and Noteworthy Flants 233

hiscing by means of longitudinal slits; pistillate florets:
sepals 3, separate, broadly obovate, navicular-conduplicate,
dark-brown or blackish, about 1.4 mm. long and O.7 mm. wide
(when flattened out), subcordate at the apex, rather densely
long-villous on the back from below the middle to the epex,
densely bearded on the back et the apex; petals 3, separate,
hyaline, elliptic, about O.8 mm. long and 0.4 m. wide,
densely long-villous on the beck, not bearded, not glandu-
liferous; style ebbreviated, about O.l mm. long, glabrous,
terminated by 3 erect stigmas about 0.1 mm. long and 3 erect
style-appendages about C.2 mm. long; ovary subglobose, about
O.3 mm. long and wide, J-angled, glabrous, 3-celled.

The type of this distinct species was collected by Julian
A. Steyermark (no. 55495) on a dry paramo at an elevation of
10,000 feet, between Buenos Aires and Paramo de las Rosas,
Lara, Venezuela, on February 11, 1944, and is deposited in
the Britton Herbarium at the New York Botanical Garden. The
species resembles P. lodiculoides Moldenke in habit, the
lutter growing in similar habitats in Colombia. The collect-
or records the common neme "flor de tierra".

PHYLA STRIGULOSA (Mart. & Gal.) Moldenke, comb. nov.
Lippia strigulosa Mart. & Gal., Bull. Acad. Roy. Brux.,
Sere As oe tens 319. 1844,

PHYLA STRIGULOSA vare PARVIFOLIA (Moldenke) Moldenke, comb.
NOVe

Phyla yucatana var. parvifolia Moldenke, Phytologia 2:
141--1 ee 19 e

PTERIDIUM LATIUSCULUM f. BERDII Moldenke, Am. Midl. Nat. 35:
313, hyponym (1946), f. nov.

Haec forma a forma typica speciei laminis non ternatis
differt.

This form differs from the typical form of the species in
having the lowest pinnae of its fronds not much larger than
those immediately above them; thus the fronds are not at all
ternate. Mr. C. A. Weatherby, of the Gray Herbarium of Har-
vard University, in & letter to me datea August 1, 1945, a-
grees that this character is probably transmissible, but ap-
parently regards it as merely a clone. The type was collect-
ed by Morris and DeEtta Berd [H. N. Moldenke 16593] in wood-
lands, North Werren, Warren County, Pennsylvania, on July
31, 1944, and is deposited in the herbarium of the Carnegie
Museum at Pittsburgh. The form occurs quite abundantly in
this region, usually mixed with the typical form, but often
in large numbers. It has a very distinct appearance in the
wild. Like the typical form, its fronds are very often heav-
ily infested by Cryptomyces pteridis (Rebent.) Rehm.

234 PHYTOLOGIA Vole 2, now 7

STACHYTARFHETA CONFERTIFOLIA Moldenke, sp. nov.

Frutex; reamis gracilibus subteretibus sarmentosis glabra-
tis; internodiis valde abbreviatis; foliis sessilibus oppos-
itis dense confertis succulentis anguste ellipticis parvis
obtusis integris, ed basin angustatis, utrinque glabris sub-
tus dense punctatis.

Small shrub; branches slender, subterete, dark, twiggy,
glabrate; twigs similer, densely foliate; internodes ex-
tremely abbreviated, 1--5 mm. long; leaves sessile, decuss-
ate-opposite, crowded, probably somewhat fleshy, nerrowly
elliptic, 7--13 mm. long, 2.5--5 mm. wide, obtuse at the a-
pex, entire, narrowed to the base, glabrous on both surfaces
and densely punctate beneath; midrib very slender, obscure
on both surfaces; vein and veinlet reticulation indiscern-
ible; inflorescence terminal, spicate, solitary; spikes ses-
sile, 3--4 cm. long, densely many-flowered; flowers closely
imbricate; rachis minutely puberulent, rather deeply sculp-
tured after the fruit has fallen; brectlets lanceolate-
ovete, about 6.3 mm. long and 2.7 mm. wide, gradually atten-
uate to the sharply acute apex, glabrous; calyx heavy-tex-
tured, tubular, about 1 cm. long and 1.8 mm. wide, glabrous
throughout, its rim 5-toothed, the teeth unequal, ovate-
triengilar, 1.3--1.8 mm. long, 0.9--1.3 mm. wide, abruptly
acute at the apex; corolla hypocrateriform, its tube slend-
er, about 1.5 cm. long and 1.5 mm. wide, glabrous, its limb
5-parted, the lobes unequal, broadly obovate-elliptic, 5--8
mm, long, 4--7 mm. wide, rounded at the apex, slightly
wavy-margined, conspicuously venose, glabrous; stamens 2,
inserted about 11.7 mm. above the base of the corolla-tube;
free portion of filaments filiform, about 0.9 mm. long;
glabrous; anthers dorsifixed near the middle, about 1.8 mm.
long; style capillary, about 2 cm. long, glabrous; stigma
capitate, about 0.5 mm. wide; ovary ovate, about 1.8 mm.
long and 1.5 mm. wide, glabrous; fruiting-calyx and fruit
not seen.

The type of this very distinct species was collected by
Louis 0. Williams and Vicente Assis (no. 6639) in a campo at
Serra da Mutuca, beyond Barreiro, Municipio of Nova Lima, at
an altitude,of 1400 m., Minas Geraes, Brazil, on April 15,
1945, and is deposited in the Gray Herbarium of Harvard Uni-
versity. .

STACHYTARPHETA STEYERMARKII Moldenke, sp. nov.

Plante lignosa; ramis acute tetragonis decussato-puberu-
lis plerumque alatis; nodis ampliatis annulatis; foliis op-
positis; petiolis dense pubescentibus paullo marginatis;
laminis leviter chartaceis elliptico-ovatis acutis arcute
serratis, supra parce breviterque pubescentibus, subtus den-
se breviterque pubescentibus.

1947 Moldenke, New and Noteworthy Plants 235

Woody plant; branches acutely tetragonal, puberulent on
alternate pairs of sides, the angles often slightly margin-
ed, often somewhat ampliate and annulate at the nodes; prin-
ciple internodes 1.5--4 cm. long; leaves decussate-opposite;
petioles slender, about 1 cm. long, rather densely pubes-
cent, somewhat margined; blades thin-chartaceous, elliptic-
ovate, 3--4 cm. long [immature?], 1.3--2.5 cm. wide, acute
at the apex, regularly sharp-serrate from almost the base to
the apex, rather sparsely short-pubescent above, densely
short-pubescent beneath; midrib very slender, plane above,
prominulent beneath; secondaries very slender, 5 or 6 per
side, ascending, slightly arcuate, plene above, very slight-
ly prominulent beneath; inflorescence terminal; peduncles to
about 3 cm. long, puberulent on one pair of sides like the
branches; floriferous portion of the spikes to about 20 cm.
long efter anthesis; rachis stout, about 4 mm. in diameter,
puberulent-strigillose on two opposite sides, deeply sculp-
tured in fruit; bractlets lanceolate, 5--7 mm. long, gradu-
ally attenuate to a long-acuminate apex, appressed or re-
curved after anthesis, minutely puberulent or glabrate, more
or less ciliolate along the margins, barely equaling the
calyx during anthesis and then more densely puberulent; cal-
yx about 5 mm. long, densely puberulent; corolla dark-vio-
let, hypocrateriform, somewhat exserted from the calyx, its
tube sbout 7 mm. long, glebrous.

The type of this species was collected my good friend,
Dr. Julian A. Steyermark (no. 54834), in dry rocky desert
hills ebove La Toma, alt. 1520--18 30 me, Loja, Ecuador, on
October 24, 1943, and is deposited in the herbarium of the
Chicago Natural History Museum. It is a pleasure to dedicate
this species to Dr. Steyermark, who has done such uniformly
splendid and valuable collecting in the southern United
States, Central America, and South America.

SYMBOLANTHUS MACRANTHUS (Benth. ) Moldenke, comb. nov.
Lisianthus macranthus Benth., Plant. Hartweg. 144. 1839.

VERBENA BAILEYANA Moldenke, hybre nov.

Herba; ramis erectis brachiatis tetragonis sulcatis al-
bo-strigillosis; nodis annulatis; petiolis obscuris late a-
latis; foliis chartaceis ovatis, ad basin cuneatis, ad api-
cem acutis, irregulariter inciso-laciniatis utrinque strig-
illosis supra scabris.

. Herb; stems erect, branched, rather sharply tetragonal,
sulcate between the angles, strigillose with short, stiff,
white, antrorsely subappressed hairs; nodes annulate; prin-
cipal internodes 2.5--5 cm. long; leaves decussate-opposite,
usually with several small ones on very abbreviated twigs in
their axils; petioles obsolete or to 2 cm. long and broadly

236 PB Ye 0) BO God oe Vol. 2, noe 7

winged, merging indistinguishably into the base of the
blade; blades chartaceous, rather uniformly bright-green on
both surfaces, 7--10 cm. long, 2.5--6 cm. wide, ovate in
outline, acute at apex, cuneately narrowed into the broadly
winged petiole at the base, irregularly and deeply incised-
laciniate, the lowermost lobes on the largest leaves often
hastate, strigillose on both surfaces with short subappress-
ed antrorse whitish hairs, scabrous above when the finger is
drawn from the apex toward tne base; midrib slender, impres-
sed above, prominulous beneath; secondaries slender, 4 or 5
per side, ascending, not much arcuate, irregularly branched,
a branch usually extending to the tip of the larger lobes;
veinlet reticulation subimpressed above, plane but visible
beneath; inflorescence a terminal panicle, the lowest
branches of which are axillary to the uppermost much reduced
leaves, the panicle about 15 cm. long and 5 cm. wide, its
branches erect or ascending, strigillose-puberulent; bracts
lanceolate, 5--8 mm. long, densely strigillose; bractlets
similar but smaller, attenuate; calyx about 2.5 mm. tong,
densely strigillose, slightly exceeding the bractlets.

The type of this hybrid was collected from cultivated ma-
' terial in the Royal Botanical Garden at Paris in 1819 and is
deposited in the Dudley Herbarium at Stanford University. It
is apparently a hybrid between V. officinalis L. and V. has-
tata L., with, in general, intermediate characters. It is
named in honor of Dr. Liberty Hyde Bailey, distinguished
worker on cultivated plants, sedges, and palms.

VERBENA CONCEPCIONIS Moldenke, sp. nov.

Herba; ramis gracilibus hirsutulis; foliis oppositis vel
suboppositis sessilibus; laminis chartaceis, ad apicem acut-
is, ad basin longe cuneatis, irregulariter 3-lobatis, lobis
j-dentatis supra rugoso-strigosis subtus strigoso-pubsscent-
ibus; spicis multifloris densis dein valde elongetis.

Herb, to about 40 cm. tall; stems erect, branched to the
base, sparingly hirsute with stiff white hairs about 1 mn.
long and standing at right angles to the stem; branches
slender, gach terninating, like the stem, in a single elon-
gated spike, hirsutulous, less so in age; principal inter-
nodes 2--6.5 om. long; leaves decussate-opposite, or the up-
per ones subopposite (like the branches), sessile; blades
chartaceous, uniformly light-green on both surfaces, 2--2.8
cm. long, 10--21 mm. wide, widest et about the middle or
slightly below, abruptly acute at the apex, long-cuneate at
the base, irregularly 3-lobed, each lobe about 3-dentate
with coarse subacute teeth, rough and strigose above, stri-
gose-pubescent beneath, the venation somewhat impressed a-
bove and prominulous beneath; peduncles slender, continuous
with the stems or branches, 2--6 cm. long, sparsely hirsut-

1947 Moldenke, New and Noteworthy Plants 237

ulous like the stems and branches; spikes many-flowered, at
first dense, later elongating to 12 cm., with the fruits
rather distant toward the base, more crowded toward the apex
and foliaceous bracts often subtending the lower pairs of
fruits; bractlets narrow-lanceolate, about 3 mm. long,
strigillose-hirsutulous, sharply attenuate-acute at the a-
pex, almost equaling the fruiting-calyx; rachis short-pubes-
cent and somewhat glandular; calyx about 3 mm. long, strig-
ose with rather long white hairs and often also somewhat
glandular-puberulent; corolla-tube about 5 mm. long, puberu-
lent outside above the calyx, its limb about 2 mm. wide;
fruiting-calyx not accrescent, strigose and somewhat glandu-
lar-puberulent.

The type of this species was collected by Louis Née (no.
57) at Concepcion, Chile, between 1789 and 1791, and is de-
posited in the herbarium of the Jardin Botanico at Madrid.

VERBENA JORDANENSIS Moldenke, sp. nov.

Herba perennis brachiatis; ramis obtuse tetragonis den-
siuscule patenteque pubescentibus brunneis; foliis opposit-
is subsessilibus; petiolis alatis; laminis ovatis triparti-
tis, partibus lobatis vel incisis, lobis angustis obtusis
supra asperulo-strigillosis subtus in reticulo venularum
patenti-pilosulis subrevolutis; inflorescentiis axillaribus
racemosis paucifloris.

Ferennial herb, about 16 cm. tall, abundantly branched
from the base, bushy; branches decussate-opposite, obtusely
tetragonal, rather densely spreading-pubescent, brown when
dry; principal internodes 1--2 cm. long; leaves decussate-
opposite, subsessile; petioles 1 mm. long or less, winged;
blades ovate, tripartite, 7--25 mm. long, 5--15 mm. wide,
each of the 3 divisions again lobed or incised, the lobes
narrow and obtuse, more or less asperulous-strigillose a-
bove, spreading pilosulous on the venation and margins be-
neath, the margins subrevolute; midrib and secondaries sub-
impressed above, prominulous beneath; inflorescence axil-
lary, racemose, the racemes few-flowered, l--2 cm. long in
fruit, usually less than 1 cm. long at anthesis; peduncles
filiform, obsolete in anthesis, to 1 cm. long in fruit,
and spreading-pubescent like the branches; bractlets ovate,
about 3 mm. long, 1 mm. wide at the base, attenuate to the
sharply acute apex, rather irregularly long-ciliate along
the margins, especially toward the base, reflexed in age;
calyx tubular, about 4 mm. long, 5-costate, very shortly 5-
apiculate on the rim, the costae spreading-pubescent, oth-
erwise glabrate, the apiculations coherent after anthesis;
corolla hypocrateriform, its tube narrow-cylindric, about 4
mn. long, the limb about 2 mm. wide, glabrous, irregularly
S-lobed; ovary subtended by a cupuliform disk about 1 mm.

238 . PHY TOL OG Is Vol. 2, now 7

in diameter, which remains in the axil of the bractlet after
the fruiting-calyx and fruit have been shed; fruiting-calyx
not enlarged, readily splitting when the fruit matures, the
apiculations remaining coherent almost up to the time of
shedding of the fruit; nutlets narrowly oblong, about 2 mm.
long, reticulate-scrobiculate on the back on the upper half,
with parallel longitudinal ridges on the lower half, glab-
rous, shiny.

The type of this most interesting and distinct specie
was collected by José Eugenio Leite (no. 3474) in wet places
and fields, Campos do Jordad, at al elevation of 1600 m.,
S&o Paulo, Brazil, in April, 1945, and is deposited in his
herbarium. The conspicuous disks beneath the ovary, which
remain in the axils of the reflexed bractlets after the
fruiting-calyx and fruit have been shed, render this species
most remarkable and show its relationship to V. thymoides
Cham.

VERBENA MARITIMA Small (Fig. 2)

Literature: Small, Bull. N. Y. Bot. Gard. 3: 436. 1905;
Moldenke, Annot. List 108. 19359; Prelim. Alph. List Invalid
Names 26. 1940; Known Geogr. Distrib. Verbenac. 5 & 101.
1942; Alph. List Invalid Names 25. 1942; Addisonia 21: 59--
60, pl. 702. 1942.

Synonyms: Glandularia maritima Small, Man. Southeast. Fl.
1138. 1933. Verbena Aubletia var. maritima Curtiss ex Mol-
denke, Addisonia 21: 60, in syn. 1942.

This species is usually considered to be a rare species
by most herbarium curators because of its very limited dis-
tribution and the general paucity of material representing
it in their collections. However, much splendid collecting
in Florida over the past 30 years has brought to light so
much material of Verbena maritima and it grows there in such
profusion that it cannot really be considered to be a little
known plant any more. Following is a list of the material of
this species which has come to me recently during the course
of my monographic studies of the group. Harris has reported
it also from Martin and Monroe Counties.

FLORIDA: Brevard Co.: F. Se Blanton 6309 (I, N), 6479
(I); A. B. Burgess 638 (N); Curtiss 1963* (Bc, C, Om, I, Up,
Vt), 5706 (Al, Bc, Fl, Io, N, N, Fo, oh een. [Fle.] (C);
Degener s.n. [Tropic, Aug. 15, 1933] (Ba); N. Hotchkiss sen.
[Feb. 5, 1935] (N); McFarlin 6608 (N); H. N. Moldenke 2192
(N, Up, Ur); O'Neill sen. [dune 11, 1929] (I), sen. [July 8,
1929] (I), sen. [south of Cocoa Beach, August 9, 1929] (Fl,
I); W. He Rhoades sen. (Cocoa, 12-8-27] (Fl); U. C. Smith s.
ns (Georgiania, Jany. 31, '91] (Up). Broward Co.: Ce C. Deam
60837 (Dm, N). Collier Co.: Sheenan a.n. [Leaning Oak] (N),
sen. [Godden's Mission, March 7, 1919] (N). Dade Co.: Bailey

1947 Moldenke, New and Noteworthy Plants

239

—- _—-

ZA SN yy

Verbena maritima Smal)

240 PHYTOLOGIA Vole 2, noe 7
& Bailey 6278 (Ba, Ba), 6388 (Ba); Ne Le Britton 220 (N),

296 (N); Buewell sen. [April 3, 1942] (Bu); Ce C. Deam 60417
(pn, N), 60940 (pm, N); A. P. Garber s.n. (Miami, May 1877]
(vt); Hawkins sen. [Royal Palm State Park, 1-25-28] (Fl);
Henderson sen. |Cape Florida] (T); Herb. Columbia Univ. s.n.
(Cape Florida] (C); Hunnewell 5835 (Ua); Lightfoot s.n. [Key
Biscayne, Apr. 28, 1917] (Ba); Be McAllister 27 (H); H. Ne
Moldenke. 549 (Go, N, Up, Ur), 586 (Go, H, N, N, Up, Ur);
Mulvania 12 (Hp); O'Neill 7596 (Bt, Du, Hp, Hp, I, N, N, St,
Ur), sen. [Jan. 30, 1933] (1); Be He Patterson sen. [Feb. 7,
1918] (Gm, Cm); J. Ke Small 2100 (N), 8123 (N), 8594 (N),
8599 (N), sen. [beach opposite Miami, November 1904] (ur);
Small & Carter 1077 (N--type, N--isotype), 2994 (N), sen.
[January 16, 1909] (We); Small, Carter, & Small 3311 (N), 5.
n. (February 1911] (H, Pl); Small & Small 5422 (Fl, N), sen.
Touly 9, 1915] (N); Small & Wilson 1961 (N); Weber & Hawkins
sen. (Homestead, 3-1-28] (Fl). Flagler Co.: O'Neill sen.
(August 7, 1929] (1); West & Arnold s.n. [Flagler Beach, 10/
10/40] (Fl). Indian River Co.: Small, DeWinkeler, & Mosier
11123 (N), sen. [April 3, 1924] (It, Mi). Lee Cow: Je Ke
Small 8347 (N). Palm Beach Co.: Bailey & Bailey 6523 (Ba,
Ba); A. Be Burgess 783 (N); W. H. Rhoades sen. [near Palm
Beach] (Hs, fie 3 Je Ke Small 2124 (N), 8509 (Go, Io, It, N);
Small, Mosier, & DeWinkeler 10891 (Up); E. West sen. [Jupi-
ter, 5-12-33] (Fl). Saint Lucie Co.: A. B. Burgess 713 (N).
Volusia Co.: B. He. Patterson sen. (Daytona, Nov. 28, 1917]
(Cm); J. K. Small 8674 (N); Smell & DeWinkeler 9856 (Mi).
County undetermined: Herb. Le Roy sen. [Florida] (C).

Explanation of Figure 2: a, Habit, x 3/4; b, flower, x 2

1/4; c, calyx and its subtending bractlet, x 2 1/4; d, cor-
olla slit open and flattened out, x 2 1/4; e, pistil, x 2
1/4; f,immature flower, abnormal. :

VERBENA MATRITENSIS Moldenke, hybr. nov.

Herbe; caulibus tetragonis densiuscule hirsutulis, pilis
albidis rigidis patentibus; foliis lanceolatis leviter char-
taceis argute acutis, ad basin cuneato-acuminatis, irregu-
lariter dentatis vel inciso-dentatis supra strigoso-scabris,
subtus strigilloso-scabrellis; inflorescentiis paniculatis.

Apparently a natural or ertificial hybrid between V. car-
olina L. and V. hastata L., exhibiting more or less inter-
mediate characters; stems tetragonal, rather densely hirsut-
ulous with whitish, stiff, spreading hairs; leaves lanceo-
late, thin-chartaceous, 4.5--7 cm. long, 1.4--2.4 cm. wide,
sharply acute at the apex, cuneate-acuminate at the base,
irr@gularly dentate or incised-dentate along the margins
from almost the base to the apex, strigose-scabrous above,
strigillose-scabrellous beneath; inflorescence paniculate,
or 1 or 2 spikes terminating short lateral branches; spikes

ra bh.

1947 Moldenke, New and Noteworthy Plante 24)

narrow, elongate, to about 7 cm. long, rather densely flow-
ered, apparently not setting seed, the short peduncles and
slender rachis puberulent with appressed gray hair; bract-
lets lanceolate, about 2 mm. long, acuminate, glabrate on
the back, sparsely ciliolate along the margins at the widest
part; calyx strigillose, slightly exceeding the subtending
bractlets; corolla-tube about 4 mm. long, its limb about 2
mm. wide.

The type of this hybrid was collected in the Royal Botan-
ic Garden at Madrid, Spain, and is deposited in the Britton
Herbarium at the New York Botanical Garden.

VERBENA NEEI Moldenke, sp. nov.

Herba parva; caulibus gracilibus decumbentibus vel ad-
scendentibus apicem versus brachiatis breviter pubescenti-
bus, pilis albis rigidis; ramis brevibus; foliis oppositis
sessilibus; laminis crasso-charteceis irregulariter inciso-
lobatis revolutis utrinque strigoso-pubescentibus, ad basin
longe cuneatis, lobis acutis; spicis sessilibus vel subses-
silibus laxiuscule multifloris.

Herb, to about 20 cm. tall; stems slender, rather harshly
short-pubescent with stiff white hairs about 0.5 mm. long,
apparently decumbent or ascending, several issuing from the
base of the plant, branched toward the apex; branches short,
terminating (like the stem) in an elongated spike; leaves
decussate-opposite, sessile; blades thick-chartaceous, 1--2
ome long, 5--12 mm. wide, widest at or below the middle, ir-
regularly incised-lobed, the margins revolute, strigose-pub-
escent on both surfaces, long-cuneate at the base, the lobes
acute; spikes sessile or subsessile, to 10 cm. long, rather
loosely many-flowered, usually about 3 spikes at the apex of
the stem, the flowers densely crowded in bud, berely imbric-
ate in fruit; bractlets lanceolate, about 2.5 mm. long, ac-
uminate at the apex, strigillose on the back, somewhat short-
er than the fruiting-calyx; calyx about 2.5 mm. long, dense-
ly strigose; corolla slightly exserted; fruiting-calyx about
3 mm. long, densely strigose.

The type of this species was collected by Louis Née (no.
108 ) -- in whose honor it is named -- on the pampas at Bue-
nos Aires, Argentina, between 1789 and 1791, and is deposit-
ed in the herbarium of the Jardin Botanico at Madrid.

X VIOLA MILLERI Moldenke, nom. nov.

Viola affinis x triloba Brainerd, Vermont Agr. Exp. Sta.
Bull. 239: 44--45. 1924; Moldenke, List Observ. Fl. Watchung
24. 1940.

VIORNA PSEUDOCOCCINEA (Schneid.) Moldenke, comb. nov.
Clematis pseudococcinea Schneid., Wien Ill. Gartenz. 29:

242 PHYTOLOGIA Vol. 2, noe 7
15. 1904; Ill. Handb. Laubholzk. 1: 279-6 1904.

WEIGELTIA SCHLIMII var. INTERMEDIA Moldenke, var. nov.

Haec varietas a forma typica speciei recedit inflorescen-
tiis 9 sessilibus vel subsessilibus, floribus 45 mm. longis,
petalis 2 mm. latis, antheribus 1 mm. longis glabris, fila-
mentis 1 mm. longis, et fructibus ca. 1 cm. in diametro.

This variety differs from the typical form of the species
in having sessile or subsessile pistillate inflorescences
(observed in fruit) and slightly larger flowers (observed in
bud only), the petals being about 5 mm. long and 2 mm. wide,
the glabrous anthers as long as the filaments, each about 1
mn. long, and the fruit about 1 cm. in diameter, smooth,
very wrinkled in drying. In the typical form of the species
the pistillate inflorescences (in fruit) have a peduncle a-
bout 5.5 cm. long, the petals are only 4 mm. long end 1 m.
wide, the filaments are about 2 mm. long, and the anthers
are very small, about 0.25 mm. long, pilose at the base,
while the fruit is only about 7 mm. in diameter, pustulate,
not wrinkled. In W. multiflora A. C. Sm. the pistillate in-
florescences are not known, but the staminate flowers have
their petals 53.5--5 mm. long and 1.9--2.5 mm. wide, the an-
thers about 1 mm. long and the filaments about 2 mm. long,
and the leaves are relatively much narrower and differ mar-
kedly in the details of their venation.

The type of this variety was collected by José Cuatrecas-
as (no. 17312) at San Isidro, altitude 5--100 m., Rfo Cajam-
bre, on the Facific coast, El Valle, Colombia, between May 2
and 5, 1944, and is deposited in the Britton Herbarium at
the New York Botanical Garden.

ADDITIONAL NOTES ON THE GENUS TIMOTOCIA. I

Harold N. Moldenke

Since the publication of my monograph of this genus in
Fedde, Repert. Sp. Nov. 39: 129--153 (1936) seventy-nine ad-
ditional specimens and photographs of specimens have been.
examined. The small number of specimens that have come to
light in these eleven years is a good indication of the
scarcity of material of this genus in the world's herbaria ~
and the rarity of the members of the group. The material
hereinafter cited is deposited in the herbaria indicated by
the following symbols: B = Botanisches Museum, Berlin; Br =
Jardin Botanique de 1'Etat, Brussels; Cb = Delessert Herbar-

|
,

1947 -Moldenke, Notes on Timotocia 243

ium, Conservatoire et Jardin Botaniques, Geneva; De = De
Candolle Herbarium, Conservatoire et Jardin Botaniques, Gen-
eva; E = Missouri Botanical Garden, St. Louis; Ed = Royal
Botanic Garden, Edinburgh; Ja = Museu Necional, Rio de Jan-
eiro; K = Royal Botanic Gardens, Kew; Kr = Krukoff Herbarium
New York Botanical Garden, New York City; Lu = Botanisk Mus-
eum, University of Lund, Lund; Mu © Botanisches Museum, Mun-
ich; N = Britton Herbarium, New York Botanical Garden, New
York City; P = Muséum National d'Histoire Naturelle, Paris;
Us = Botaniska Institutionen, Uppsala; W = United States
National Herbarium, Smithsonian Institution, Washington; and
Z = He N. Moldenke Herbarium, Watchung, New Jersey.

TIMOTOCIA Moldenke

References: Casselia Nees & Mart., Nov. Act. Fhysico-med.
Acad. Caes. Leopold.-Carol. Nat. Cur. 11: 73, pl. 6, figs. A
& B. 1823; Reichenb., Conspect. Reg. Veg. 1: 117. 1828;
Walp., Repert. 4: 40. 1845; Schau. in A. DC., Prodr. 11:
527. 1847; Paxt., Fl. des Serres pl. 361. 1848; Paxt., Mag.
Bot. 15: 75. 1849; Schau. in Mart., Fl. Bras. 9: 1743, pl.
32. 1851; Bocq., Rev. Verb. 141, pl. 16. 1861--1863; Benth.
in Benth. & Hook. f., Gen. Pl. 2 (2): 1148. 1876; Junell,
Symb. Bot. Upsal. 4: 18. 1934; Moldenke in Fedde, Repert.
Sp. Nov. 39: 129. 1936; Moldenke, Alph. List Invalid Names
12. 1942. Timotocia Moldenke in Fedde, Repert. Sp. Nov. 39:
129--153. 1936; Chron. Bot. 3: 311. 1937; Alph. List Invalid
Names 12. 1942; Known Geogr. Distrib. Verbenac. 38, 40, 41,
74, & 100. 1942; Prelim. Alph. List Invalid Names 14. 1940;
Alph. List Citations 237 & 264. 1946.

Junell in the reference cited above states that the flor-
al morphology, especially that of the gynoecium and androe-
cium, and the vegetative morphology point to a close relat-
ionship between this genus and Ghinia Schreb. The two genera
are, however, widely separated in the classification of Bri-
quet. Reichenbach, in the reference cited above, places the
genus (as Casselia Nees) among the accepted genera in his
section Verbeneae of the family Labiatae.

TIMOTOCIA CHAMAEDRYFOLIA (Cham. ) Moldenke

On page 145 of my monograph four specimens of Saint-
Hilaire s.n. [Olho d'Agoa, 1816--1821] are cited from the
Paris herbarium. The figure should be 3, since one of these
sheets is now in the Britton Herbarium at New York.

Illustrations: Warming, Dansk. Vid. Selsk. Skrift, ser.
6, 6: 197. 1892; Drude, Okol. Pflanzen [Die Wissensch. 1:]
66. 1913.

Additional citations: BRAZIL: Goyaz: J. E- Pohl 442 (N).
Minas Geraes: Saint-Hilaire s.n. [Olho d'Agoa, 1816--1821]
(N); Sellow 1517 [Macbride photos 17576] (Kr--photo of type,

244 PE.YoT OL.OG. TA Vol. 2, noe 7

P--isotype), 1519 (B, B, B).s Rio de Janeiro: Sellow sen.
[Peyal, 1818] (B, B)- a

TIMOTOCIA CONFERTIFLORA Moldenke

The Macbride photograph cited below is erroneously label-
led "Gardner 3371" and “Rasselin 3371".

Additional citations: BRAZIL: Goyaz: G. Gardner 3369
([Macbride photos 34298] (Br--isotype, Kr--photo of isotype).

TIMOTOCIA CONFERTIFLORA var. LACINIATA Moldenke

Tnis plant has been misidentified in some herbaria as
Lantana brasiliensis Link.

Additional citations: BRAZIL: Goyaz: G. Gardner 35570
[Macbride. photos 34297] (Br--isotype, Sd--isotype, Kr--photo
of isotype).

TIMOTOCIA GLAZIOVII Briq. & Moldenke

This species has been misidentified in some herbaria as
"Lantana turneraefolia Cham.", "Lippia turnersefolia Cham.",
and "Scrophul.

Additional citations: BRAZIL: Goyaz: Glaziou 21890 [Mac-
bride photos 24626, in part] (B, Br, K, Kr--photo); J» Be
Pohl 2158 [Herb. Imp. Vindob. 142; Macbride photos 34296]
cai of type). State undetermined: J. E. Pohl sen. ©
(Br e

TIMOTOCIA HASSLERI (Briq.) Moldenke

Material of this species is mixed with Basistemon spinos-
us (Chod.) Moldenke, of the Scrophulariaceae, on the Berlin
sheet of Hassler 7450, cited below.

Additional citations: PARAGUAY: Hassler 7550, in part
(B), 7889 [Mecbride photos 24627] (Cb--isotype, Kr--photo of
type, N--fragment of type, P--isotype), 10760 (Cb, Cb, F).

TIMOTOCIA HYMZNOCALYX (Briq.) Moldenke

Additional citetions: PARAGUAY: Fiebrig 4092 [Herb. Mon-
uce 4147] (Cb, Cb, Ed, Mu); Hassler 7637 [Macbride photos
24628] (Cb--isotype, Kr--photo of type, P--isotype). CULTI-
VATED: Paraguay: Rojas 2406 [Herb. Parag. Jard. Bot. Parag.
10834] (Mu).

TIMOTOCIA INTEGRIFOLIA (Nees & Mart.) Moldenke

References: Casselia semiserrate Hort. ex Moldenke, Pre-
lim. Alph. List Invalid Names 14, in syn. 1940; Alph. List
Invalid Names 12, in syn. 1942.

Illustrations: Paxt., Flore des Serres, ser. 1, 4: pl.
361(colored). 1848; Paxt., Mag. Bot. 15: 75 (colored). 1849.
Additional citations: BRAZIL: Piauhy: G. Gardner 2272
{N). Rio de Janeiro: L. Riedel 404 (De); Riedel & Luschnath

1947 Moldenke, Notes on Timotocia 245

1328 (N); Ule sen. [Oct. 1899] (B). State undetermined: Lund
394 (Cb). CULTIVATED: Belgium: Hort. Mons. e.n. [18393 type
coll. of Casselia semiserrata Hort.] (Br). ILLUSTRATION,
colored (Br).

TIMOTOCIA INTSGRIFOLIA var. FISCHERI (Mart. ) Moldenke

The Macbride photograph 20347 cited on page 137 of my
monograph as a photograph of an isotype of this variety is
actually a photograph of the type. The collection label on
the Krukoff Herbarium sheet cited below is inscribed "Bra-
zil", apparently in error, for the plant of which this is a
photograph was collected in cultivation in Germany, though
it may have originated in Brazil.

Additional citations: CULTIVATED: France: Herbs Baillon
sen. [Hort. Paris] (P). Germany: Lucae a.n. (Herb. Kummer;
Herb. Monac. 1281; Macbride photos 20347] (Kr--photo of
type, Mu--photo of type).

TIMOTOCIA MANSOI (Schau. ) Moldenke

References: Casselia Mansii Schau. ex Moldenke, Prelim.
Alph. List Invalid Names 14, in syn. 1940; Alph. List In-
valid Names 12, in syn. 1942. Casselia Mansoi Schau. in A.
DC., Prodr. 11: 527. 1847; Schau. in Mart., Fl. Bras. 9: 75,
pl. 32. 1851; Junell, Symb. Bot. Upsal. 4: 18. 1934. Cassel-
ia peduncularis Mart. ex Moldenke, Prelim. Alph. List Inval-
id Names 14, in syn. 1940; Alph. List Invalid Names 12, in
syn. 1942.

Illustrations: Mart., Fl. Bras. 9: pl. 32, fig. 2. 1851;
Booq., Rev. Verbenac. pl. 16. 1861--1863; Briqe in Engl. &
Prantl, Nat. Pflanzenfam. 4 (3a): 157. 1895.

Additional citations: BRAZIL: Goyaz: G. Gardner 5371 (Br,
Cb, Cb, P); Weddell 2789 (P). Mattogrosso: Collector undes-
ignated s.n. [Morro do Ernesto Cuiaba; as “Casselia Mansii
Schau."] (Br); Malme 2524 (Lu, Us); Silva-Manso ene [Herbs
Martius 1025; Herb. Monac. 421; Macbride photos 24629] (Br--
isotype, De--type, Kr--photo of isotype, Mu-~-isotype, N--
photo of type, P--isotype, Z--photo of type). State undeter-
mined: Herb. Baillon sen. (F); J. Ee Pohl 144 [type coll. of

Casselia peduncularis Mart. ] (Br), Sen. (Br). BOLIVIA: Santa

(Seer es

Cruz: Steinbach 5 (Cb). ILLUSTRATIONS: Mart. Fl. Bras.

pl. 32 B).

TIMOTOCIA ROSULARIS (Sandw.) Moldenke

This species is described by Archer and Gehrt as having
long tuberous roots and pink flowers with the odor of cinna-
mon (Cinnamomum zeylanicum).

Additional citations: BRAZIL: Mattogrosso: Archer & Gehrt
152 (N, W), sen. [Herb. Inst. Biol. S. Paulo 36320] (N);
Malme 2449a (B, E--photo, N--photo, Z--photo).

246 PB Y¥ 27-0. L ©: G56 Vol. 2, now 7

TIMOTOCIA SERRATA (Nees & Mart.) Moldenke

References: Casselia serrata Nees & Mart., Nov. Act.
Physico-med. Acad. Caes. Leopold.-Carol. Nat. Qur. 11: 75--
76, pl. 6, fig. A. 1823; Junell, Symb. Bot. Upsal. 4: 18 &
19. 1934.

Junell, in the reference cited above, discusses the gyn-
oecium morphology of this species.

Illustrations: Nov. Act. Physico-med. Acad. Caes. Leopold
-Carol. Nat. Cur. ll: pl. 6, fig. A. 1823; Symb. Bot. Upsal.
4s fig. hole. pl. I, fig. le 1934.

Additional citations: BRAZIL: Bahia: Wied-Neuwied s.n.
-(Brasilia] (Br, Lu). Minas Geraes: Dusen sen. (Herb. Rio de
Janeiro 32253] (Ja); Glaziou 13060 [Macbride photos 24626,
in part] (Kr--photo). .

TIMOTOCIA VERONICAEFOLIA (Cham.) Moldenke
Additional citations: BRAZIL: Minas Geraes: Sellow 1518
(Macbride ‘photos 17577] (Kr--photo of type & isotype).

TIMOTOCIA ZELOTA Moldenke
Additional citations: BRAZIL: Minas Geraes: Blenchet ples
(Cb--isotype).

ADDITIONAL NOTES ON THE GENUS AMASONIA. II

Harold N. Moldenke

AMASONIA L. f.

Taligalea and Amasonia are both included among the accep-
ted genera in the section Verbeneae of the family Labiatae
by He G. L. Reichenbach in his Conspect. Reg. Veg. 1: 117

AMASONIA CAMFESTRIS (Aubl.) Moldenke |
Additional citations: SURINAM: Maguire 23780 (N). BRAZIL:
Maranh&o: G. Don H.136 ["85"] (Br).

AMASONIA HIRTA Benth.

The type collection of this species was made “in Brazil
by Pohl and Langsdorff" according to Bentham's original des-
cription, not by Schomburgk as erroneously stated by me on
page 205 of my monograph. The Pohl s.n. [Villa Boa; Herb.
Monac.e 931] collection cited by me on the same page is ac-
tually from the state of Goyaz.

Additional citations: BRAZIL: Goyaz: Ule 451 [Herb. Rio

1947 Moldenke, Notes on Amasonia 247

de Janeiro 32275] (Ja). Mattogrosso: Martius 583 [Herb. De
Candolle 827; Herb. Monac. 929; Macbride photos 7886 &
20346] (Kr--photo, Kr--photo). Minas Geraes: Temberlik s.n.
(F). S&o Paulo: L. Riedel s.n. [Ville Franca, June 1834) (W).

“ AMASONIA LASIOCAULOS Mart. & Schau.

References: Moldenke, Prelim. Alph. List Invalid Nemes 5
& 42. 1940; Alph. List Invalid Names 4 & 43. 1942; Known
Geogr. Distrib. Verbenac. 36 & 86. 1942; Phytologia 2: 91.
1945.

Cuatrecasas collected this species at an altitude of 230
m. in Colombia, and describes the bracts and calyx as red
and the corolla as yellow, blooming in September. The "Ducke
85" cited on page 210 of my monograph is an error for Huber
85.
~ Additional citations: COLOMBIA: Vaupes: Cuatrecasas 7017
(Ww).

AMASONIA OBOVATA Gleason

References: Fedde, Bot. Jahresber. 59 (2): 416. 1939;
Moldenke, Known Geogr. Distrib. Verbenac. 32 & 86. 1942.

The species is described by Steyermark as a "shrubby
herb" or shrub, growing at 200--1095 m. altitude, with a
single, simple, erect stem 4 to 5 feet tall, membranous
leaves that are dark- or dull-green above and pale dull-
green or dull-purple beneath, bracts dark-red or deep rose-
red, corolla pale-greenish, filaments greenish-white, and
anthers brown-lavender, blooming in August. The species is
doubtfully distinct from A. arborea H.BeK.

Additional citations: VENEZUZLA: Amazonas: Steyermark
57871 (F--1205147, N), 57980 (F--1205146, N).

AMASONIA SPRUCEANA Moldenke

References: Moldenke, Known Geogr. Distrib. Verbenac. 30,
32, 36, 71, & 86. 1942.

Cuatrecasas describes this species as having "bracts and
flowers red". He collected it et an altitude of 200 m. in
Colombia, in flower and fruit in September.

Additional citations: COLOMBIA: Vaupes: Cuatrecasas 6853
(Ww). VENEZUELA: Amazonas: Spruce 3288 [Macbride photos
28391] (F--photo of isotype, Kr--photo of isotype).

Excluded species:

Amazonia integerrima Spreng. apud Stendl., Contrib. Us. Se
Net. Herb. 25: 1335. 1926 = Braveisia integerrima (Spreng. )
Standl.

248 PHYTOLOGIA Vol. 2, no. 7

ADDITIONAL NOTES ON TH GENUS PETREA. III

Harold N. Moldenke

PETREA Houst.

This genus is placed in the section Verbeneae of the fam-
ily Lebiatee, as an accepted genus, by H. Ge Le Reichenbach
in his Conspect. Reg. Veg. 1: 117 (1828), where he writes the
name "Fetrea Le" It is classified in the same way by Reichen-
beach in Méssler, Hendb. Gewaéchsk., ed. 1, 1: xxvi (1827) and
ed. 3, 1: lxxv (1833).

An excluded species is Fetrea scandens Nee, in herb.,
which is a synonym of Coffea arabica Le. of the Rubiaceae.

PETREA ARBOREA HeBeKe

Steyermark records the common name “nacareno" for this
plant and describes it as a shrub 5 feet tall with chartace-
ous leaves, wrinkled along the outer margin, and bracts deep-
purple, growing at altitudes of 1065--1220 m.

Additional citations: VENEZUELA: Lera: Steyermark 55546
(F--1205142 ).

PETREA ASPERA Turcz.

Steyermark records the common name "flor de mayo" én des-
cribes the plant as a high-climbing liana with "firmly mem=-
branaceous-chartaceous" or "subcoriaceous-chartaceous" leaves
that are deep- or rich-green above and dull-green or paler
rich-green beneath, the calyx rich-lavender with purplish-
blue midrib, the corolla deep-lavender, blooming in April at
altitudes of 230--820 m. in rocky upland chaparral and valley
savannas.

Additional citations: VENEZUELA: Sucre: Steyermark 62368
(F--1205703, N), 62801 (N).

PETREA BRACTEATA Steud.

A common name for this species in British Guiana is " gand-
paper vine" and the plant is described as a soft gray “rope”
‘with stems 1--2 cm. thick, growing in the crowns of small
trees by creeks; with leathery, supple or stiff, rugose, sca-
hrous leaves, the upper surface of the young ones “navy-blue
green like the spike"; flowers in small axillary or terminal
dark purple-green spikes, the corolla tubular, purple or vio-
let-purple, heiry in the throat; the epicalyx membranous,
violet or pale-blue when fully expanded.

Additional citations: BRITISH GUIANA: Forest Dept. Br.
Guiana 4470 (F.1734] (N), 4471 [F.1735] (N).

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on the suitability of manuscripts will be solicited, if necessary, from quali- 3
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qQuiries may be addressed to the magazine or to either editor. a

Designed to expedite botanical publication

June, 1947 No. 8

CONTENTS

Guzason, H. A., Nokia on some American plants....... he ae
7 ILEASON, H. A., A letter from Ferdinand von Mueller................291

GLEASON, H. A., New or noteworthy melastomes,
ie Bee CHMCTUY Pe CUAG OF CAN a. .5cescschunus dens thdenbeete eardcatgv lamer
i

Y

: [OLDENKE, H.N,, Notes on new and noteworthy plants. I1........306
Mo .penkeE, H.N., The known geographic distribution of the
re) : members of the Verbenaceae and
| Avicenniaceae. Supplement 6 ....ccrcrsesssessesserse DOO
M nENES, H. LN, The known geographic distribution of the
Eriocaulaceae. Supplement 1 ....ccsssescssscseeessee. O49

Published by H. A. Gleason and Harold N. Moldenke __ ini
The New York Botanical Garden,
Bronx Park, New York 58, N. Y.
Price of this number $1.00; per volume, $5.00 in advance

Vol. 2, No. 7, was issued April 22, 1947

AW
a
ia
i

Natt ie
wey
ea

3 LIBRARY
NEW YORK
BOTANICAL

GARDEN

DEBUTS OF WOODY PLANTS IN CULTIVATION

Data on Introduction and First Appearances -- in relation
to those in Rehder's MANUAL OF CULTIVATED TREES AND SHRUBS,
Ed. 1940.

P. Je van Melle

Tnese data are submitted by way of a casual contribution
rather than of an exhaustive study. They represent an acoumr
lation in the compiler's notes -- many of them by-products
of other inquiries. The years or blanks in the right-hand
columns are those occurring in the MANUAL. Except in the few
instances marked "(*)" the nomenclature employed is that of
the MANUAL.

The presentation of these data in relation to that work
implies no criticism of it. On the contrary, the MANUAL is
regarded as 6 great central structure, about which minor
contributions like this cluster themselves like little shops
between buttresses of a cathedral. The method saves needless
duplication of information conveniently available. Only such
data are given here as are at variance with, or not given in
the MANUAL. Data of this kind should perhaps always be re-
garded as more or less tentative -- subject to adjustment
upon further investigation. Some allowances should be made
for misidentifications of materials in the sources cited.
Reasonable care has been exercised to avoid cases of dis-
cernible misidentifications.

Intriguing as is the subject of plant debuts, it appears
not to have been explored exhaustively. Fartly because of
the greater accessibility of English literature, much is
known, comparatively, about introductions into England, and
little about those into continental Europe, the records of
which are, moreover, for a large part not readily available.
Too little is known popularly about the important introduc-
tions from Japan by von Siebold and by Maximowiocz, and those
by Russian travellers, from Siberia, Mongolia and North-
China. Specialized investigation might bring out worth-while
facts about seeds and living plants received in Paris from
the French missionary, Armand David, of whose introductions
little is known. Among them is probably, for instance, the
comparatively important Pfitzer Juniper of gardens.

Much study remains to be made of the history of plant in-
troductions before it can be viewed in a proper perspective
and in relation to explorations projected from many lands.

249

-

250 Pa YT 0:30 Gtt sé \ Vol. 2, noe 8

A few titles of periodicals, catalogs and other works ap-
pearing in the lists in more or less abbreviated form

A.H-Be eeeoeevevovoeveoes@

Ann. Soc. Agricult.
Bretschneider weceoe
Bull. F.S- @eeeeseosee2d

Cox SE ot OL Fe

Gartfl. e@eseeeneaeee 06

Hort. Berole ceoccee

Hort. Breite eeeccs

Hort. Dinegro cece.

Illustr. Hortic. ee

Leiden eeseeeooe oe ee8

Montreuil ecccccces

Annales d'Horticulture et de Botanique.
Leiden, 1858--1862. Published by von
Siebold and W. H. de Vriese. Devoted
largely to plants brought from Japan
and from Java.

Annales de la Société d'Agriculture et
de Botanique de Gand.

E. Bretschneider - History of European
Botanical Discoveries in China. 1898.

Bulletin de la Federation des Socistes
d'Horticulture en Belgique.

Ee. He Me Cox - Plant Hunting in China.
1945.

Gartenflora.

Hortus Berolinensis. A catalogue of the
Botanical Garden at Berlin.

Hortus Breiterianus, A catalog of a
semi-commercial botanical collection
maintained in Leipsig by Mmristian
August Breiter. 1817.

A catalog, by D. Viviani, of a botani-
cal collection privately owned by J.
Car. Dinegro, in Genoa. 1802.

L' Illustration Horticole.

Many data otherwise unannotated of
plants cultivated in Leiden have been
derived from a jubilee publication:
Hortus Academicus Lugduno Batavus, by
H. Veendorp and Baas Becking, 1938.
Other data have been derived from old
indices of this Academic Garden.

Refers to a "Catalogue des Arbres dans
les jardine de M. Lemonnier a Mont-
reuil - copie sur de ms. de l'auteur,
dont Me. le Prof. Adrien de Jussieu

- i . : | }
1947 van Melle, Woody Plants 251

est en possession." It is dated 1774,
We found it, in the library of the New
York Botanical Garden, bound in a
"Catalogue méthodique des vegéetaux
cultivés dans le Jardin des Plantes de
la Ville de Versailles" » by F. He
Philippar, 1843.

Pavlovak .eeceeesee Refers to a catalog, by I. A. Weinmann,
of an Imperial Garden in Pavlovsak,
near Petrograd.

Prince eeeeeeeeeren Catalogs of the Wm. Prince nursé6ry =
the "Linnsean Botanic Garden" =
Flushing.

Rev. Hortic. .seee.ee Revue Horticole.

Soc. Gardeners .... "Catalogue of Trees, Shrubs, Plants
and Flowers both exotic and domestic
which are propagated for sale in the
Gardens near London". By "A Society of
Gardeners". 1730. - The similarity of
the style of this work to that of
Philip Miller's Gardeners Dictionary
suggests thet he compiled or edited it.

Von Siebold & Co. . The catalogs referred to include the
"Kruidkundige Naamlijsten" in the 1844
and 1845 yearbooks of the Nederlandsch
Koninklijke Maatschappij tot Aan moed-
iging v. d. Tuinbouw. (These Yearbooks
are the “Annuaires Pays-bas" of some
authors).

Von Siebold ....... “Sur l!Etat de l'Horticulture su Japon
et sur l'Importance des plantes usuel-
les et d'ornament introduites et cult-
ivées dans le Jardin d'Acclimation de
M. Ph. von Siebold a Leide". A bro-
chure by von Siebold. 1863.

Uppsala .seceesceeee Hortus Upsalensis, 1748. - A catalog, by
‘ Linneeus, of his garden at Uppsala.

Vol. 2, Ne 8

PHYTOLOGIA

252

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Vol. 2, noe 8

PHYTOLOGIA

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255

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Vol. 2, no. 8

PHY TO DOG I%

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257

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1947

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Vol. 2, noe 8

PEE POLOG bk

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259

van Melle, Woody Plants

1947

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Vol. 2, noe 8

PHYTOLOGIA

260

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PHYTOLOGIA

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Vol. 2, noe 8

PHYTOLOGIA

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277

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1947

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ADDITIONAL NOTES ON BLAKEA AND TOPOBEA

j

H. Ae Gleason

1. To anyone who has examined the anthers, the distinct-
ion between Blakea and Topobea is obvious. Nevertheless it
is impossible to refer a plant without anthers to either
genus, since the various patterns of leaves, inflorescence,
bracts, and calyx are often repeated in both. It is quite
probable that some species described without flowers have
been assigned to the wrong genus. I described Topobea alt-
ernifolia without knowledge of the flowers and twenty years
elapsed before the collection of flowering specimens showed
my error and led me to transfer the species to Blakea,

Cogniaux admitted twenty-four species of Topobea i: in his
monograph of 1891. Some of these he had not seen himself,
nor were the anthers described by the original author.
Whether these species actually belong to Topobea can only be
surmised. They must have been assigned there solely on their
general facies and that is completely unreliable.

Since then sixteen species have been proposed and not yet
transferred. Of these barbate Gl., Brenesiana Standl.,
brevibractea Gl., Auspidete Gl., discolor Hochr., Durandiana
Cogn., floribunda Gl., longisepala Gl., pubescens Gl., and
rupicola Hoehne undoubtedly belong to this genus, while fer-
ruginea Gl., Maurofernandeziana Cogn., pluvialis Standl.,

_ rosea Gl., Storkii Standl., and urophylles Standl. may or may
not belong, so far as my present knowledge of them is con-
cerned.

2. When Blakea calyptrata was described, it was noted
that a second sheet from a thousand meters lower elevation
differed in no "technical" characters except the width of
the bracts; it was given a varietal name. Now a third sheet
has been seen which again differs in no important characters
except the shape of the leaf. While the type of the species
came from the western slope of the Cordillera Occidental,
alt. 960-1180 m., this was collected on the eastern slope at
an altitude of 2500 m. I propose verietal rank for it also.

BLAKEA CALYPTRATA Gl. var. OVATA, ver. nov. A typo dif-
fert foliis ovato-lanceolatis acuminatis.

Type, Cuatrecasas 21684, collected at Hoya del Rfo Cali,
en La Palma, Dept. del Valle, Colombia.

3. Examination of additional material and repeated com-
pearison with older collections still convince me that the
species mentioned by me (Bull. Torrey Club 72: 1. 1945) form

279

\

' ,
/

280: PHYTOLOGIA Vole. 2, now &

a group easily segregated by their general ainiiaeeee and
therefore a practical group for purposes of identification.
I am not satisfied, however, that the species are actually
phylogenetically related, as stated by me earlier (Ann. Mo.
Bot. Gard. 28: 434. 1941).

Blakea Andreana Cogn. was the first species of the group
to be described. The type was collected near Vijes in the
valley of the Cauca River, alt. 1800 m. So far as foliage is

concerned, it is duplicated by Cuatrecasas 22188, from the

western Cordillera of El Valle, alt. 1950-2000 m., and his
21599 from the same general region, alt. 2250-2260 m. The
upper surface of tne leaf lacks white punctation, but is
sparsely marked with black dots which look like the mouth of
pits, but which are actually only saucer-shaped depressions.
The primary veins curve inward near the end of the leaf to
form a half circle, the blade is broadly rounded at the sum-

_ mit,-and the terminal apiculum is lacking or rudimentary.

Plants of the Zastern Cordillera, referred by me (1945)
to Be Andreana, are more or less white-punctate on the upper
leaf-surface, the blade is short-scuminate, and the primary
veins meet the midvein at a very acute angle. They may be
described as new.

BLAKEA ORIENTALIS sp. nov. Arbor 20 m. alta. Fetioli
crassi, 2--3 cm. longi, minute furfuracei. Laminae subcoria-
ceae, ellipticae, usque ad 18 cm. longee 9 cm. latae, abrup-
te breviterque acuminatee, basi late cuneatae, J-nerviae
jugo marginali neglecto, supra glabrae subnitentes sparse
albo-punctatae, subtus brunnescentes fere glabrae. Fedunculi
solitarii complanati glabri ca. 4 cm. longi. Bracteae fere
aequales, ad medium connatae, coriaceae, lete rotundatae,
glabrae, 2 cm. longae. Sepala bracteas 8 mm. excedentia,
semicirculearia, late rotundata, coriacea, rubescentia. Feta-
la rosea obovata subcoriacea 4 cm. longa. Antherse late
semiobovatae, 8 mm. longae; connectivo basi in calcar rectum
breve producto.

Type, Lawrance 153, in high Fieadt region of Mt. Chapon,
Boyacé, Colombia, alt. 2250 m. I also refer here tentatively
Killip & Smith 20197 from Norte de Santander, in which the
leaves are proportionately broader, more abruptly acuminate,
and very densely white-punctate above and the leaves and se-

pels much thinner in texture.

4, TOFOBEA REDUCTA sp. nov. Liana, ramis gracilibus juv-
enilibus, petiolis, et pedunculis tenuissime furfuraceis.
Folia valde dimorpha; petioli majorum usque ad 7 mm. longi;
laminae ellipticae chartaceae, 5--8 cm. longae, 2.5--3./7 cm.
latae, ceudato-acuminatae, basi obtusae, 5-pli-nerviae, ut-
rinque glabrae subnitentes; petioli minorum vix 1 mm. longi;
laminae ovatae, 3--& mm. longae. Flores solitarii, peduncu-

1947 Gleason, Blakea & Topobea 281

lis ca. 15 mm. longis. Bracteae ad basin distinctae, ca. 3
mm. longae. Hypanthium poculiforme, 2.5 mm. longum. Calycis
tubus ca. 0.5 mm. longus; lobi late rotundati ca. 1 mm. lon-
gi. Fetala ovata, 5 mm. longa. Antherae crasse subulatae, 2
mn. longae, basi tuberculo dorsali ornatae.

Type, Cuetrecasas 21082, from Rfo Calima, in the Choco
region of El Valle, Colombia, alt. 30--50 m. Since the spec-
imen exhibits only a single flower, no dissection has been
made and the dimensions stated above are approximate. The
anthers are clearly visible and leave no doubt of the gener-
ic position of the plant. Only three species with anisophyl-
lous leaves have hitherto been described. Of these T. glab-
rescens Tr. has sessile, cordate-clasping leaves; T. insig-
nis Tr, has much larger 5-nerved leaves, setose stem, and
bracts longer than the calyx. T. anisophylla Tr., to which
our plant is most closely related, has subsessile leaves
broadly rounded or subcordate at base, much longer peduncles
and bracts equaling the calyx.

NOTES ON SOME AMERICAN PLANTS

He Aw Gleason

Sida Elliottii and Sida inflexa.

Sida Elliottii is a well known species of the southeast-
ern states, represented in the larger herbaria by ample ser-
ies of specimens. Such manuals as Gray, seventh edition,
Britton & Brown, second edition, and Small give its range as
extending north to Virginia and Missouri. A recent collec-
tion of the Virginian plant by Fernald has led him to exem-
ine the species carefully and as a consequence to segregate
the plants of Virginia, Missouri, Tennessee, and one collec-
tion from Alabama as Sida inflexa Fern.

The differences between S. Elliottii and the proposed
species are stated by Fernald (Rhodora 40: 463, 464) as
shown below.

1. Se Es (a) Stems nearly glabrous, (b) 1.5--8 dm. tall.

O«
E. (a) Cauline leaves linear, (b) mostly 1.5--5 cm.
long, (c) 1.5--7 mm. wide.

i. (a) Foliis lanceolato- vel lineari-oblongis, (b)

2.5--6 cm. longis, (c) 0.4--2 om. latis.

Oo

282 PHYTOL OG Ta Vol. 2, noe 8

5. Se (a) Flowers mostly solitary in the axils and (b) on
pt fave up to 2.5 cm. long.
Se ie (a) Floribus plerumque corymbosis terminalibus, (b)
pedunculis ad 1./ cm. longis.
E. Calyx at most strigose on the ribs at base.
Calycibus basi plus minusve villoso-hirsutis.
. Carpels (a) with prominent erect teeth, (b) glab-
ous or nearly so on the beck.
S. i. Carpellis apice valde incurvatis, (b) dorse hispid-

Of these characters, impressive in their totality, no. 1
may be neglected as descriptive but not diagnostic, since
the dimensions of one are mostly included in those of the
other. Under no. 2, the actual dimensions overlap very much,
but the ratio of length to width seems derivable from the
stated figures at 7--10 times as long as wide in S. Elliott-
ii and 3--6 times in S. inflexa. Under no. 3, the ~ second
part of the statement refers only to maximum dimensions.

The first part, although purely qualitative refers to a con-
dition which can generally be recognized, although there is
no surety thet a corymbose inflorescence does not eventually
become axillary by elongation of the internodes. Numbers 4
and 5, especially the latter, appear to afford the best
diagnostic characters.

The Britton Herbarium contains forty sheets of S. Ell-
iottii in the broad pre-segregation sense, of which six : are
from Tennessee and Missouri and are referable by Fernald's
citations to S. inflexa. Also among the forty are three from
Alabama, from which state Fernald also cites S. inflexa. The
remaining thirty-one are from Florida and the Gulf Coast and
are referable to S. Slliottii according to the geographic
distribution stated by Fernald. These sheets have been care-
fully examined by me. Leaf-width has generally been measured
by eye-piece micrometer under a magnification of 10. Pubes-
cence of calyx and carpels has been observed under a binocu-
lar magnifying 235 times. Not every sheet illustrates all of
the characters stated by Fernald.

2. Ratio of length to width of leaf. Sixteen Gulf Coast
plants show the broadest leaves 3--6 times as long as wide,
thereby corresponding to the character of the more northern
S. inflexa. Fifteen show the broadest leaf seven times as
Tong as wide, or more, agreeing with Fernald's stetement for
S. Elliottii. The narrowest upper leaves, so far as observ-
ed, are 8--24 times as long as wide. In the Alabama plants,
the lower leaves are 2.4--35 times, the upper 12--15 times as
long as wide. In the Tennessee and Missouri plants, the low-
er are 3--6, the upper 8--12 times as long as wide, in the
latter features transcending the figures stated by Fernald

1947 Gleason, American Plants 283

Por Se inflexa.

3. Corymbose or axillary flowers. In 10 Gulf Coast sheets
the flowers may be described as corymbose, a character ad-
duced for S. inflexa. In 18 they are axillary, as stated for

Se Blliottii. In two Alabama sheets they are axillary; in

one they are corymbose. In six Tennessee and Missouri plants
(S. inflexa) they are all corymbose.

~ 4, Pubescence of the calyx. Fernald states that the calyx
of S. Elliottii is at most strigose on the ribs at base. No
strigose calyx was observed in all 40 sheets. leven Gulf
Coast plants (S. Slliottii) are distinctly villous with
soft, spreading, slender hairs on the midrib of each sepal,
as stated for S. inflexa; 19 are not villous. Of the six
Tennessee end Missouri plants (S. inflexa) all are villous
on the midribs. Fernald does not state thse pubescence of the
surface of the calyx. In two sheets from the Gulf Coast and
one from Alabama, it is glabrous; in two sheets from the

Gulf Coast and one from Tennessee (S. inflexa) it is nearly

glebrous; in all others it is distinctly but minutely stel-
late.

Sa. Direction of the beak. In 11 sheets from the Gulf
Coast (S. Elliottii) the beaks are erect; also in two from

Alebama, and in four (Se inflexa) from Tennessee and Missou-

ri. In three sheets, all from the Gulf Coast (S- Zlliottii),
they are incurved, but the incurving is apparently due to
the direction of pressure when the plants were dried. In the
other sheets mature carpels are not present or not easily
visible. |

Sb. Pubescence of the carpels, In every sheet where cer-
pels are exhibited the beaks of the carpel are minutely

hispidulous. Also in every sheet the back of the carpel be-

low the base of the beaks is glabrous.

In summary, carpels witn naturally incurved beaks and
hispid on the back do not exist in our specimens of the spe-
cies, even in the specimens cited by Fernald as S. inflexa.
Villous calyces exist in the specimens referable to S. in-
flexa but also in a third of the Gulf Coast plants. Corynm-
bose inflorescence exists in S. inflexa, but also in a
third of the Gulf Coast plants. In the Gulf Coast region
there is no correlation between villous celyces and corym-
bose inflorescence. The distinction between the two in pro-
portions of their lesves is largely fictitious.

In conclusion, I oan not recognize S. inflexa Fern. as a
species, nor even as a variety or form.

Sium suave.

Sium suave Walt. is a widespread species across the
northern states and adjacent Canada from the Atlentic to the

Gen.

284 PRE Eh: POG A Vol. 2, now 8

Pacific. When well grown in our northern wet sunny meadows,
or farther west in the open prairie "sloughs", it becomes a
great husky plant up to 2 meters tall or even more, with a
stem 3 om. in diameter neer the base and very prominently
and sharply angled. Under such favorable circumstances the
principal leaves may be 3 dm. long with as many as 17 leaf-
lets and these up to 17 cm. long and 5 cm. wide.

But the plant is exceedingly variable in stature, leaf-
let-shape, and dimensions. The leaflets may be linear and
only 2 mm. wide, the stem may be slender and weak, the um-
bels comparatively few. The umbels nevertheless retain their
usual appearance, except for an occasional reduction in the
number of primary rays, and the character of the rays, the
pedicels, and the fruit seems to be as uniform as one ex-
pects to find within a single species. Whether these varia-
tions are genetic and consequently heritable, or are caused
by « variable environment, such as amount of shade, depth of
water or the water table, or competition from surrounding
vegetation, is as yet unknown. Long familiarity with the
plant in the field has led me to believe that much of its
variability is caused by tne environment. I have no proof
thet such is the case; it is merely an impression based on
cumulative experience.

Sium Carsonii Durand has been recognized by some as a
species since its first publication in 1867 and is still so
considered by Mathias and Constance in 1945. It was reduced
to varietal rank by Stevens in 1910 end to a mere form of S.
suave by Fassett in 1921. Fassett's disposition, with which
I agree, was accepted by Fernald in 1943 (in print) or earl-
ier.

Sium floridanum was described by Small in 1933 in his us-
ual sketchy fashion and based on two specimens in the herb-
arium of The New York Botanical Garden, both from the Chip-
ola River in Florida. sxtracting his contrasting cnaracters
both from nis key and his descriptions and placing them to-
gether, we have the following, with the upper line of each
pair referring to S. suave, the lower to S. floridanum:

_l. Flant 6--19 dm. tall, stout.

1. Flant smaller and more slender.

2. Dileted petiole auriculate at the top.

2. Dilated petiole oblique at the top.

3. leaflets 11--17, linear to linear-lanceolate or rarely
wider.

3. leaflets 3--11, ovete to elliptic-lanceolate.

4, Leaflets saliently sharp-serrate or incised.

4, Leaflets finely appressed-serrate.

5. Umbel-rays slender.

5. Umbel-rays filiform.

1947 Gleason, American Plants 285

6. Larger corollas fully 2 mm. wide.
6. Larger corollas less than 2 mm. wide.

As to the significance of tnese differences, let us exam-
ine the type and compare it with northern plants always ac-
cepted as S. suave.

Pig. 1 leaf-margins

1. The type specimen and a duplicete of it are the tops
of plants. They do not indicate that the plant was smaller.
The thickness of the stems, in comparison with plents of S.
suave measured at the same distance from the sumnit, does
not indicate that the plants were significantly more slen-
der. Some plants of S. suave, as Blake 5456 from New Bruns-
wick, are much stouter; others, as Hartmann 215 from New
Hampshire, are equally slender. bet.

2. The petiole of the type of S. floridanum does taper
gradually to the summit. In S. suave it may taper gradually,
Or be abruptly narrowed, or very distinctly auriculate.

3. Tne largest leaves of the type heve only 9 leaflets.
On tne upper and uppermost leaves tnis number is progress-

286 PHYTOLOGIA Vol. 2, no. 8

ively reduced to 7, 5, 3, and 1. In S. suave there may be as
many as 17; other plants show, as far as the dried specimens
indicate, a lower maximum, often 11 or 9, in one instance 7,
while the leaves are again progressively Femnges upward to
5, 3, or even l.

4, The leaf-serration of S. suave varies veontla and ap-
parently without relation to any other structural feature.
Six leaf-margins of northern plants are shown in our Fig. l
(p. 285), all magnified 8 diameters. The type of S. florida-
num is very uniform in the character of its serration and a
typical centimeter is also shown in our figure. I trust that
the reader will try to decide for himself, on the basis of
Small's statements, which part of the figure represents S.
floridanum before turning to the clue in the last paragraph
below.

5. Small's statement needs more explicit statement before
it can be discussed.

6. Measurements of dry flowers have been made repeatedly,
selecting always those that were pressed open to exhibit
their full width, but making no allowance for the distention
caused by flattening of the hypanthium. The type of S. flor-
idanum veries from 18 to 23 units on my eye-piece microm-
eter, while S. suave varies from 18 to 26. This corresponds
to approximately 1./--2.2, or 1.7--2.4 mm.

Mathias and Constance add two characters:

7. Bracts 6--10, 3--15 mm. long.
7- Bracts 28-5, 2--5 mm. long.
8. Rays 10--20.

8. Rays 6--10.

In the type of S. floridanum the bracts on 3 umbels are 2,
5, and 5; their lengths are 27; 3.6, and 6.6 mm. The bracts
in S. suave are usually either 5 or 8, the larger number
holding for somewhat more then half the plants; umbels with 3
or 2 bracts are occasional and one bractless plant has been
seen.

Fernald in 1943 added another character, or rather defined
one suggested by Small:

9. Feduncle, primary rays, snd pedicels angulate.
9. Feduncle, primary rays, and pedicels filiform.

The statement concerning S. suave is correct as to ped-
uncle and primary rays. The pedicels may or may not appear
angled when dry, but every specimen which I have examined af-
ter boiling has been essentially terete. The type of S. flor-
idanum has the peduncle and primary rays just as strongly
angled as in inflorescences of similar size in S. suave. The

‘

1947 ; Gleason, American Plante. 287

pedicels also appear angled when dry.

It seems clear to me that there are no definite morpho-
logical features associated with S. floridanum which might
justify its maintenance as @ species. Its sole character is
thet of general frailty or debility in contrast with the
sturdiness and virility of the usual type and such a charac-
ter might easily be the result of a sheded environment on a
typically sun-loving plant. The same conclusion is suggested
by Fernald's statements thet its petioles are widely spread-
ing, while those of S. suave are ascending.

Tne source of the seven leaf-margins illustrated in Fig.
1 (p. 285) is: (a) Jones 12593, from Illinois; (b) Hartman
215 from New Hampshire; (c type of S. floridanum; (d) Fer-
nald & Wiegand 5954 from Newfoundland; (e) Deam 21292 from
Indiena; (f) Gleason & Gleason 167 from deen eed and (g)
Senn 1516 from Ontario.

Rhamnus lanceolatus.

Rhamnus lanceolatus Pursh is a well known species of the
Middle West, where it inhabits rich moist soil and is, in
general, the sole representative of the genus. From southern
Illinois soutnward its range overlaps with that of R. caro-
linienus Walt.

Fursh's description reeds:

R. inermis, arborescens; foliis lanceolatis serrulatis
utrinaue acutis subtus pubescentibus. On the side of hills:
Tennessee. Lyon. v.e S- in Herb. Lyon. Berries black.

Pursh had four other species in his Flora and it is note-
worthy that this is the only one without mention of floral
characters and the only one of our three native species
which he had not seen growing.

Twenty-four yesrs later Torrey and Gray knew this plent
directly from Kentucky end Missouri specimens and accepted
Fursh's statement thet it grew also in Tennessee. Their des-
cription must have been written primarily from the actual
material at hand, but tnrough deference to Pursh's "foliis..
--esubtus pubescentibus" they stated "more or less pubescent
beneath." At the same time and place they described as new
Re parvifolius, with pubescent leaves and tetrandrous flow-
ers, based on & Barton specimen from Harper's Ferry, W. Va.
The type of the species is at the New York Botanical Garden
and confirms what Torrey and Gray wrote in tneir appendix
two years later: "We have reason to suspect that this plant
is not distinct from R. lanceolatus." Other specimens a-
greeing with Barton's type occur from southern Fennsylvania
to Alabama. All have lesves densely pubescent at anthesis

288 PH YT OL OG:DA Vol. 2, noe 8

with apparently golden-brown hairs and remaining pubescent
at maturity.

West of the mountains, where the species is comparatively
common, the leaves may or may not be sparsely pubescent at
anthesis and are regularly glabrescent by maturity. Finding
no other character to distinguish the two populations, I
propose to recognize them as well marked geographical vari-
eties:

Rhamnus lanceoletus Pursh, var. lanceolatus. Speciei pears
typica, foliis molliter pubescentibus.

RHAMNUS LANCEOLATUS Pursh, var. GLABRATUS Gl., var. nov.
Foliis juventute glabris vel parce villosulis, maturitate
glabris. Rich moist woods at low elevations, west of the
mountains, Kentucky and Tennessee to Nebraska and Arkansas.
Type, Deam 787, Brookville, Franklin County, Indiana, in
Herb. N. Ye Bot. Gard.

Triadenum.

The few species of Triadenum, although segregated generi-
cally more than a century ago, have often been considered as
forming merely one section of the large genus Hypericum.
Britton adopted the genus Triadenum in the Illustrated Flora
and was followed by Small and Rydberg. The characters of the
genus are well known. The petals are imbricate rather than
convolute; the stamens ere only nine and are united into
three fascicles of three stamens each; these fascicles al-
ternate with three conspicuous hypogynous glands; the-petals
are pink, flesh-color, or greenish instead of yellow. It is
purely a matter of personal opinion whether these characters
are considered of sufficient importance to warrant the seg-
regation of a genus. In my pereonal opinion they are, and I
am accordingly discussing our American species under the
generic name Triadenum.

For many yeare our plants have been classified in two
species, T. petiolatum (Walt.) Britt. and T. virginicum (L.)
Raf.

As to the first of these, Fernald showed clearly in 1936
that it consisted of two populations, to one of which he
gave a varietal neme, maintaining the other as the typical
element of the species without a distinctive name. Walter
had described them both as species of Hypericum in 1788 but
unfortunately one of his names, H. petiolatum, was @ homonym
antedated by H. petiolatum L. The oldest valid name for this
species under the genus Hyperioum is H. Walteri Gmel. The
other of Walter's species, He tubulosum, has been described
in recent literature as T. longifolium | Small, over which the
specific epithet tubulosum has more than a century priority.

It is again purely a matter of personal opinion whether

\

1947 Gleason, American Plants 289

these two populations are considered as two species, or as
two varieties of one species, or as a single species. In my
opinion they are species. Fernald has pointed out correctly
the difference in the leaves. The sepals of tubulosum aver-
age about 1 mm. longer and are almost always acute. The
leaves of tubulosum have no superficial glands and also lack
the translucent glands found in the other species.

While valid names for both are available in Hypericum,
new combinations are necessary when the plants are placed
in Triadenum.

TRIADENUM WALTERI (Gmel.) comb. nov. Hypericon Walteri
Gmel. Syst. Nat. 2: 1159. 1791. Hypericum petiolatum Walt.
Fl. Car. 191. 1788; not H. petiolatum L. 1762. H. tubuloeum
Walt. var. Walteri Lott, Jour. Arnold Arb. 19: 279. 1936.
Triadenum petiolatum Britt. Ill. Fl. 2: 437. 1897.

TRIADENUM TUBULOSUM (Walt.) comb. nov. Hypericum tubulos-
um Walt. Fl. Car. 191. 1788. T. longifolium Small, Bull.
Torrey Club 25: 140. 1898. H. petiolatum var. tubulosum
Fern. Rhodora 38: 436. 1936. H. Walteri var. tubulosum Lott,
Jour. Arnold Arb. 19: 151. 1936. |

Fernald has also indicated the differences between the
northern and southern forms of T. virginicum. These differ-
ences are so clear-cut that Spach separated the plants spec-
ifically a century ago and one can only wonder why they were
neglected by all (so far as I know) American botanists for
an even hundred years. On the basis of these differences
Fernald proposed to distinguish the plants varietally, but
here again I believe that he did not go far enough and that
we shall do better to treat them as species. Again a new
combination is necessary.

TRIADENUM FRASERI (Spach) comb. nov. ericum Fraseri
Spach, Ann. Scie Nat. Bot. II. 5: 168. ea H. virginicum
var. Fraseri Fern. Rhodora 38: 434, 1936.

I have measured 155 fruits of the aggregate species un-
der an eyepiece micrometer, using a magnification which per-
mitted accuracy to the tenth of a millimeter. The sepals of
the northern T. Fraseri vary from 2.8 to 4.9 mm. long,
measured from the sinus, with an average length of 3.7/7 m.
Furthermore theee sepals are always obtuse and usually ac-
tually rounded at the summit. Also, they aré wider at the
middle than at the base and consequently commonly appear
elliptic or spatulate in general outline. The sepals of the
southern T. virginicum vary from 4.3 to 8.4 mm. long and av-
erage 5.86 mm. ; only one plant exhibited sepals less than 5
mm. long. These sepals taper toward the summit, which is
sometimes acuminate, more commonly acute, and rarely narrow-
ly obtuse, Because of the long terminal taper they appear
lanceolate or oblong-lanceolate in outline.

The styles of the northern T. Fraseri, as they persist on

290

Length of styles

2.6-35.0

{ 4

RA -Y“T Oub O GTA

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aaa
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+ wo

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Ft on 868 AF 9

il

ad
oN WM Fk
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a
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B98 SB Bh BEL Bien, os OR cae ie Vine @
ory ty Oo NM 8
i aoe aoe? Oy as Bee Beef
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6.6-7.0

Vol. 2% noe §

7.1-7.5
7.6-8.0
8.1-8.5

1947 | Gleason, American Flants 291

the fruit, vary from 0.6 to 1.5 mm. long and average 1.06
mm. Measurements were made only on the fruit, since they are
rarely completely visible in flowers without boiling and
dissecting. The styles of T. virginicum vary from 1.9 mm.
(two instances) to 3.5 mm. and average 2.72 mm.

When the dimensions of the sepals and styles are plotted
together [see the chart on page 290], they show that the
aggregate is composed of two completely separate popula- |
tions.

The distribution of the aggregate is in some waye similar
to that of Sarracenia purpurea, which consists of a southern
population chiefly confined to the coastal plain and a nor+
thern one extending far inland. T. virginicum (L.) Raf. is -
essentially a plant of the coastal plain from Nova Scotia to
the Gulf coast, but extends inland across New York into
southern Ontario and reappears in northern Indiana. T. Fre-
seri (Spach) Gl. is essentially a boreal plant, extending
from Newfoundland and Labrador to Manitoba and southward to
Connecticut, New York, northern Indiana, end Nebraska, or at
higher altitudes to West Virginia.

= = 22 2 2 2 2 2 2 2& 2 2 2 5s 2 2 2 ses 2 @&

A LETTER FROM PERDINAND VON MUELLER

He Ae Gleason

There recently came into my possession a hand-written
letter from the noted Australian botanist, Sir Ferdinand Ja-
kob Heinrich von Mueller, addressed to B. Daydon Jackson
(1), which has some biographical and bibliographical inter-
est. Von Mueller was born in Germany in 1825, emigrated at
an early age to Australia, became a British subject, devoted
his life to a study of the flora of Australia, was govern-
ment botanist for forty-four years and director of the Mel-
bourne Botanic Garden for sixteen years, was created Baron ~
by the King of Wiirtemberg in 1871, was knighted by Queen
Victoria, and died in 1896. His career therefore shows some
parallelism with those of Sir Richard and Sir Robert Schom-
burgk, who were also born in Germany. The letter reads:

21.11.83.

Herewith, dear Mr. Jackson, I beg to send you a copy of
part of a letter from Dr. Fournier (2), just received, con-
cerning the priority of Vahea over Landolphia, as this ques-
tion will interest you not only for these genera but in many
other respects for your nomenclator (3). I aleo forward a
copy of the important prospectus, found by Dr. Fournier, as

\

292 PHY TO:L.OGEA Vol. 2, no. 8

this renders it conclusive, how far the “illustration des
genres" had actually appeared up to 1796. Ae you and Mr.
Britten (4) took such an interest in clearing up the dates
(5) of Rees' Cyclopaedia (6), you likely will give some at-
tention to this question now.

I intended to have written a letter on the subject in
Trimen's (7) journal or rather now Britten's journal: of Bot-
any (8), but I have been suffering for fully two months from
so severe a bronchial catarrh, that I had to keep to my
rooms all that time; the cough has slightly abated now, but
as there is some emphysema, it is doubtful whether I will
recover. My parents both died early on phthisis, and heredi-
tary inclination to that fatal disease brought me out to
‘this warmer clime so soon as I left the University. In the
wet season it is however even here too cold for me. I am
just going up into a mild forest region with the hope of
benefitting from the moist equable air there. I feel very
weak. Am very sorry to hear of Mr. Bentham's failing
strength, but hope, that after some rest he will still be
able to resume his grand phytographic labours.

Regardfully your
Ferd. von Mueller.

Only a part of Fournier's letter was copied and sent to
Jackson. It reads:

Paris, le 24 Septembre 1883.
Tres honoré Monsieur-

Je reponds d'abord a la partie de votre lettre qui con-
cerne 1'@poque de publication de la partie botanique de
1l'Eneyelopedie (10). J'ai correspondu déja sur ce sujet
pour vous avec rn de Candolle (11), qui a du vous transmet-
tre ma réponse. De plus, en rendant compte dans le Bulletin
de le Societe de France t. XXIX, Revue, C p. 137 (1882), de
votre Literary Reference to the Caoutchoue Vaheas, j'ai
tranecrit le texte de Millin, qui prouve que la genre Vahea
a 6té publié anterieurement a 1797.

Aujourd'hui je me trouve en possession de nouveaux ren-~
seignements, grace a@ un prospectus publié le es Nov. 1796
par 1'éditeur de 1'Encyclopedie, prospectus qu ‘un heureux
hasard & fait entrer dans ma bibliotheque. Je vous en ad-
resse ci-joint un extrait.

Publicatior
de la soixantieme livraison de 1'Encyolopédie
a@ Paris, rue des Poitevins: No. 18.
le l-er Frimaire, an cinquieme de la Republique frangaise
(le lundi 21 Novembre 1796, vieux style)

Cette livraison est composée

1° de la dix septieme des planches d'histoire naturelle,
formant la septieme centurie de celles de la botanique, par
le citoyen Lamarck (12), de l'Institut national, professeur

1947 Gleason, Letter from von Mueller 293

et administrateur du Muséum d'Histoire Naturelle.

Le volume de planches, qui fait partie de cette livrai-
son, est le septieme centurie de celles de la _botenique, et
presente la continuation de 1'ouvrage intitulé: Illustration
des genres. Ces planches sont gravees avec le plus grand
goin, et la plupart des figures sont originales.

Elles offrent, depuis la planche 601 juequ'é la 700° in-
clusivement, 164 genres, parmi lesquels il s'en trouve quan-
tité de nouveaux. . oe Les genres exposés dans cette liv-
reison appartiennent a le Diadelphie entiere, & @ la Polyadel-
phie et & la plus grande partie de las Syngénesie. . . « «
Dens la soixante-unieme livraison nous publierone le prem-
fere partie du tome IV® du Dictionnaire de botanique, par le
citoyen Lamarck.

The letter, with its accompanying correspondence, there-
fore becomes of bibliographic value, indicating definitely
that the publication of plates 601--700 of the Illustration
des Genres actually appeared in 1796. The accepted date of
publ ication has heretofore been 1797.

One can only guess the history of the letter since 1883,
or the early part of 1884, when it reached Mr. Jackson.
Probably it was mislaid among some herbarium specimens, sent
to America with duplicates, lay unobserved for more than
forty years, and finally came to light among some old papers
at the Biological Station of the University of Michigan.

Footnotes

(1) B. Daydon Jackson (1846--1927), editor of the Index Kew-
ensis.

(2) Bugene Pierre Nicolas Fournier (1834--1884).

(3) = Kewensis, 1893--1895, with supplements 1--8, 1901-
1934.

(4) James Britten (1846--1924), editor of Journal of Botany
1880-- l 924 °

(5) Jour. Bot. 15: 107, 108. 1877; 18: 87, 88. 1880.

(6) Abraham Rees, The cyclopaedia; or universal dictionary
of arts, sciences, and literature. London, 1806--1820.

(7) Henry Trimen (1843--1896), editor of Journal of Botany
1871--1879.

294 PHYTOLOGIA Vol. 2, no. 8

(8) Journal of Botany, founded 1863, now in its eightieth
volume.

(9) George Bentham (1800--1884).

(10) Encyclopédie méthodique. Botanique par M. le Chevalier
de Lamarck. Paris, 1783--1817.

(11) ‘i phonse Louis Pierre Pyramus de Candolle (1806--1893).

(12) Sead Baptiste Antoine Pierre Monnet de Lemarck (1744--
1829).

NEW OR NOTEWORTHY MELASTOMES, CHIEFLY ECUADOREAN

H. Ae Gleason

There is in South America a group of seven species within
the genus Meriania which have a conspicuous habital similar-
ity. This is due, on superficial examination, to their broad
5-nerved leaves, usually elliptical in shape, their inflor-
escence with well developed central axis, and the densely
pubescent, cinereous or ferruginous hypanthium. Closer exam-
ination reveals other features of similarity of a more tech-
nical nature. The hairs of the hypanthium are wholly or
partly enlarged at base and there roughened or stellate,
while the terminal portion is slender, smooth, and curved-
ascending. The hairs of the lower leaf-surface are conspicu-
ously or minutely stellate at base, with an erect simple
bristle.

In the four species which inhabit Venezuela and Colombia
we find also a strong similarity in the stamens (Fig. 1).
The anthers are flattened tangentially; the thecae are
scarcely in contact, so that the connective is visible from
the anterior side. On the posterior side the connective is
not elevated; it is prolonged straight back. The Feruvian
species has a connective which appears quite different, but
which can be readily homologized with the more northern spe-
cies. In the Bolivian species we find a real divergence.
Here the connective is prolonged into a large basal spur and
a smaller dorsal spur. This plant, M. boliviensis Cogn., was
originally assigned by its author to © the section Bumeriania,
characterized by unappendaged anthers and solitary flowers.
Later, in his monograph of the family, he placed it in sec-
tion Umbellatae, with paniculate flowers but with unappen-
daged anthers. Apparently Cognieaux never made a dissection

1947

Fig. l

Gleason, New or Noteworthy Melastomes 295

A.
Be
Ce
De

a erie
igs

Lateral views of stemens, X 5
Meriania cuneifolia, isomorphic
M. colombiana, isomorphic
M. boliviensis, dimorphic
ouintuplinervis, dimorphic

Fl

296 PHYTOLOGIA Vol. 2, nog!

of the flowers.

It is not my present purpose to attempt to decide shethay
the division of Merienia into five sections is valid, or
whether the structure of the anthers should take precedence
over habit and pubescence in determining intrageneric rela-
tionships. I merely point out that there are seven species
which resemble each other strongly in these latter features.
Since these features are so patent, the group will be recog-
nized by any one trying to identify these species or any
still undescribed species of similar nature. The seven may
be separated by the following keys, the first based wholly
on vegetative characters, the second utilizing the structure
of the stamens.

Leaves 5-nerved, conspicuously dentate.
Pubescence cinereous, that of the veins on the lower leaf-
surface no longer than that of the surface.
Flowers NUMETOUBs ceovccccccccccsceeMe Weberbaueri Macbr.
Flowers apparently SOLITATY. scucsscvesecsle loxensis Gl.
Pubescence ferruginous, that of the veins on the lower
leaf-surface much longer and coarser than that of the
BUITECOcscceccccccscccccccces oe Steye rmarkii Gl. ined.
Leaves S-pli-nerved, entire or very obscurely denticulate.
Leaves subacute to rounded at base, broadest at or near
the middle.
Exterior teeth scarcely projecting beyond the sepals....
coc ccccccccccsccccscccccvccoeMe Guintuplinervis Naud.
Exterior teeth projecting 2--3 mm. beyond the sepals.
Pedicels 15--20 mm. long; leaves softly subtomentose
beneath, ACULC ce cceececcccccccccccle colombiana Gl.
Pedicels 5--8 mm. long; leaves very sparsely pubescent
beneath, short-acuminate.....+eM. boliviensis Cogn.
Leaves long-cuneate at base, broadest much above the
Middl@sescsccevccrccscccrcccvccsessceeMe cuneifolia Gl.

Connective below the tnecae terete or channeled on the lower
(anterior) side, the dorsal spur minute or lacking.
Filament attached to the very base of the connective.
Leaves obtuse or rounded at base..ceceseeeMe colombiana.
Leaves cuneate at Daa icceiancesnibier cide cuneifolie.
Filament attached near the middle of the connective.
Exterior teeth scarcely projecting beyond the sepals;
leaves S-pli-nervedeseccscccccsceeeoMe Quintuplinervis.
Exterior teeth conspicuously projecting; leaves 5S-nerved
CeCe rerrrccccccccccscsresessccccesoMe Steye rmarkii.
Connective elevated into a very flat, triangular or 2-lobed,
basal spur.
Leaves S-nerved, conspicuously dentate; connective not
lobed.

1947 Gleason, New or Noteworthy Melastomes 297

:

m

lateral views of stamens, X 5
A. Meriania pallida, dimorphic

Be Me macrophylla, dimorphic

C. M. Steyermarkii, isomorphic
D. Me loxensis, isomorphic
B. Me Weberbaueri, dimorphic Fig. 2

. ' ~
,
j ‘ a

‘

298 PHYTOLOGIA Vol. 2, no. 8

Spur of the connective extending well forward on the
back of the anthereceeseecccscscccecseeMe Weberbaueri.
Spur entirely below the base of the anther. Me loxensis.
leaves S-pli-nerved; connective of the larger stamens 2-
LobedecccccecccccccsccccsevessesoveseceseMe boliviensis.

The mention of M. Steyermarkii at this place is under no
circumstances to be construed as publication, which will be
effected elsewhere; no Latin diagnosis is here provided.

MSRIANIA CUNEIFOLIA sp. nov. Sect. Umbellatae. Caules,
petioli, folia subtus, paniculae, et hypanthia dense pubes-
centes, pilis basi incrassatis barbellatis, apice simplici-
bus. Dentes exteriores calycis ultra sepala producti. Stam-
ina satis dimorpha; antherae complanatae; connectivum infra
thecas rectum, dorse minute calcaratum, basi ima «d filamen-
tum affixum.

Shrub 1.5 m. tall. Fanicle, hypanthia, stems, and lower
leaf-surface softly cinereous, the hairs slender, smooth,
and curved-ascending above an enlarged, roughened or stel-
late base. Petiole 2--3 cm. long. Blades oblanceolate, up to
15 cm. long and 4 cm. wide, abruptly and sharply acuminete,
entire, long-cuneate at base. Panicle terminal, the 5S-merous
flowers in subumbellate terminal clusters on pedicels 4--8
mm. long. Hypanthium campanulate, 5.2 mm. long to the torus.
Calyx irregularly ruptured to the torus, the lobes 8.5 mm.
long, pubescent like the hypanthium, slightly thickened e-
long the median line but with no developed exterior teeth.
Petals "buff-salmon", rotund, 13 mm. long. Stamens dimorph-
ic; filaments 6.4 or 8.5 mm. long, flat, becoming concave at
the summit; thecae 8.3 or 5.3 mm. long, strongly flattened
tangentially; connective prolonged straight back 2.5 mm.,
channeled on the lower side, affixed to the filament at its
very base, bearing a large or very small, broadly conic, ob-
tuse or rounded basal dorsal spur. Style straight, 21 mm.
long; stigma truncate.

Frov. Santiago-Zamora, Ecuador, dense forest between
Campanas and Arenillas, altitude 2195 meters, Steyermark
53543. The species is further contrasted with its apparent
relatives in the preceding paragraphs.

MSRIANIA LOXENSIS sp. nov. Sect. Umbellatae. Caules, pet-
ioli, folie ad nervos subtus, et hypanthia dense sed tenui-
ter pubescentes, pilis basi incrassatis barbellatis, apice
simplicibus. Dentes exteriores calycis ultra sepala bene
producta. Stamine isomorpha; antherae complanatae; connecti-
vum infra thecam in calcar dorsalem tuberculatum elevatum.

Shrub 3\m. tall. Stems, veins of the lower leaf-surface,
petioles, and hypanthia thinly cinereous, the hsirs slender,

/

\

1947 Gleason, New or Noteworthy Melastomes 299

smooth, and curved-ascending above an enlarged roughened or
minutely stellate base. Petioles 10--15 mm. long. Bledes
firm, elliptic, up to 10 cm. long and 5 cm. wide, acute,
denticulate in the distal half, rounded at base, very ob-
scurely 5-pli-nerved, densely stellate-furfuraceous on the
veins beneath, on the surface very minutely and sparsely
stellate, a few of the hairs ending in a very short erect
bristle. Flowers 5-merous, apparently solitary, on a pedicel
8 mm. long. Hypanthium campanulate, 8 mm. long to the torus,
very thick-walled. Calyx-tube prolonged 1--1.5 mm.; sepals
broadly ovate, thin, 4.5 mm. long, acute; exterior teeth ad-
nate nearly to the summit of the sepals, projecting 3.5--5
mm. Petals “deep salmon-vermillion", obovate, 27 mm. long.
Stamens isomorphic; filaments strongly flattened; anthers
subulate, tangentially flettened, 9.4 mm. long; connective
prolonged down the back as a sharp narrow ridge, greatly di-
lated immediately below the thecae, and below the summit of
the filament prolonged 3.3 mm. into a flattened or subconic
obtuse organ strongly tuberculate toward the tip. Ovary sup-
erior, 10-costete; stigme truncate.

Frov. Loja, Scuador, Sotobosque, between Tambo Cachiyacu,
La Entrada, and Nudo de Sabanillas, Steyermark 54468.

MERIANIA PALLIDA sp. nov. Sect. Fachymeriae. Arbor 20 m.
alta, ramis juvenilibus 4-angulatis pallide furfuraceo-
puberulis. Fetioli usque ad 6 cm. longi, scuto dorsali orna-
ti. Laminae firmae, ellipticae, usque 29 cm. longae 14 cm.
latae, obtusae, integrae, basi rotundatee, 3-nerviae, jugo
conspicuo marginali neglecto, supra glabrae opacae, subtus
griseae arcte stellato-tomentosulae; venae secondariae supra
planae, subtus elevatae, 5--8 mm. dissitae, sub angulo 80°
divergentes. Fanicula ca. 1 dm. longa; rachis compressa pul-
verulenta; pedicelli 5-8 mm. longi. Hypanthium late poculi-
forme, ad torum 5 mm. longum, primum sparse griseo-stellula-
tum, mox glabrescens. Calycis tubus 1.5 mm. longus, truncat-
us; dentes exteriores minuti, ca. 0.1 mm. longi. Petala ros-
ea, ca. 15 mm. longa, inequilatera, late oblonga. Filamenta
glabra torta, basi iata, ad apicem angustata, 7 vel 7.5 mm.
longa. Antherae subulatae, 9 vel 8 mm. longae, poro dorso-
termineli dehiscentes. Connectivum ad dorsum antherse basin
versus oe infra antheram in calcar assur-
gentem 4 vel 3.4 mm. longum productum; appendix dorsalis in
ser. ext. ad basin connectivi, subulata, 6 mm. longa, apice
bifurcata; in ser. int. ultra medium connectivi, subulata,
5-5 mm. longa.

Type, Cuatrecasas 15567, from Dept. del Valle, Colombia,
G&rdillera Occidental, vartiente occidental, Hoya del rfo
Sanjuniquin, lado izquierdo, La Laguna, bosques, 125C--1400
me. alt., described as "Arbol 20 m., 30 om. didm.; hoja cori-

300 PHYTOLOGIA 5 Vol. 2, now 8

dcea, gruesa, fragil verde esmeralde en el haz, palida; cen-
icientto blenquecina en e] enves; pétalos caérdeno vivo muy
brillantes; caliz verde o purpureo; corteza grisdceo amaril-
lenta palida; madero amarilla."

The plant was originally identified by me as M. macro-
phylla (Benth.) Triana. While certainly closely related to
that species, it differs in such important respects that its
recognition as a species is necessary (see Fig 2, p. 297).

In M. macrophylla the two spurs of the connective are of ap-

proximately the same size, the leaves are shorter and more
ovate, ferruginous rather than cinereous; in M. pallida the
two spurs are very unequal, and the subulate anterior spur
is bifurcate in the lerger stamens, the leaves are elongate
and elliptic and distinctly cinereous.

CALYPTRELLA DENTICULATA sp. nov. Folia elliptica vel obo-
vato-elliptica, utrinque acuminata, 5-nervia. Flores longe
pedicellata, S-meri. Calyx ante anthesin apice S-dentatus,
ad anthesin non circumscissus, irregulariter ruptus in lobos
3--5 triangulares. Antherse 5.5 mm. longae. Stylus 17 m.
longus.

2 ae

Fig. 3 Calyptrella denticulata, stamens X 6

Shrub up to 4.5 m. tall, the young stems, petioles, pan-
icle, hypanthium, and lower leaf-surface stellate with minu-
te hairs about O.1l mm. across. Leaves elliptic or obovate-
elliptic, up to 19 om. long and 8 cm. wide, abruptly short-
acuminate, entire, tapering to the base, S-nerved or weakly
S-pli-nerved with an additional pair of marginal veins,
glabrous above, soon glabrescent beneath except for a little
persistent stellate pubescence along the nerves. Panicle
terminal, 3--6 cm. long, many-flowered, its branches tending
to nod. Hypanthium cup-shaped, about 4 mm. long to the tor-
us, firm-walled, thinly stellate. Sepals in bud closely con-
nate to the summit, where the minute exterior teeth project
slightly, at anthesis irregularly ruptured into 3--5 broadly
triangular lobes with convex side, the tube about 1 m.
long, the lobes about 2 mm. long. Fetals obliquely subro-
tund, 9 mm. long, 10 mm. wide. Filaments flattened, 5.6 mm.
long, opening by a minute pore; connective extending along

1947 Gleason, New or Noteworthy Melastomes 301

the thecae as a slender sharp ridge, below the thecae great-
ly thickened and prolonged 1.5 mm., bearing a thick dorso-
basal spur. Ovary nearly free, S-celled; style slender, 17
mm. long; stigma punctiform.

Prov. El Oro, Ecuador, forested slopes between Pampa de
los Cedros, northeast of San Pablo, and Curtincapa, alti-
tude 2285--2430 meters, Steyermark 53809; his number 54167,
also from Prov. El Oro, is the same. Number 52781, collected
a short distance to the north in Frov. Azuay, shows no point
of difference in the flower, but the leaves are conspicuous-
ly S-pli-nerved, the inner pair of veins arising about 15
mm. above the base of the leaf.

This is the eighth species of Calyptrella to be describ-
ed. (An unpublished name under this genus is attached to He
H. Smith 3, found in many herbaria; the plant does not be-
long to this genus or even to this tribe of the family.) The
eight may be distinguished by the following key.

Fetals ovate to lanceolate, acute or acuminate.
Flowers 6-merous; panicles 1--3 dm. long.
Pedicels 2--6 mm. long; Ecuador to Bolividecccsccccccees
ec vccccccscccccccsvecesseeGe Cucullata (Don) Triana.
Fedicels obsolete, or less than 1 mm. long.
Leaves about half as wide as long; MexicOscecccceccecs
C. Galeottii Naud.
Leaves about three-fourths as wide as long, or wider;
Costa Rica, Colombia.....sC. cycliophylla Donn. Sm.
Flowers 4-merous.
Panicle 2 dm. long; leaves rounded at base, 7-nerved,
stellate-puberulent beneath; Peru....C. robusta Cogn.
Panicle 5--10 cm. long; leaves acute or obtuse at base,
3-nerved (excluding the marginals).
Teaves coriaceous, minutsly lepidote beneath; Feru....
ce ceercceccvcoccccccccvescccoseveece tristis Triana.
Leaves thin, glabrous beneath; Feruicesecsccccccesccces
Co oe ee rerccresececsesereseceeeeele gracilis Triana.
Petals obovate to subrotund.
Flowers 4-merous; petals 4 mm. long; leaves 3-nerved,
rounded at base; Colombia....ssseeeeeeCe littoralis Gl.
Flowers S-merous; petals 9 mm. long; leaves S-nerved, nar-
rowed to the base; Ecuador..c.eeceeeeCe denticulata Gl.

sf

MICONIA ZAMORENSIS sp. nov. Sect. Amblyarrhena. Panicula
cum hypanthio longe glanduloso-hirsuta. Sepala patula, obo-
vata, dentibus exterioribus subuletis. Petala late rotunda-
to-obcordata. Ovarium setis ca. 10 glanduliferis coronatum;
stylus tenuissime villosulus; stigma peltatum.

Stem, petioles, and branches of the panicle freely hir-
sute with slender spreading hairs 2--3 mm. long, those of

~

502 -PHYTOLOGIA Vol. 2, no. 8

the panicle mostly gland-tipped, those of the petioles most-
ly simple, those of mature stems entirely simple. Fetioles a
1.5--3.5 cm. long. Blades thin, elliptic-oblong, up to 12.5 |
by 6.5 om., acuminate, minutely serrulate, rounded or broad-

Fig. 4 Miconia zamorensis, style and stamens X 10

ly obtuse at base, S-nerved or weakly S-pli-nerved, hirsute
with yellowish hairs 2--2.5 mm. long, those of the upper
side avoiding the veins, those of the lower side on the
veins onlye Panicle about 1 dm. long, including the long pe-
duncle, loosely branched and few-flowered; actual pedicels
only 0.5 mm. long. Flowers S-merous. Hypanthium broadly cup-
shaped, 2 mm. long to the torus. Sepals round-obovate, 1.6
mm. long from the sinuses, much exceeding the subulate ex-
terior teeth. Fetals 2./ mm. long, 3.35 mm. wide. Stamens
isomorphic; filaments flat, gradually tapering from a wide
base, glabrous; anthers oblong, 4-celled, 2.4 mm. long, o-
pening by a ventro-terminal pore; connective simple. Ovary
inferior, crowned by about 10 erect glandular setae; style
(immature) 4 mm. long, obscurely villosulous; stigma pelt-
ate, not angled, 1.1 mm. in diameter.

"Shrub 5 feet tall; petals white; filaments white; anth-
ers yellow; calyx greenish-white; pedicels and peduncle pale
salmon; leaves membranous, shining and deep green above,
pale green below." Prov. Santiago-Zeamora: high wooded slopes
above Valladolid, altitude 2100--2400 meters, Steyermark
54701. Among the 141 described species of this section, the
great majority of which are represented in the herbarium of
the New York Botanical Garden by authentic specimens, de-
tailed drawings, or notes, not one has similarly glandular-
hirsute pubescence. In foliage and especially in inflores-

1947. ' Gleason, New or Noteworthy Melastomes 303

cence, M. zamorensis resembles M. Killipii Gl. of Colombia,
and M. megastigma astigma Gl. of Ecuador. Both of these have glandu-
lar filaments and styles and anthers of entirely different
shape.

MICONIA BARBIPILIS sp. nov. Sect. Amblyarrhena. Folia ov-
ata, supra bullata asperrima, subtus, sicut caulis, rachis,
et hypanthium, pilis conicis basi dense barbatis obtecta.
Filamenta stylusque sparse glanduloso-puberula. Stigma late
pelteatum S-angulatum.

A shrub 3 meters tall. Stem stoutly 4-engled, densely
‘ferruginous with stoutly conic or nearly ovoid hairs barbel-
late at base, slender above. Fetioles similarly pubescent,
35--7 cm. long. Blades ovate, up to 25 cm. long and 15 cm.
wide, subacuminete, broadly rounded at base, 7-nerved; upper
surface bullate, the principal bullae terminated by a conic
ascending hair about 0.5 m. long; lower side foveolate, the
veins all marked by a row of barbellete hairs like those of
the stem but shorter. Fenicle 15 cm. long, sparsely branch-
ed, pubescent like the stem. Flowers 5S-merous, sessile, sub-
tended by ovate brects 3.5--4 mm. long. Hypenthium cup-
shaped, thick-walled, about 5 mm. long to the torus, densely
beset with ovoid ascending hairs about 0.5 mm. long and bar-.
bellate at the base. Calyx-tube prolonged about 0.8 mm.; se-
pels semicircular, thin, about 0.9 mm. long above the sinus-
es; exterior teeth continuous, pubescent like the hypanthium
but more sparsely, terminating in a very short conic projec-
tion. Petals white, obliquely obovate, about 5 mm. long and
nearly as wide. Stamens isomorphic; filaments broad and
flat, sparsely and minutely glandular-puberulent; anthers
oblong, 4-celled, 3.3 mm. long, opening by a minute ventro-
terminal pore; connective simple. Ovary inferior, apparently
S-celled; style columnar, at least 5 mm. long, densely
glandular-puberulent; stigma peltate, S-angled, 2.1 mm.wide.

"Shrub 10 feet tall; petals white; calyx dull olive-
green; leaves deeply and finely rugose both sides, dull
buff-green below, dark green above; anthers yellow." Prov-
ince Santiago-Zamora, trail between Failas and El Pan, alti-
tude 2255--3445 meters, Steyermark 54308.

In Cogniaux' monograph there is a group of twelve species
described as "folia supra appendicis crassis conicis vel py-
remidatis strigosa" or "folia supra bullis setiferis pustu-
lata." Our plant is related to these species and to the four
recently described members of the same group, M. frontinoana
Gl., M. trichrona Macbr., M. Fennellii Gl., and | M. pseudo-
radula | Cogn. & Gl. Among these M. barbipilis is the only
species with barbellate pubescence, as described above.

There is in the Andes of Ecuador and Colombia a small
group of species in the section Amblyarrhena of the vast ge-

Se

304 PHYTOLOGIA’ Vol. 2, no. 8

us Miconia which not only have the same general aspect, as

seen mounted on herbarium sheets, but also agree in certain :

points of structure. They probably constitute a distinct
species-group. At least four of them seem to be apparently
low plants, almost herbaceous of stem, freely and diffusely
branched. The other two are variously advertised as shrubs,
low trees, or trees, ususlly with no statement of height,
although one specimen is designated as a tree four feet
tall. When dry, all species have a dull green or bluish
green cast. The leaves are thin, ovate, and prominently re-
‘ticulate on the lower surface. The petals are broadly obo-
vate, slightly retuse, and nearly equilateral; the flat fil-
aments are not geniculate and taper uniformly from a broad
base to a narrow summit. The plump anthers tend to be
slightly obovate; they are essentially isomorphic, but in
the epipetalous series the connective narrows toward the
base, while in the episepalous series it broadens and is ob-
scurely bilobed; in all but one species it is prolonged
briefly below the thecae into an inconspicuous dorsal lobe.
The filaments and style are glabrous; the stigme is capit-
ate. The six species may be separated by the following brief
key.

Pubescence of the hypanthium and panicle stellate, either
wholly or with simple hairs also....M. peychrophila Naud.
Pubescence of the hypanthium and panicle entirely of un-
branched hairs.
Pubescence entirely of long spreading unbranched hairs.
Hairs partly or chiefly gland-tipped.
Exterior teeth thick and rounded, not surpassing the
BOPAlLSecccscccesevesccccccsceeMe caesia Cogne & Gle
Exterior teeth subulate, much longer than the sepals..
ecccersccccesccsccscccccccceMe nigripes Cogn. & Gl.
Hairs all simple.
Leaves plane; exterior teeth not projecting beyond the
sepals; flowers S-merouSesecccceceseMe subalpina Gl.
Leaves bullate; exterior teeth projecting; flowers 4-
MOTOUBsccccccccccccccsccvscccele acalephoides Naud.
Pubescence of minute incurved hairs; flowers erage Fi
cece c coer cscescccccesccccscsccsccccccscccoMe inna ta Gl.

M. scabriuscula Cogn., a Bolivian species which I have
not seen, was stated by the euthor to be related to M. aca-
lephoides. It is said to have bullate leaves and a long-set~
ose calyx, as in that species, but 5-merous flowers. The ex-
terior teeth were not mentioned by RognAayns

MICONIA INNATA ep. nov. Sect. Amblyarrhena. Frutex 6 dm.
altus, cauli cum petiolo pubescente, pilis flexuosis usque

1947 Gleason, New or Noteworthy Melastomes 305

‘ad 1 mm. longis. Petioli 1--2 cm. longi. Laminae tenues, o-
vatae, opace virides subtus pallidiores, obtusae, irregular-
iter crenulatae, basi rotundatae vel subcordatae, 5-nerviae
vel fere 5S-pli-
nervise, supra fere

glabree, subtus ad
venas sicut cauli
pubescentes. Panicu-
la pyramidalis 5--6
cme longa, minute
pubescens, pilis in-

curvis 0.2 mm. long-

is. Flores 4-meri.
Hypanthium tubulos-
um, ad torum 2 mn.

longum, sicut panic-
ula pubescens. Caly-
cis tubus O.2 mm.
productus; sepala
trieangularia obtusa,
a sinibus 0.7 me
longa; dentes exter- —
Fig. 5 Miconia innata, stamens X 18 iores rotundate
crassa, ca. 0.2--0.3
mme in diametro.
Petala obovata alba, 2 mm. longa. Stamina fere isomorphea;
filamenta complanata, 1.6 mm. longa; thecae oblongae obtusae
4-loculares, poro satis lata terminslis dehiscentes; connec-
tivum minutissime productum in lobum dorsalem, in stem. ser.
ext. obscure bilobum, in ser. int. angustatum. Stigma capit-
atum.
Frov. Santiago-Zamora, Ecuador, between Pailas and El

Pan, altitude 2255--2445 meters, Steyermark 54309.

MICONIA HIRSUTIVENA sp. nov. Sect. Cremanium. Caules,
petioli, et basibus venarum ma jorum longe hirsuta. Flores 5-
meri. Antherae isomorphae, obovato-oblongse, 2-loculares,
connectivo basi producto in lobum unicum dorsalem late obo-
vatum. Stylus clavatus; stigma truncatum. Folia elliptica a-
cuminata 43-nervia glabra, venis exceptis.

Shrub 5m. tall, the stems roughly hirsute with simple
hairs about 3 mm. long. Fetioles 6--10 mm. long, similarly
hirsute. Blades thin, elliptic, up to 12 cm. long by 5 cm.
wide, slenderly acuminate, entire, obtuse or subrotund at
base, S-nerved with an additional pair of marginal veins,
glabrous on both sides except for the hirsute bases of the
primary veins. Panicle about 1 dm. long, merely furfurace-
ous. Flowers S-merous, all on pedicels 1--1.5 mm. long. Hy-
panthium cup-shaped, 1.8 mm. long to the torus, glabrous.

306 PHYTOLOGIA Vol. 2, no. 8

_ Calyx-tube nearly erect, 0.8 mm. long; sepals truncate-
triangular, about 0.4 mm. long; exterior teeth merely total-
ly adnate thickenings. Petals obovate, inequilateral, white,
1.9 mm. long. Stamens isomorphic; filaments flat, 45.5 mm.
long, tapering from a broad base, geniculate at two-thirds
of their length, glabrous; anthers oblong, 1.5 mm. long;
connective greatly thickened below and prolonged about 0.3
mm. below the thecae, not lobed. Style gradually enlarged
distally, glabrous, 3.5 mm. long; stigma truncate.

Prov. El Oro, Ecuador, between Paccha and Puente Grande,
altitude 1830--2430 meters, Steyermark 54142. The species
appears related to M. divergens Triana, in which the panicle
and upper leaf-surface are pilose and the flowers smaller.

NOTES ON NEW AND NOTEWORTHY PLANTS. II

Harold N. Moldenke

AEGIPHILA FARINOSA Moldenke, sp. nov.

Arbor; ramulis crassis tetragonis cavis dense ochraceo-
farinosis; petiolis crassis dense ochraceo-farinosis; lamin-
is late ellipticis vel subobovatis breviter acuminatis, ad
basin attenuato-acutis vel breviter acuminatis, integris
supra parce farinosis glabrescentibus, subtus farinosis; in-
florescentiis axillaribus vel supre-axillaribus bifurcatis
fulvo-farinosis; calyce truncato integro vel minutissime 4-
apiculato.

Tree to 8 m. tall; bark almost flat, gray-ochraceous,
succulent, clear-ochre in section; wood pliant, white;
branchlets apparently stout, tetragonal, hollow, ampliate
and flattened at the nodes, densely ochraceous-farinose,
slightly tuberculate-lenticellate; nodes not annulate; prin-
cipal internodes 3--4 cm. long; leaves decussate-opposite;
petioles stout, 3--4 cm. long, densely ochraceous-farinose;
blades membranous-chartaceous, clear-green above when fresh,
somewhat lighter beneath, broadly elliptic or very slightly
obovate, 13.5--24 cm. long, 6--10 cm. wide, short~-acuminate
at apex, attenuate-acute or snort-acuminate at base, entire,
sparsely farinose above but glabrescent in age except for
the densely farinose midrib, sparsely farinose’ beneath, more
densely so on the midrib and larger veins; midrib stout,
prominulous above, very prominent beneath; secondaries slen-
der, 9 or 10 per side, arcuate-ascending, arcuately joined
in many loops some distance from the margins, plane above,
prominulous beneath; vein and veinlet reticulation conspicu-

1947 Moldenke, New and Noteworthy Plants 307

ous on both surfaces and very slightly prominulous, usually
densely farinose; inflorescences axillary or supra-axillary,
2 per node, several times bifurcate, their branches densely
tawny-farinose, stout, firm, widely divergent; bractlets and
prophylla linear-elongate, 1--7 mm. long, very densely
tawny-farinose; calyx campanulate, about 4 mm. long and
wide, densely tawny-farinose, its rim truncate, entire or
very obscurely 4-apiculate; corolla hypocrateriform, yellow-
ish-white, violet outside at the extremities, the tube
broadly cylindric, about 4 m. long, glabrous or slightly
farinose outside, often slightly farinose at the throat
within, its rim 4-parted, the lobes broadly elliptic, about
5 mms long and 4 mm. wide, entire, glabrous on both surfaces
or slightly farinose at the base; stamens exserted; fila-
ments filiform, white, projecting about 1 m. from the cor-
olla-limb when this is erect, about 5 or 6 mm. when this is
wide-spreading; anthers yellowish-white; pistil included.
The type of this interesting species was collected by
José Cuatrecasas (no. 21689) at La Falma on the right bank
of the Rfo Fichindé, Hoya del Rfo Cali, on the eastern
slopes of the Cordillera Occidental, El Valle, Colombia, at
an altitude of 2500 m., on July 24, 1946, and is deposited
in the Britton Herbarium at the New York Botanical Garden.

ALONSOA WARSCEWICZII f. COCCINEA Moldenke, f. nov.

Haec forma a forma typica speciei corollis coccineis re-
cedit. - This form differs from the typical form of the spe-
cies in its scarlet corollas. The type was collected by me
(no. 7827) from cultivated plants at Watchung, Somerset Co.,
New Jersey, on July 30, 1933, and is deposited in the Brit-
ton Herbarium at the New York Botanical Garden.

ALOYSIA CHIAFPENSIS Moldenke, sp. nov.

Frutex; ramis ramulisque obtuse tetragonis dense hirsutu-
lis; internodiis valde abbreviatis; petiolis gracillimis
densissime hirsutulis vel villosulis; laminis chartaceis
lanceolatis, ad apicem acutis vel rotundatis, serrulatis, ad
basin acutis, supra rugosis et dense pustulato-strigosis,
subtus densissime tomentellis; inflorescentiis congestis.

Shrub, apparently considerably branched; stems subterete
or obscurely tetragonal, gray, glabrate; branches and
branchlets obtusely tetragonal, brownish, densely hirsutu-
lous, not resinous-punctate, less densely so in age; nodes
not annulate; principal internodes mostly greatly abbreviat-

ed, 2--30 mm. long, occasionally to 6.5 cm. long on vigorous

shoots; leaves decussate-opposite, apparently caducous;
leaf-scars large, prominent, corky, more or less lunate, on
divergent sterigmata; petioles very slender, 1--4 mm. long,
very densely hirsutulous or villosulous with white hairs;

308 PHYTOLOGIA i Vol. 2, no. 8

leaf-blades (immature?) chartaceous, uniformly green on both.
surfaces, lanceolate, 1--2.5 cm. long, 4--9 mm. wide, acute
or rounded at apex, uniformly serrulate from apex to base
with small blunt teeth, acute at base, rugose above and
densely strigose with white, pustulate-based, antrorse hairs
very densely tomentellous or short-pubescent beneath, the
hairs on the larger venation beneath sometimes strigose on
younger leaves; the very slender midrib and about 7 pairs of
close ascending-divergent secondéries somewhat impressed a-
bove and prominent beneath, the abundant veinlet reticula-
tion also.more or less impressed above and prominent be-
neath; inflorescence axillary, 2--6 per node, greatly con-
gested toward the tips of the branchlets, but apparently al-
so produced on entirely leafless branches, about equaling
the leaves where these are present (but the leaves may still
be immature), divergent or drooping, very densely many-
flowered, 1--3 cm. long; peduncles very slender, 5 mm. long
or less, very densely hirsutulous; brectlets large, foliace-
ous, conspicuous, lanceolate, about 6.5 mm. long and 2.5 mm.
wide, 3-nerved, attenuate-subacuminate et apex, rather
densely short-pubescent with microscopically glandular-capi-
tate hairs and sericeous-villosulous with much longer whit-
ish antrorse hairs on the back, only microscopically puberu-
lent on the inner surface; calyx cupuliform, 2-parted, 1--
1.2 mm. long, densely setulose-hirsute on the outside with
stiff widely spreading hairs as long as or longer than the
diameter of the calyx, each segment navicular, obtuse at a-
pex, glabrous within; corolla infundibular or hypocrateri-
form, about 3.8 mm. long in all, its tube cylindric, about
O.7 mm. wide at base, constricted immediately above the ov-
ary to 0.3 mm., ampliate to 0.9 mm. at apex, puberulent or
short-pubescent with spreading hairs from just above the ov-
ary to the apex on the outer surface, densely pubescent
within, its limb about 1.5 mm. wide, 5S-lobed, the lobes un-
equal, the largest less than 1 mm. wide and O.5 mm. long,
rounded, undulate-margined, glabrous on the inner and pubes-
cent on the outer surface; stamens 4, didynamous, inserted
about 0.8 mm. below the mouth of the corolla-tube; filaments
practically obsolete; anthers very smell, apparently effete;
pistil one; style capillary, about 2.1 mm. long, glabrous;
stigma lateral, oblique, about 0.4 mm. long; ovary subglo-
bose, about 0.6 mm. long and wide, glabrous.

The type of this rrmarkable species was collected by Carl
Albert Purpus (no. 10519) on rocky banks at Monserrate,
Chiapas,: Mexico, 1 in Marcel March, 1925, and is deposited in the
Britton Herbarium at the New York Botanical Garden. The spe-
cies is obviously related to A. barbata (T. S. Brandeg.)
Moldenke, of Baja California, and may like that species be
polygamo-dioecious. It differs from A. barbata in its pubes-

1947 Moldenke, New and Noteworthy Plants 309

cence, leaf-size and shape, and especially the much smaller
size of its flowers.

ALOYSIA REICHII var. TRILOBATA Moldenke, var. nov.

Haec varietas a forma typica speciei foliis semper 3-
lobatis recedit. - This variety differs from the typical
form of the species in having all its leaves 3-lobed.

The type was collected by Rodolfo Wagenknecht (Looser
4238) at Rfo Turbio, dept. Elqui, Coquimbo, Chile, on Octo-
ber 19, 1940, and is deposited in the Britton Herbarium at
the New York Botanical Garden. It was inaccurately reported
by me in Lilloa 6: 312 (1941) as typical A. Reichii Molden-
ke, whose leaves are mostly unlobed.

ALOYSIA TERNIFOLIA Moldenke, sp. nov.

Frutex; ramulis gracilibus subtetragonis striatis griseis
adpresso-pubescentibus; foliis ternatis; petiolis gracilli-
mis densiuscule adpresso-pubescentibus; laminis chartaceis
ellipticis abrupte acutis vel obtusiusculis, apicem versus
6--12-dentatis, supra minute scabrello-puberulis, subtus
molliter puberulis; inflorescentiis spicatis multifloris.

Shrub; branchlets slender, subtetragonal, somewhat stri-
ate-ridged, gray, appressed-pubescent with very short gray-
ish hair; nodes annulate; principal internodes 2.5--4 cm.
long; leaves ternate; petioles very slender, 1--5 mm. long,
rather densely appressed-pubescent with very short white or
grayish hairs like the branchlets, slightly margined especi-
ally toward the apex; blades chartaceous, bright-green, only
very slightly lighter beneath, elliptic, 4--7.5 cm. long,
1.2--4 cm. wide, abruptly acute or bluntish at apex, the
lower 2/3 of the margin entire, the upper 1/3 with 6--12
broad and rather blunt antrorse teeth, minutely scabrellous-
puberulent above under a handlens, softly puberulent with
obscure hairs beneath, slightly dense on the larger venatim;
inflorescence spicate, in the upper leaf-axils, 2 or 3 per
node, /--9 cm. long, many-flowered, rather dense, all except
the very lowest flowers more or less imbricate; peduncles
very slender, 1.5--4 cm. long, densely white-puberulent; ra-
chis very slender, densely white-puberulent; bractlets lan-
ceolate, 1.5--35 mm. long, 0.5 mm. wide, long-acuminate, ap-
pressed-puberulent on the back; calyx deeply 2-labiate, the
lips divergent, about 2.8 mm. long, finely appressed-puberu-
lent on the outside, each usually 2-lobed or 2-toothed at
the apex;. corolla hypocrateriform, its tube 3.5--4 mm. long,
about 1 mm. wide at the base, ampliate to 2 mm. at the cent-
er and from there to the apex, glabrous outside, densely to-
mentose within, the limb 2-lipped, the upper lip 2-lobed,
the lower lip 3-lobed, the lobes lingulate-orbicular, 1.5--2
mm. long and wide, the margins slightly undulate, glabrous

310 PHY TOL 0G T Aw Vol. 2, no. 8

outside, pubescent at the base within; stamens 4, inserted

near the apex of the corolla-tube, included, didynamous;
filaments obsolete or to 0.5 mm. long; anthers 2-celled;

- gtyle stoutish, about 1.35 mm. long, glabrous, firm; stigma

capitate, slightly 2-lobed; ovary obovate, about 1 mm. long

and 1.3 mm. wide, glabrous, 2-celled.

Tne type of this very distinct species was collected by
Fer Karl Hjalmar Dusen (no. 4228) at a rivulet at Iteiacooa,
near Ponta Grossa, Santa Cruz, Argentina, March 17, 1904,
and is deposited in the herbarium of the Naturhistoriske
Riksmuseum at Stockholm.

ALOYSIA VIRGATA ver. PLATYPHYLLA (Briq.) Moldenke, comb.nov.

Lippia virgata var. elliptica Briq., Ann. Conserv. &
Jard. Bot. Genev. 7-8: 304. 1904.

CALLICARPA CANDICANS var. PERRYANA (Dop) Moldenke, comb.nov.

Callicarpa cana ver. Perryana Dop, Bull. Soc. Hist. Nat.
Toulouse 64: 504, 1932.

CHLOANTHES GRANDIFLORA Moldenke, sp. nov.

Frutex, caulibus densissime lanato-tomentosis; internod-
iis abbreviatis; foliis sessilibus crassis, ad apicem obtus-
is vel subacutis, integris utrinque densissime lanato-tomen-
tosis; floribus solitariis vel fasciculatis axillaribus.

Apparently shrubby; stems subterete, very densely lanate-
tomentcse, less densely so at the apex; principal internodes
abbreviated, about 1 om. long below the inflorescences and
to 2.5 cm. long among the inflorescences; leaves decussate-
opposite, sessile, thick-textured, 1.8--3 em. long, 8--15
mn. wide, obtuse or subacute at apex, entire, very densely
white-lanate-tomentose on both surfaces or becoming merely
densely stellate on both surfaces; midrib and a few long-
ascending secondsries sometimes barely visible through the
tomentum; flowers solitary or in small clusters in the upper
ten leaf-axils, the lower ones often borne on stout white-
lanate peduncles 1--2 cm. long; calyx campanulate, deeply 5-
parted almost to the base, the lobes equal, elliptic, about
8 mms long, 3.5--4 mm. wide, acute at the apex, densely
lanate-tomentose on the outer surface with white hairs, ven-
ose; corolla large, showy, tubular, the tube about 2 cm.
long, slightly asymmetrical and curvate, about 4 mm. wide at
the base and to above the ovary, ampliate to 12 mm. near the
apex, glabrous or very obsoletely puberulent outside, glab-
rate within except for the densely villous-tomentose ring a-
bove the ovary, venose, the limb 2-lipped, the upper lip 2-
lobed, the lower lip 3-lobed, the lobes ovate-orbicular, 3--
4 mm. long, 4--5 mm. wide, rounded, puberulent on the outer
surface, venose; stamens 4, inserted at about the middle of

1947 Moldenke, New and Noteworthy Plants 311

the corolla-tube, included, didynamous; filaments flattened,
6--7 mm. long, glabrous; anthers bifid; style capillary, a-
bout 17 mm. long, glabrous, included or equaling the corol-
la-tube; stigma unequally and shortly bifid; ovary small, a-
bout 1 mm. long and wide, farinose-pulverulent.

The type of this species was collected by J. Mauritzon
somewhere in Western Australia in September, 1936, and is
deposited in the herbarium of the Naturhistoriska Riksmuseum
at Stockholm.

CONGEA CHINENSIS Moldenke, sp. nov.

Frutex scandens; ramis dense villoso-pubescentibus; nodis
annulatis; petiolis dense villosis; laminis ellipticis acum-
inatis, ad basin rotundatis vel subcordatis, integrise supra
leviter pilosulis subtus plusminusve dense piloso-purictulat-
is; inflorescentiis paniculetis; bracteolis involucri 4 el-
lipticis vel suboblanceolatis ad basin cmnatis utrinque
dense albo-tomentosis obtusis vel subacutis.

Woody vine; stems branched; branches rather slender, sub-
terete or very obtusely tetragonal, densely villous-pubes-
cent with more or less appressed antrorse brownish hair,
less so in age; principal internodes 3.5--6 cm. long; nodes
distinctly annulate; leaves decussate-opposite; petioles
rather stout, 5 mm. long or less, densely villous like the
branchlets; blades dark-green and brunnescent in drying a-
bove, lighter beneath, chartaceous, elliptic, 7--10.5 cm.
long, 3.5--4.2 cm. wide, acuminate at apex, usually rounded
or subcordate at base, entire, lightly pilosulous above,
more densely pilose on the larger veins, more or less dense-
ly pilose-punctulate beneath, more densely so on the larger
veins; midrib very slender, slightly prominulent above,
prominent beneath; secondaries slender, 4--6 per side, arcu-
ate-ascending, plane above, prominulent beneath; vein and
veinlet reticulation obscure above, subprominulent beneath;
inflorescence racemose, in pairs in the upper axils, forming
a large terminal panicle; peduncles slender, 5--9 cm. long,
densely villous like the branches, annulate and bracteate at
the nodes of the rachis; bracts foliaceous, el liptic-lanceo-
late, 1.5--6 cm. long, 4--20 mm. wide, decreasing in size
toward the apex of the inflorescence, in pairs at the nodes,
similar to the leaves in color, texture, and pubescence or
more densely pubescent on both surfaces; pedicels slender,
6--15 mm. long, densely villous, two per node; involucre
composed mostly of 4 bractlets, elliptic or slightly oblan-
ceolate, connate at base to forma cup about 6 mm. high, the
free portions 2--2.5 cm. long, obtuse or subacute at apex,
densely white-tomentose on both surfaces; flowers about 5
per head, densely white-villous-tomentose with short appres-
sed hairs on the outer surface.

512) - PHYTOLOGIA Vol. 2, now 8

The type of this distinct species was collected by H. T.
Taai (no. 52611) in Ytinnan, China, in 1932, and is deposited
in the herbarium of the Naturhistoriska Riksmuseum at Stock-
holm. It was determined as C. tomentosa Roxb. by Re C.
Ching, and so distributed, but differs pronouncedly from
thet species in the large involucral cups and in other char-
acters. Its involucres are similar to those of C. connata
Fletcher and C. siamensis.Fletcher, of Thailand, which, how-
ever, may be distinguished at once by their uniformly ae
parted involucres.

HELIPTSRUM ROSEUM f. ALBUM (Le H. Bailey) Moldenke, stat.
nove
Helipterum album Hort. ex L. H. Bailey, Cycl. Am. Hort.
3:3 72 ° 190 e

LANTANA EHRENBERGIANA Moldenke, sp. nov.

Frutex; ramis obtuse tetragonis submarginatis glabris;
ramulis gracillimis numerosis tetragonis antrorse strigosis
glabrescentibus; internodiis abbreviatis; petiolis strigos-
is; laminis firme chartaceis ovatis obtusis, ad basin sub-
truncatis, serratis supra strigosis rugosis subtus dense
griseo-tomentellis; capitulis subglobosis; bracteis ovatis
foliaceis strigillosis acutis vel breviter acuminatis.

Shrub; branches obtusely tetragonal, slightly margined on
the angles, glabrous, gray; branchlets and twige very slend-
er, numerous, tetragonal, antrorsely strigose-pubescent, be-
coming glabrescent in age; principal internodes 0.5--2.8 cm.
long, mostly quite abbreviated; leaves decussate-opposite ;
petioles 1--4 mm. long, densely antrorse-strigose; blades
firmly chartaceous, rather dark-green above, lighter beneath
ovate, to 3 cm. long and 2 cm. wide, obtuse at apex, sub-
truncate at base and slightly cuneately attenuate into the
petiole in the middle, coarsely serrate along the margins
from base (except the cuneate prolongation) to apex with
rounded rather appressed antrorse teeth, strigose above and
usually rugose, densely gray-tomentellous beneath; midrib
slender, impressed above, prominulent beneath; secondaries
slender, 4 or 5 per side, impressed above, prominulent be-
neath; vein and veinlet reticulation impressed above, usual-
ly not obvious beneath; inflorescence axillary toward the
tips of the twigs, apparently usually one per node, ascend-
ing or erect; peduncles very slender, 1--4 cm. long, strig-
ose; heads subglobose, 1--1.5 mm. wide, densely flowered;
bracts ovate, foliaceous, to 7 mm. long and 5 mm. wide, ac-
ute or shortly acuminate, strigillose; corolla slightly sur-
passing the brects.

The type of this species was collected by Carl August Eh-
renberg -- in whose honor it is named -- at Santo Domingo,

a ae

: : :
1947 Moldenke, New and Noteworthy Plants S58.

Dominican Republic, Hispaniola, in or before 1839, and is
deposited in the Meisner Herbarium at the New York Botanical
Garden.

LANTANA MSARNSII var. CONGOLENSIS Moldenke, var. nov.

Haec varietas a forma typica speciei recedit laminis fol-
iorum tenuiter chartaceis vel submembranaceis supra scab-
rellis subtus plusminueve leviter puberulis; inflorescentiis
2--6 aggregatis, pedunculis gracilibus in longitudine valde
variabilibus plerumque 2.5--3 cm. longis; spicis usque ad 3
em. elongatis; bracteis lanceolatis attenuato-acuminatis
laxe puberulis.

This variety differs from the typical form of the species
in its thin-cnartaceous or even submembranous leaf-blades
which are scabrellous above and more or less lightly puberu-
lent beneath. The inflorescences ere 2--6 per node, the
slender peduncles very variable in length, usually 2.5--3
cm. long. The spikes elongate to 3 cm. after anthesis, and
the bracts are lanceolate, /7--10 mm. long, 2--3 mm. wide at
the base, attenuate-acuminete at the apex, loosely puberu-
lent.

The type of this variety was collected by Feller (no. A.
46) at Congo da Lemba, Belgian Congo, on April 24, 1913, and
is deposited in the herbarium of the Jardin Botanique de
l'Etat at Brussels. A common name is “disisusu na bakala".
The corolla is described as wnite by the collector, who also
states that the plant is boiled down for vapor baths.

LANTANA MEARNSII var. LATIBRACTZOLATA Moldenke, var. nov.

Haec varietas a forma typica speciei recedit pedunculis
usque ad 1 cm. longis ternatis et bracteis /7--9 mn. longis,
4--5 mm. latis, ad apicem triangulari-acutis rectis imbrica-
tis densiusculs breviterque pubescentibus ciliatis.

This variety differs from the typical form of the species
in its uniformly snort-peduncled spikes, the peduncles only
1 cm. long or less, 3 per node, and its broadly ovate bracts
which are 7--9 mm. long and 4--5 mm. wide at the base, ab-
ruptly narrowed to the triangular-acute apex, erect, imbri-
cate, rather densely snort-pubescent and ciliate-margined.
The leaves are thin-chartaceous, scabrellous-puberulent a-
bove, densely tomentellous beneath.

The type of this variety was collected by Joseph Charles
Corneille Bequaert (no. 5490) in the steppes at the edge of
& leke, Kabare, Belgian Congo, on August 29, 1914, and is
deposited in the herbarium of the Jardin Botanique de l'&tat
at Brussels. The corolla is described by the collector as
red-violet.

LIPFIA CHSVALIERII Moldenke, sp. nov.

314 PHYTOLOGIA 3 Vol. 2, no. 8

Herba (?); ramis gracillimis stramineis subteretibus ob-
scure strigillosis; foliis ternatis vel verticillatis sub-
sessilibus; laminis chartaceis oblanceolatis acutis, ad bas-
in cuneato-attenuatis, argute serratis supra parciuscule
strigosis, subtus dense strigosis; inflorescentiis axillar-
ibus 3--6 aggregetis; capitulis oblongo-cylindricis densis-
sime flavo-tomentosis; bracteolis ovatis acuminetis.

Herbaceous (7); branches very slender, stramineous, sub-
terete, striate, rather obscurely strigillose; nodes annu-
late; principal internodes elongate, 5--14 cm. long; leaves
ternate or in 4's, subsessile; blades chartaceous, often
rather thin, bright-green a grayish-green beneath, ob-
lanceolate, 4--6 cm. long, 1.3--2 cm. wide, acute at apex,
cuneate-attenuate at base, sharply serrate except at and
near the base, the teeth small and rather obtuse, antrorse,
rather sparsely strigose above, much more densely so be-
neath; midrib very slender, usually plane above, prominu-
lent beneath; secondaries very slender, 4 or 5 per side,
ascending, almost indiscernible above, prominulous beneath;
vein and veinlet reticulation indiscernible above, rather
obscure beneath, flat; inflorescence axillary only, 3--6

per node in the uppernost 2 or 3 nodes, usually shorter

than the subtending leaves; peduncles slender, 5--15 mm.
long, densely white-pubescent with antrorse hairs; heads
oblong, cylindric, 4--11 mm. long, 5--6 mm. wide, very
densely yellow-tomentose, all save the lowest bractlets
completely hidden by the yellow tomentum; lowest bractlets
ovate, about 3 mm. long and 2 mm. wide, acuminate, densely
tomentose on the back; corolla about 4 mm. long in all,
its limb about 2 mm. wide.

The type of this species was collected by August J. B.
Chevalier (no. 67) -- in whose honor it is named -- at
Toukota, French Soudan, French West Africa, on December
28, 1898, and is deposited in the herbarium of the Jardin
Botanique de l'Etat at Brussels.

LIPPIA DOMINGENSIS Moldenke, sp. nov.

Frutex decumbens; caulis prostratis gracilibus saepe ad
nodos radicantibus glabris; ramis tetragonis dense brunn-
eo-puberulis resinosis glabrescentibus; foliis oppositis
vel ternatis numerosis parvissimis; petiolis grecillimis
strigilloso-puberulis resinosis; laminis crassiusculis o-
valibus vel suborbicularibus, ad apicem rotundetis, ad ba-
sin acutis, supra valde bullatis rugosisque scaberrimisque
reculariter dentatis, subtus adpresso-pubescentibus.

Prostrate or spreading shrub; stems to 4 feet long,
slender, gray, often rooting at the nodes, glabrous, the
bark exfoliating in age; branches numerous, short, very
slender, tetragonal, densely brownish-puberulent, resin-

1947 Moldenke, New and Noteworthy Plants 315
, ‘

ous, glabrescent in age; branchlets and twigs numerous, very
slender, tetragonal, densely brownish-puberulent, resinous;
nodes annulate; principal internodes abbreviated, 3--20 mm.
long; leaves decussate-opposite or ternate, numerous, very
small; petioles very slender, 1--2.5 mm. long, strigillose-
puberulent, resinous; blades very. small, rather thick-tex-
tured, bright-green above, somewhat lighter beneath, oval or
suborbicular, 4--15 mm. long, 3--11 mm. wide, rounded at a-
pex, acute at base, deeply bullate above, the margins regu-
larly dentate with small rounded rather spreading teeth, ru-
gose and very scabrous above, appressed-pubescent or strig-
ose beneath and somewhat resinous; midrib slender, deeply
impressed above, very prominent beneath; secondaries very
slender, 3--6 per side, ascending, rather straight, deeply
impressed above, very prominent beneath; tertiaries very
slender, connecting the secondaries and at right angles to
them, rather straight and subparallel, deeply impressed a-
bove, prominulent beneath; inflorescence axillary, sparse,
less than 1 cm. long in all, about equaling the subtending
leaves; peduncles very slender, 4--5 mm. long, densely pub-
erulent and resinous; heads few-flowered, subglobose, not
elongating in fruit; bractlets elliptic or oblanceolate,
2.5--53 mm. long, 1 mm. wide, obtuse or subacute at apex,
resinous-puberulent; corolla white, its tube 3--4 mm. long,
slightly surpassing the subtending bractlets; corolla-limb
1--1.5 mm. wide.

The type of this species was collected by Richard A. and
Ee S. Howard (no. 8110) at the edge of a limestone ravine
in pine woods along the trail between Federnales and Aceit-
ial, alt. 4200 feet, prov. Barahona, Dominican Republic,
between August 8 and 12, 1946, and is deposited in the
Britton Herbarium at the New York Botanical Garden. The
collectors note that only 4 flowers open at a time in each
head.

LIPPIA LEFIDA Moldenke, sp. nov.

Planta pumila, ad basin lignosa, usque ad 15 cm. alta;
caulibus gracilibus subteretibus dense breviterque pubes-
centibus saepe glanduliferis; foliis oppositis sessilibus
ellipticis obtusis argute serratis, ad basin subacutis vel
obtusis, supra dense breviterque pubescentibus, subtus
albo-tomentosis; inflorescentiis capitatis; bracteis magnis
ovato-ellipticis acutis glanduloso-pubescentibus imbricatis

Dwarf plant, apparently from a woody base, to 15 cm.
tall; stems slender, subterete, densely short-pubescent
with spreading often gland-tipped hairs, brown macroscop-
ically, but glistening-silvery microscopically; nodes 2--4,
not annulate; internodes 2.5--5.5 cm. long, or the very
lowest abbreviated to 1 cm. or less; leaves 1--4 pairs, de-

316 Po Ye T0c EB) Ore Bak Vol. 2, now 8 ©

}
cussate-opposite, sessile, elliptic, 7--14 mm. long, 4--8
mm. wide, obtuse at apex, sharply serrate, subacute or ob-
tuse at base, densely short-pubescent above and white-
tomentose beneath; inflorescence capitate, usually two at
each of the 1 or 2 upper nodes and a single terminal one;
peduncles very slender, 1--3.5 om. long, densely glandular-
pubescent like the stems; heads hemispheric, about 1 cm.
long and 1.5 cme wide, densely many-flowered; bracts large,
ovate-elliptic, about 5 mm. long and wide, acute at apex,
glandular-pubescent, overlapping; corolla rose, hypocrateri-
form, projecting about 5 mm. beyond the bracts, its limb 5
mm. or more wide.

The type of this little species was collected by Ae Fe Me
Glaziou (no. 21891) near the encampment at Corrego do Brejo,
Goyaz, Brazil, in March or April, 1883, and is deposited in
the herbarium of the Jardin Botanique de l'Etat at Brussels.

LIFPIA SCHLIEBENI Moldenke, sp. nov.

Fruticulus; caulibus ramisque griseis obtuse tetragonis
scabrellis; ramulis tetragonis brunneis breviter pubescenti-
bus vel puberulis; foliis oppositis numerosis; petiolis den-
se puberulis; laminis chartaceis ovetis acutis dense serru-
latis, as basin acuminatis, supra scabris bullatisque, sub-
tus adpresso-canescento-puberulis; inflorescentiis axillari-
bus spicatis usque ad 2 cm. elongatis.

Dense bush; stems and branches gray, obtusely tetragonal,
scabrellous; twigs tetragonal, short-pubescent or puberu-
lent, brownish; nodes not annulate; leaf-scars elevated,
corky; principal internodes 2--15 mm. long on twigs, to 5
cme long on the main stem; leaves decussate-opposite, numer-
ous; petioles slender, 1--6 mm. long, densely puberulent;
blades chartaceous, bright-green above, lighter beneath, o-
vate, 1.5--5 cm. long, 5--15 mm. wide, acute at apex, dense-
ly serrulate, acuminete at base, scabrous and bullate above,
appressed canescent-puberulent beneath; midrib slender,
deeply impressed above, prominulent beneath; secondaries
slender, 3--5 per side, ascending, not much arcuate, deeply
impressed above, prominulent beneath; vein and veinlet re-
ticulation beautifully conspicuous and deeply impressed a-
bove, prominulent beneath; inflorescence axillary, abundant,
spicete, 2 per node; peduncles slender, 2--6 cm. long, den-
sely puberulent, glebrescent in age; spikes subcapitate dur-
ing anthesis, elongate to 2 cm. in fruit, densely many-flow-
ered; brects ovete, numerous, reflexed during anthesis, a-
bout 5 mm. long and 3 mm. wide, sttenuate at apex, densely
puberulent on both surfaces; corolla hypocrateriform, white,
surpassing the subtending bract by about 3 mm., densely pub-
escent on the outer surface, its limb about 3 mm. wide

Tne type of this species was collected by H. J- Schlieben

\

1947 | Moldenke, New end Noteworthy Plants 317

(no. 5596) -- in whose honor it is named -- at Mucraplateau,
Bakari, 80 km. west of Lindi, alt. 600 m., Tanganyika Terri-
tory, on October 26, 1934, and is deposited in the herbarium
of the Jardin Botanique de 1'Etat at Brussels.

LIPPIA STROBILIFORMIS Moldenke, sp. nov.

Fruticulus (7); caulibus gracilibus tetragonis sulcatis
strigillosis; ramis paucis brevibus; foliis oppositis; peti-
olis gracilibus strigosis; laminis firme chartaceis lanceo-
latis vel ellipticis acutis regulariter serruletis, ad basin
acutis, supra scabris subbullatisque, subtus puberulis res-
inoso-granulatisque; inflorescentiis axillaribus spicato-
strobiliformibus numerosis; bracteis magnis perspicuis.

Stems slender, tetragonal, ridged and sulcate, strigil-
lose; branches few, short; nodes not annulate; principal in-
ternodes 3.5--6 cm. long; leaves decussate-opposite; petio-
les slender, 2--5 mm. long, strigose; blades firmly charta-
ceous, grayish-green on both surfaces, lanceolate or ellip-
tic, 3--6 cm. long, 1--2 cm. wide, acute at apex, regularly
serrulate along the margins, acute at base, scabrous and
slightly subbullate above, puberulent and resinous~-granular
beneath; midrib slender, subprominulous above, prominulous
beneath; secondaries slender, 6--8 per side, ascending, only
slightly arcuate, subimpressed above, prominulous beneath;
vein and veinlet reticulation subimpressed above, subpromin-
ulous beneath; inflorescence axillary, spicate-strobiliform,
2 per node, abundant; peduncles slender, firm, ascending,
1--5.8 cm. long, tetragonal, strigillose; spikes very dense-
ly capitate, to 2 cm. long and 1.5 cm. wide; bracts large
and conspicuous, very numerous, very closely imbricate and
appressed antrorsely, ovate, about 7 mm. long, 4--5 mn.
wide, abruptly acute or subacuminate at apex, very densely
appressed-pubescent with antrorse hairs which project prom-
inently beyond the margins and form what appears macroscop-
ically like a light border to the bracts; corolla hypocret-
eriform, fts tube 7--8 mm. long, densely puberulent outside,
the limb about 4 mm. wide.

The type of this handsome species was collected by Cap-
tain Storms (no. 3) at Karema, Tanganyika Territory, and is
deposited in the herbarium of-the Jardin Botanique de 1'Etat
at Brussels. The long slender roots are remarkable because
of the large number. of sessile nodules which they bear.
These nodules are 2--4 mm. in diameter.

LIFPIA STROBILIFORMIS var. ACUMINATA Moldenke, var. nov.

Haec varietas a forma typica speciei recedit foliis saepe
ternatis, inflorescentiis saepe ternatis, spicis dense capi-
tatis, brecteis valde patentibus non adpressis lanceolato-
ovatis longe acuminatis dense puberulis.

318 PHYTOLOGIA yol. 2, neue

This variety differs from the typical form of the species
in having often ternate leaves and inflorescences and in
having the bracts of its dense capitate spikes wide-spread-
ing, not appressed, lanceolate-ovate, about 1 om. long, 4--
4.5 mm. wide, long-acuminate at the apex, densely puberu-
lent, without a light border. The mature spikes are 2 cm.
wide and the peduncles are to 4.5 cm. long.

The type of this variety was collected by Pere Hyacinthe
Vanderyst (no. 17168) in the region of Panzi, Belgian Congo,
in 1925, and is deposited in the herbarium of the Jardin

Botanique de 1'&tat at Brussels.

LIPPIA STROBILIFORMIS var. PARVIFOLIA Moldenke, var. nov.

Haec varietas a forma typica speciei recedit foliis par-
vioribus, petiolis 1 mm. longis vel obsoletis, laminis ell-
ipticis 1.5--3 cm. longis, 6--14 mm. latis; bracteis paten-
tibus vel reflexis ovatis non adpressis breviter acuminatis.

This variety differs from the typical form of the species
in its smaller leaves, the petioles being 1 mm. long or ob-
solete, the blades elliptic, 1.5--3 cm. long, 6--14 mn.
wide; inflorescences 2--4 per node, 1--5 cm. long; and the
bracts spreading or reflexed, closely imbricate but not ap-
pressed, ovate, 7--8 mm. long, 3--4 mm. wide, short-acumin-
ate at the apex, densely appressed-puberulent, sometimes
subrevolute along the margins, without « lighter margin or
with an obscure one.

The type of thie variety was collected by Pere Hyacinthe
Vanderyst (no. 23423) et Merode, Belgian Congo, and is de-
posited in the herbarium of the Jardin Botanique de L'Etet
at Brussels.

LIPPIA WOODII Moldenke, sp. nov.

Herba; caulibus plerumque simplicibus dense puberulis;
foliis oppositis paucis; petiolis strigoso-puberulis; lam-
inis chartaceis rectis anguste ellipticis supra scabris
subbullatisque, subtus dense puberulis vel breviter pubes-
centibus; inflorescentiis capitato-spicatis multifloris

Herb; stems subterete toward the base, subtetragonal
toward the apex, mostly unbranched, densely puberulent,
less densely so in age; nodes not annulate; principal inter-
nodes 3--14.5 cm. long; leaves decussate-opposite, few, usu-
ally with a very much abbreviated branchlet and a few small
leaves in their axils; petioles slender, 1--5 mm. long,
strigose-puberulent; blades chartaceous, rather grayish-
green on both surfaces, epparently erect on the stem, nar-
rowly elliptic, 4--6 cm. long, 8--12 mm. wide, scabrous and
subbullate above, densely puberulent or short-pubescent be-
neath; midrib slender, impressed above; secondaries very
slender, 4--6 per side, impressed above, ascending, not much

- — - ‘

1947 Moldenke, New and Noteworthy Plants 319

arcuate, prominulous beneath; veinlet reticulation impressed
above, prominulous beneath; inflorescence capitate-spicate,
2 per node at the uppermost 2 or 3 nodes, about equaling the
subtending leaves; peduncles slender, 2--4.5 cm. long,
densely short-pubescent with brown hairs; spikes capitate,
about 1 cm. long, 1.2--1.4 cm. wide, densely many-flowered;
bracts narrowly lanceolate, the lowest about 7 mm. long, 2--
2.5 mm. wide, long-acuminate or caudate at apex, densely
short-pubescent with subappressed hairs, far surpassing the
flowers; corolla hypocreteriform, 3--4 mm. long, densely
short-pubescent with whitish hairs outside, the limb 1--1.5
mn. wide. .

The type of this species was collected by J. Buchanan (J.
Medley Wood 6937) at Blantyre, Nyassaland, and is no. 83373
in the herbarium of the Chicago Natural History Museum.

LYCHNIS COELI-ROSA f. COERULEA Moldenke, f. nov.

Haec forma a forma typica speciei corolla coeruleis rece-
dit. - This form differs from the typical form of the spe-
cies in its sky-blue corollas.

The type was collected by me (no. 10021) from cultiveted
plants at Ville Elsinore, Watchung, Somerset Co., New Jersey
on July 31, 1937, and is deposited in the Britton Herbarium
at the New York Botanical Garden.

MENTHA GENTILIS f. VARIEGATA Moldenke, f. nov.

Haec forma a forma typica speciei foliis albo-variegatis
recedit. - This form differs from the typical form of the
species in having its leaf-blades variegated with white.

The type was collected by me (no. Béle ) from cultivated
plants et Leonia, Bergen Co., New Jersey, on July 15, 1935,
and is deposited in the Britton Herbarium at the New York
Botanical Garden.

NICOTIANA ALATA var. GRANDIFLORA f. RUBSLLA Moldenke, f.nov.

Haec forma a forma typica varietatis corolla rubellis re-
cedit. - This form differs from the typical form of the var-
iety in its pink corollas.

The type was collected by me (no. 8122) from cutlivated
plants at Villa Zlsinore, Watchung, Somerset. Co., New Jer-
sey, on July 25, 1934, and is deposited in the Britton Her-
barium at the New York Botanical Garden.

NIGELLA DAMASCENA f. PLENIFLORA Moldenke, f. nove

Haec forma e forma typica speciei corollis plenis rece-
dit. - This form differs from the typical form of the spec-
ies in its "doubled" corollas.

The type was collected by me (no. 30 38 ) from cultivated
plants at Villa Zlsinore, Watchung, Somerset Co., New Jer-

320 Pine -Och, O atk Vol. 2, noe 8

sey, on August 9, 1926, and is deposited in the Britton Her-
barium at the New York Botanical Garden.

NOLANA ATRIPLICIFOLIA f. ALBA (Fletcher) Moldenke, comb-enov.
Nolana paradoxa var. alba Fletcher in L. H. Beiley, Cycl.
Am. Hort. 510 1092. 1906.

AROSA DAWSONI Moldenke, nom. nov.
Rosa multiflora Thunb. x R. borboniana Desp. ex Rehd.,
Man. cult. Trees & Shrubs, ed. 2, 445. 1940

XROSA FELICITA Moldenke, nom. nov.

Rosa sempervirens Le x Re chinensis Jacq. ex Rehd., Man.
Cults. t. Trees & Shrubs, ed. 2, 448. 1940. ;
RUDBSCKIA HIRTA f. FLENIFLORA Moldenke, f. nov.

Haec forma a forma typica speciei capitulis plusminusve
toto ligulatis recedit. - This form differs from the typical
form of the species in having its flower-heads with several
to many supernumerary series of rays.

,.The type was collected by me (no. 2058 ) in a grassy field
in the valley between the First and Second Mountains, Watch-
ung, Somerset Co., New Jersey, on July 27, 1924, and is de-
posited in the Britton Herbarium at the New York Botanical
Garden.

SANVITALIA PROCUMBENS f. PLENIFLORA Moldenke, nom. nov.
Sanvitalia procumbens var. flore-pleno Hort. ex Barclay
in Le H. Bailey, Stand. Cycl. Hort. 3: 3071. 19535.

STACHYTARFHETA AMFLSXICAULIS Moldenke, sp. nov.

Herba suffrutescens; caulibus ut videtur simplicibus ter-
etibus dense puberulis; foliis oppositis amplexicaulibus;
laminis chartaceis brunnescentibus ovatis, ad apicem attenu-
atis vel acuminatis, serratis, ad basin abrupte angustatis,
supra minute pulverulis vel glabrescentibus, subtus dense
puberulis; inflorescentiis solitariis spicatis densifloris.

Suffrutescent herb; stems apparently simple, terete,
densely puberulent, straight; principal internodes 9--10 cm.
long; leaves decussate-opposite, amplexicaul at base; blades
chartaceous, dark-green on botn surfaces, brunnescent in
drying, ovate, 4.5--8.5 cm. long, 1.9--4 cm. wide, attenuate
or acuminate at apex, serrate along the margins with appres-
sed rounded very regular teeth, abruptly narrowed at base
into a broadly winged petiole about 1 cm. long and 8 mm.
wide, cordate-clasping around the stem, the auricles of the
opposite leaves overlapping each other, minutely and very
inconspicuously pulverulent or glabrescent above, densely
puberulent beneath; midrib slender, plane above, very incon-

\

1947 Moldenke, New and Noteworthy Plants 321

spicuously prominulent beneath; secondaries very slender, a-
bout 5 per side, arcuate-ascending, obscure above, slightly
prominulent and densely puberulent beneath; vein and veinlet
reticulation indiscernible above, the tertiaries only pro-
minulous beneath; inflorescence terminal, solitary, spicate;
peduncles short, 1--2 cm. long, puberulent-pulverulent;
spikes (immature) 6--11 cm. long, densely flowered; rachis
puberulent, shellowly sculptured in age, ridged; bractlets
lanceolate, about 1 cm. long, long-attenuate or caudate at
apex, puberulent; calyx tubular, about 12 mm. long, puberu-
lent, irregularly toothed at apex, the teeth 1--2 mm. long,
triangular, sherply acute; corolla hypocrateriform, blue,
the tube projecting 5 mm. or more from the calyx, glabrous.

The type of this very distinct species was collected by
Auguste Frangois Marie Glaziou (no. 13063) at Congonhas do
Campo, Minas Geraes, Brazil, in June or July between 1861
and 1895, and is deposited in the herbarium of the Jardin
Botanique de 1l'Etat at Brussels.

TECTONA GRANDIS var. GLABRIFOLIA Moldenke, var. nov.

Haec varietas a forma typica-speciei recedit foliis
subtus ut videtur glabris valde pallidis subargenteis.

This variety differs from the typical form of the species
in having the puberulence on the lower leaf-surface so
closely appressed and obscure as to impart to the leaves a
glabrous appearance except under the microscope. The lower
leaf-surfaces are also very pale, almost silvery.

The type of the variety was collected by John W. Gilles-
pie (no. 4544) on the overland trail to the other side of
the island, in the mountains south of Levuka, Ovalau, Fiji
Islands, at an altitude of 250 meters, on January 31, 1928,
and is deposited in the Britton Herbarium at the New York
Botanical Garden. The collector states that the plants were
doubtless introduced.

TITHYMALOFSIS IFPECACUANHAE f. LINZARIS Moldenke, f. nov.

Haec forma a forma typica speciei foliis linearibus rub-
ris recedit. - This form differs from the typical form of
the species in its linesr and red leaf-bledes.

The type was collected by me (no. 10478) in sand along a
roadside at Smithtown, Suffolk Co., New York, on May 29,
1938, and is deposited in the Britton Herbarium at the New
York Botanical Garden.

VERBENA HUNZIKERI Moldenke, sp. nove

Herba brachiata, ad basin sublignosa; caulibus procum-
bentibus; remis numerosis gracilibus adscendentibus irregu-
lariter laxeque piloso-pubescentibus glabrescentibus; peti-
Olis elongatis gracillimis dense patento-pubescentibus; lam-

322 PHYTOLOGIA Vol. 2, no. 8

inis deltoideo-ovatis, ad apicem attenuatis, ad basin trun-
catis, crasse dentatis supra parce pilosis subtus dense pil-
oso-pubescentibus; inflorescentiis terminalibus depresso-
spicatis multifloris.

Branching herb, somewhat woody at the base; stems procum-
bent; branches numerous, slender, ascending, irregularly and
loosely pilose-pubescent with whitish hairs of various
lengths and standing out almost at right angles to the
stems, glabrescent in age, tetragonal; nodes annulete; prin-
cipal internodes 1.5--5 cm. long; leaves decussate-opposite;
petioles elongate, very slender, 6--15 mm. long, densely
spreading-pubescent like the younger branches; blades thin-
chartacesous, rather uniformly bright-green on both surfaces
or slightly lighter beneath, deltoid-ovate, 1.8--3.5 cm.
long, 9--18 mm. wide, regularly narrowed from the broad
base to the attenuate apex, truncate at base, coarsely and
rather irregularly sharp-toothed along the margins with an-
trorse teeth, sparsely pilose above, densely pilose-pubes-
cent beneath, less densely so in age and the hair then most-
ly concentrated on the larger venation; midrib very slender,
plane above, slightly prominulous beneath; secondaries very
slender, 4--6 per side, ascending, only slightly arcuate,
obscure or very slightly subimpressed above, obscure or
slightly prominulous beneath; inflorescence terminal, de-
pressed-spicate, many-flowered; peduncles slender, sontinu-
ous with the stem, rather densely pilose-pubescent like the
stems, often with some gland-tipped hairs, 2--3 cm. long;
spikes up to about 3 cm. long and 3.5 cm wide in anthesis;
brectlets narrowly lanceolate, about 4 mm. long and 1 m.
wide, glabrate except for the long-ciliate margins, sharply
attenuate at apex; calyx tubular, about 8 mm. long, short-
pilose on 4 parallel ribs, otherwise subglabrate, the sharp-
ly acute teeth 1--1.5 mm. long, irregular; corolla-tube 1.3
--1.8 cm. long, more or less puberulent outside, its limb to
1.3 cm. wide, the lobes deeply cordate.

The type of this species was collected by Armando T. Hun-
ziker (no. 6812) -- in whose honor it is named -- in the al-
der formation along the highway between Alto del Clavillo
and Alpachiri, Tucuman, Argentina, on September 18, 1946,
and is deposited in the Britton Herbarium at the New York
Botanical Garden. The species is obviously related to V.
phlogiflora Cham. and V. incisa Hook. whose densely pubes-
cent calyxes at once distinguish them.

VSRBENA’ LINDBERGI Moldenke, sp. nov.

Herba; caulibus 1 m. altis brachiatis obtuse tetragonis
reflexo-hispidulis; ramis adscendentibus tetragonis sulcatis
hispidulis; petiolis 1--2 mm. longis breviter hispidulis vel
obsoletis; laminis chartaceis oblongo-lanceolatis acutis, ad

1947 Moldenke, New and Noteworthy Plants 523

basin abrupte acutis vel subtruncatis, remote serrulatis,

supra strigilloso-scabris, subtus secus venulis sparsiuscule

hispidulis; inflorescentiis spicatis paniculatis paucifloris
juventute conglobatis, senectute usque ad 1.5 cm. elongatis.

- Herb; stems 1 m. or more tall, branched above, obtusely
tetragonal, hispidulous with reflexed hairs about 1 mm.
long, wearing off at the base of the stem in age; branches
decussate-opposite, slender, ascending, tetragonal, sulcate
in drying, hispidulous like the stems; nodes annulate; prin-
cipal internodes 5.5--14 cm. long; leaves rather sparee,
decussate-opposite, sessile or subsessile; petioles, when
present, 1--2 mm. long, short-hispidulous; blades chartace-
ous, dark-green above, slightly lighter beneath, oblong-
lanceolate, 2--4 cm. long, 6--11 mm. wide, acute at apex,
abruptly acute or subtruncate at base, rather remotely ser-
rulate along the margins, the points of the appressed teeth
4--5 mm. apart on mature leaves, strigillose-scabrous with
whitish antrorse bulbous-based hairs above, rather sparsely
short-hispidulous along the venation beneath; midrib indis-
cernible above, very inconspicuously prominulous beneath,
very slender; secondaries very slender, about 3 per side,
ascending, indiscernible above, very indistinctly promim-
lous beneath; vein and veinlet reticulation indiscernible
above, obscure beneath; inflorescence spicate, terminating
each lateral branch and in a panicle of about 6 branches at
the apex of the stem; peduncles slender, tetragonal, sulc-
ate in drying, hispidulous, 1--7.5 cm. long, those termina-
ting the branches usually quite short; spikes rather few-
flowered, densely conglobate when young, elongating to a-
bout 1.5 cm. in fruit and the lower fruits then imbricate
but not especially densely so; bracts in the terminal pan-
icle foliaceous, lanceolate, 0.5--2 cm. long, 1--5 mm.
wide, attenuate to the sharp apex, short-hispidulous on
both surfaces, sessile, in decussate-opposite pairs; bract-
lets lanceolate, about 2.5 mm. long and 1 mm. wide, gparse-
ly puberulent, the margins often ciliolate, attenuate at
apex, about half as long as the fruiting-calyx; calyx tubu-
lar, 4--5 mm. long, short-toothed, spreading-puberulent
with antrorse hairs; corolla-tube about 5 mm. long, its
limb about 1.5 mm. wide.

The type of this species was collected by Gdsta A. Lind-
berg (no. 136) -- in whose honor it is named -- in swamps
at Ribeiro dos Buggris, in the neighborhood of Caldas, Min-
as Seraes, Brazil, on November 1, 1854, and is deposited in
the herbarium of the Jardin Botanique de 1'Etat at Brussels.
XVERBENA OSTENI Moldenke, hybr. nov.

Planta hybrida naturelis; caulibus gracilibus plusminus-
ve dense breviterque pubescentibus, novellis patento-pubes-

324 Po. Y -7-0.4:0 GK Vol. 2, now 8

centibus vel hirsutulis; pedunculis ca. 1.5 cm. longis vel .
subobsoletis dense patenti-hirtellis; capitulis dense pauci-
floris; bracteolis lanceolatis longe attenuatis ciliatis.

A natural hybrid between V. peruviana (L.) Britton and V.
pletensis Spreng., with intermediate characters; stems slen-
der, more or less densely short-pubescent, the younger parts
spreading-pubescent or hirtellous; petioles about 1 mm.
long, hirtellous; blades small, ovate, 1--1.5 cm. long, 4--9
mm. wide, coarsely dentate, pustulate-scabrous and very
sparsely or more densely strigose-hirsutulous above, scat-
tered-pubescent or hirsutulous beneath, especially on the
larger venation; peduncles about 1.5 cm. long or almost ob-
solete, densely spreading-hirtellous with hair of several
lengths; heads densely rather few-flowered; bractlets lance-
olate, 5--6 mm. long, long-attenuate, densely short-pubes-
cent, long-ciliate on the margins; calyx about 1 cm. long,
densely hirsutulous, irregularly apiculate; corolla light-
red, its tube glabrous, about 15 mm. long, its limb about 15
mm. wide.

The type of this natural hybrid wes collected by Cornel.
Osten -- in whose honor it is named -- between plants of the
perent species at Coquimbo, dept. Soriano, Uruguay, on Nov-
ember 16, 1894, and is deposited in the herbarium of the
Museo de Historia Natural at Montevideo.

VERBENA PERUVIANA f. ROSEA Moldenke, f. nov.

Haec forma a forma typica specieicorollis roseis recedit.
- This form differs from the typical form of the species in
having pink instead of bright-scarlet corollas.

The type was collected by Cornel. Osten at Arroyo Grande,
dept. Soriano, Uruguay, on October 3, 1895, and is deposited
in the herbarium of the Museo de Historia Natural at Monte-
video.

XANTHOXALIS EUROPAEA var. ATROPURPUREA (Planch.) Moldenke,
combe NOVe

Cxalis corniculata var. atropurpurea Planch., Fl. Serres
12: 47,'1857.

XYLOPHACOS PURSHII ver. INTERIOR (Me EB. Jones) Moldenke,
comb. nov.
Astragalus purshii ver. interior M. &. Jones, Astragalus
222. 1923.

XYLOPHACOS PURSHII var. LSUCOLOBUS (M. &. Jones) Moldenke,
combe nove
Astragalus purshii var. leucolobus M. E. Jones, Contrib.
West. Bot. 10: 61. 1902.

THE KNOWN GEOGRAPHIC DISTRIBUTION OF THE MEMBERS OF THE VER-
BENACEAE AND AVICENNIACBAE. SUPPLEMENT 6

Harold Ne Moldenke

Continued studies of herbarium material of the Avicennia-
ceae, Symphoremaceae, Stilbaceae, and Verbenaceae have
brought to light 481 new country or island records in these
groups, 443 new state, province, or depertment records, and
222 new county or parish records not previously recorded by
me in my tabulation of the known geographic distribution of
the accepted members of these groups [Moldenke, H. N., The
known geographic distribution of the members of the Verbena-
ceae and Avicenniaceae, 104 pp. 1942; Supplement 1, 4 pp.
1943; Supplement 2, in Bot. Gaz. 106: 158--164. 1944; Sup-
plement 3, in Castanea 10: 35--46. 1945; Supplement 4, in
Am. Journ. Bot. 32: 609--612. 1945; Supplement 5, in Bol.
Soc. Venez. Cienc. Nat. (in press)]. The specimens on which
these records are based will all be cited in my forthcoming
monographs of the genera involved, or in the supplements
thereto, but as it will probably be some years before these
generic monographs are all published, it has been thought
advisable to make the specific and varietal records avail-
able to students working on the flora of given areas. The
7,000 herbarium specimens on which these new records are
based are deposited in the herbaria of the New York Botani-
cal Garden, Missouri Botanical Garden, United States Natio-
nal Museum, Cornell University, New York State Museum, Uni-
versity of Texas, L. H. Bailey Hortorium, Jardin Botanique
de l'Etat at Brussels, Museo de Historia Natural at Monte-
video, University of Miami, Chicago Natural History Museum,
Gray Herbarium of Harvard University, Botaniska Trédgard at
Goteborg, Museo Paranagsnse at Curitiba, Universidad Nacional
de México at Mexico City, Instituto Miguel Lillo at Tucuman,
Princeton University, Jardin Botanico at Madrid, Naturhist-
oriska Riksmuseum at Stockholm, Instituto Darwinion at San
Isidro, Southern Methodist University, Vanderbilt Univer-
sity, and the private herbaria of Harry Ahles, Jose Bugenio
Leite, C. Le Lundell, and Mery Thais,

As in previous installments of these records, an asterisk
(*) following a name indicates that the plant is endemic to
that country or island, as far as now known.

CANADA :
Quebec:
Verbena hastata L. (Assomption County)
Nootka Island:
525

326 PHYTOLOGIA Vol. 2, no. 8

Verbena lasiostachys Link
Verbena robusta Greene

UNITED STATES OF AMERICA:

New Hampshire: .
‘Verbena hastata L. (Carroll County)
New York:
Verbena urticifolia var. leiocarpa Perry & Fernald
(Westchester County)
Pennsylvania :
Fhyla lanceolata (Michx.) Greene (Dauphin & Mifflin
Counties
Marylend:
Verbena hastata L. (Baltimore County.)
North Carolina:
Phyla nodiflora (L.) Greene (Iredell County)
South Caroline:
Fhyla lanceolata (Michx.) Greene (Berkeley County)
Verbena canadensis (L.) Britton (Abbeville County)

Verbena urticifolia var. leiocarpa Perry & Fernald :
Lexington County )
Georgia:

Callicarpa americana L. (Lowndes County)
Phyla nodiflora (L.) Greene (Glynn County)
Verbena hastata L. (Chatham County)
Florida:
Clerodendrum indicum (L.) Kuntze (Pinellas County)
Lantana Camara var. mista (L.) Le He Bailey (Pinellas & |
Polk Counties)
Lantana ovatifolia Britton (Martin County)
Phyla nodiflora (Le) Greene (DeSoto County)

Alabama:
Verbena bonariensis L. (Crenshaw County)
Ohio:

Phyla lanceolata (Michx.) Greene (Licking County)
Illinois:

Phyla lanceolata (Michx.) Greene (Hancock, Ferry, &

Whiteside Counties)

x Verbena illicita Moldenke (Cass County)

Verbena stricta Vent. (Bureau County)
Indiana :

Verbena bracteata Lag. & Rodr. (Madison County)

Verbena simplex Lehm. (Jay County)

Verbena urticifolia L. (Marshall & Vermillion Counties)
Iowa :

Verbena simplex Lehm. (Benton County)

Verbena urticifolia L. (Poweshiek County)
Kentucky:

Phyla lanceolata (Michx.) Greene (Jefferson County )

x Verbena Blanchardi Moldenke (Warren County)

/

1947 Moldenke, Geographic Distribution 3 327

x Verbena Engelmannii Moldenke -- to be deleted
Tennessee :
Verbena simplex Lehm. (Blount County)
Verbena urticifolia L. (Lewis County)
Michigan:
Verbena hastata L. (Eaton County)
Wisconsin: °
x Verbena Deamii Moldenké (Pierce County)
Minnesota :
Verbena hastata L, (Meeker & Yellow Medicine Counties)
Verbena simplex Lehm. (Rock County)
South Dakota :
Verbena bracteata Lag. & Rodr. (Roberts County)
Verbena hastata L. (Devel & Tripp Counties)
Verbena stricta Vent. (Devel County)
Kansas:
x Verbena Perriana Moldenke (Sedgwick County )
Verbena Trightii A. Gray (Finney County)
Missouri:
Phyla lanceolata (Michx.) Greene (Greene County)
Louisiana:
Verbena brasiliensis Vell. (Tensas Parish)
Verbena canadensis (L.) Britton (Orleans Parish)
Verbena Halei Small (Plaquemines Parish)
Nevada: |
Aloysia Wrightii (A. Gray) Heller (Clark County)
Colorado:
Verbena ambrosifolia Rydb. (Weld County)
Verbena stricta Vent. (Zl Paso County)
Verbena Wrightii A. Gray (Boulder County)

Nebraska :
Verbena stricta Vent. (Oteo County)
Oklahoma :

Verbena bracteata Lag. & Rodr. (Garvin County)
Verbena canadensis (L.) Britton (Garvin County )
Texas:
Aloysia ligustrina (Lag.) Small (Atascosa, Coke, Gilles-
pie, Pape. & zavalla Counties)
Aloysia ligustrina var. Schulzii (Standl.) Moldenke (Jim
Hogg, Jim Wells, Kinney, & Zapate Counties)

Aloysia Wrightii (A. Gray) Heller (Coke & Live Oak Coun-
ties)

Callicarpa americana L. (Freestone, Hardin, & Shelby

Counties )

Callicarpa americana var. lactea F. J. Muller (Jasper
perce

Citharexylum brachyanthum (A. Gray) A. Gray (Zapata
Sines)

Lantana horrida H.B.eK. (Karnes, Kenedy, & Tarrant Count-

328

PHYTOLOGIA Vol. 2, no. 8

ies) |

Lantana macropoda Torr. (Bexar, Jim Hogg, & La Salle
Counties)

Phyla cuneifolia (Torr.) Greene (Dallam, Hale, & Lubbock
Counties

Phyla incisa Small (Calhoun, Frio, Kenedy, La Salle,
Mitchell, Palo Pinto, & Williamson Counties)

Phyla lanceolata (Michx.) Greene (Bowie, Colorado, Don-
ley, Kaufman, & Matagorda Counties)

Phyla nodiflora (L.-) Greene (Hardin & McLennan Counties)

Phyla nodiflora var. reptane (H.B.K.) Moldenke (Val Ver-
de County

Phyla strigulosa (Mart. & Gal.) Moldenke (Cameron, Hid-
algo, & Wood Counties)

Phyla strigulosa var. parvifolia (Moldenke) Moldenke

Burnett, Hidalgo, Presidio, Starr, & Uvalde Counties )

Phyla yucatana Moldenke -- to be deleted

Phyla yucatana var. parvifolia Moldenke -- to be deleted

Tetraclea Coulteri A. Gray (£1 Paso & Tom Green Counties)

Tetraclea Coulteri var. angustifolia (Woot. & Standl.)
A. Nels. & Macbr. (Brewster County)

Verbena ambrosifolia Rydb. (Reeves County).

Verbena bipinnatifida Nutt. (Andrew, Bastrop, Borden,

Deaf Smith, Hardman, Jefferson, Mason, McLennan, Rand-
all, & San Patricio Counties)

Verbena bonariensis L. (Jasper County)

Verbena bracteata Lag. & Rodr. (Bowie, Comal, Hale, Hud-
speth, Mitchell, Swisher, & Wood Counties)

Verbena brasiliensis Vell. (Hardin, Jefferson, Orange, &
Wharton Counties)

Verbena canadensis (L.) Britton (Jasper & Upshur Coun-
ties)

Verbena canescens var. Roemeriana (Scheele) Ferry (Cole-
man & Guadalupe Counties

Verbena ciliata Benth. (Brooks & MacMullen Counties)

Verbena Cloveri Moldenke (LaSalle County )

Verbena Cloveri var. lilacina Moldenke -=- to be deleted

Verbena Halei Small (Aransas, Brooks, Coleman, Jeffer-
son, & Wharton Counties)

Verbena Matthesii Turez. -- to be deleted

Verbena perennis Wooton (Hudspeth County)

Verbena plicata Greene (Andrews, Atascosa, Jim Wells,

. Sterling, Uvalde, Wilbarger, & Winkler Counties)

Verbena pumila Rydb. (Baylor, Ector, Jack, Kinney, Lub-
bock, Medina, Montague, Randall, Red River, Ward, Wi-
chita, & Wilbarger Counties)

Verbena quadrangulate Heller (Willacy County)

Verbena Runyoni Moldenke (Brazoria County)

Verbena scabra Vahl (Hardin County)

1947 Moldenke, Geographic Distribution 329

Verbena tenuisecta Briq. (Angelina County)

Verbena urticifolia L. (Wise County)

Verbena urticifolia var. leiocarpa Perry & Fernald (Cass
County

Verbena Wrightii A. Gray (Hudspeth County)

Verbena xutha Lehm. (Bastrop, Calhoun, Ellis, Fayette, &
Hardin Counties)

Vitex ees var. heterophylla (Franch.) Rehd. (Brazos

County
Vitex Negundo var. incisa (Lam.) C. B. Clarke -- to be

deleted

New Mexico:

Tetraclea Coulteri var. angustifolia (Woot. & Standl.)
A. Nels. & Macbr. (Eddy County
Verbena bipinnatifida Nutt. (Dona Ana County)
Verbena bracteata Lag. & Rodr. (Baca County)
Verbena Halei Small (Otero County)
Verbena plicata Greene (Eddy County)
Verbena Wrightii A. Gray (Luna County)
Arizona:
Lantana macropoda Torr. (Santa Cruz County)
Verbene canadensis (L.) Britton -- to be deleted
Verbena carolina L. (Gila County)
Verbena ciliata var. pubera (Greene) Perry (Gila County)
Verbena plicata Greene (Yavapai County)
Washington: ‘
Verbena hastata L. (Clarke County)
Oregon:
Verbena bracteata Lag. & Rodr. (Coos, Hood River, &
Sherman Counties)
Verbena hastata L. (Columbia County)

Verbena lasiostachys var. scabrida Moldenke (Curry &
Jackson Saartame

California:

Lantana Camara var. mista (L.) L. He Bailey (San Diego
County

Fhyle incisa Small (Fresno & Imperial Counties)

Phyla lanceolata (Michx.) Greene (Contra Coste County)

Phyla nodiflora (L.) Greene (Fresno County)

Phyla nodiflora var. canescens (HeB.K.) Moldenke (Imper-
ial, Kern, Merced, & Yolo Counties)

Phyla nodiflora var. reptans (H.BeK.) Moldenke (Sacramen-
to County

Phyla nodiflora var. rosea (D. Don) Moldenke (Amador
County

Verbena brasiliensis Vell. (Solano County)

Verbena lasiostachys f. albiflora Moldenke (San Matec
County )*

Verbena lasiostachys var. scabrida Moldenke (Alameda, Co-

550 PHYTOLOGIA Vol. 2, no. 8

lusa, San Mateo, & Siskiyou Counties); delete the "*"
Verbena lasiostachys var. septentrionalis Moldenke (Ama-
dor, Del Norte, Nevada, San Benito, Santa Barbara,
Shasta, Trinity, & Tulare Counties)
Verbena menthaefolia Benth. (Riverside County)
Verbena officinalis L. (Amador County)
Verbena robusta Greene (Merced & San Benito Counties)
MEXICO:
Aloysia barbata (T. S. Brandeg.) Moldenke --. delete Chi-
apas
Aloysia chiapensis Moldenke (Chispas)* ;
Bouchea prismatica (L-) Kuntze (Michoacan)
Burroughsia appendiculata (Robinson & Greenm.) Moldenke
(San Luis Potos
Citharexylum Alteamiranum Greenm. (Hidalgo)
Citharexylum Berlandieri B. L. Robinson (Oaxaca)
Citharexylum flabellifolium S. Wats. (Carmen Island)
Citharexylum Schottii Greenm. -- delete the "*"
lantana bipinnatifida Sessé & Moc. -- to be deleted
EE ETE var. aculeata (L.) Moldenke (Colima & Sina-
loa
Lantana frutillea Moldenke (Guanajuato)
Lantana hispida H.B.K. (Chiapas)
Lantana involucrata L. (Socorro Island)
Lantana origanoides Mart. & Gal. -- to be deleted
Lantana repens Sesse & Moc. -- to be deleted
Lantana acorta Moldenke (Oaxaca & Sinaloa)
Lantana trifolia L. (Chiapas)
Lantana tuxtlensis Seseé & Moc. -- to be deleted
Lantana velutina Mart. & Gal. (Tlaxcala)
Lippia alba (Mill.) N. E. Br. (Guanajuato)
Lippia practeosa (Mart. & Gal.) Moldenke (Chiapes, Jalis-
co, Oaxaca, & Puebla)*
Lippia callicarpaefolia H.eBeK. (Veracruz)
Lippia graveolens H.eB.Ke (Chiapas & Tamaulipas)
Lippia hypoleia Briq. (Oaxaca)
Lippia nutans Be Le Robinson & Greenm. -- to be deleted
Lippia Palmeri S. Wats. (Magdalena Island)
Lippie Palmeri var. spicata Rose (Sonora & Tiburon Island)
Lippie substrigosa Turcz. (Tabasco)
Lippia umbellata Cav. -- delete Sonora
Phyla incisa Small (Baja California, Federal District, &
Sinaloa)
Fhyla lanceolata (Michx.) Greene (Sinaloa)
Fhyla nodiflore (L.) Greene (Nayarit)
Phyla scaberrima (A. Le Juss.) Moldenke (Campeche & Vera-
cruz
Phyla strigulosa (Mart. & Gal.) Moldenke (Campeche, Guer-
rero, Michoacan, Nuevo Leon, San Luis Potos{, Vera-

1947 Moldenke, Geographie Distribution 331

cruz, & Yucatan)
Fhyla strigulosa var. parvifolia (Moldenke ) Moldenke (Hid-
algo, Jalisco, Mexico, Michoacan, Morelos, Nuevo Leon,
San Luis Potos{, Sinaloa, & Tamaulipas)
Phyle yucatana Moldenke -- to be deleted
Phyla yucatana var. rvifolia Moldenke -- to be deleted
Prive grandiflora (Ort.) Moldenke (Oaxace )
Verbena canescens HeB.K. (México)
Verbena carolina L. (Querétero)
Verbena ciliata Benth. (San Pedro Nolasco Island)
Verbena elegans H.B.K. (Michoacan)
Verbena gracilis Desf. (Guanajuato)
Verbena longifolia Mart. & Gal. (Coahuila)
Verbena neomexicana (A. Gray) Small (Sonora)
Verbena neomexicana var. hirtella Perry (Baja California)
Verbena perennis var. Johnstoni Moldenke (Coahuila)
Verbena Wrightii A. Gray (San Luis Fotosf)
GUATEMALA 3
Citharexylum hirtellum Standl. (Izabal)
Lantana trifolia L. (Retalhuleu)
Phyle stoechadifolia (L.) Small (Guatemala)
Phyla strigulosa (Mart. & Gel.) Moldenke (Alta Verapaz, El
Feten, & Izabal)
Phyla yucatana Moldenke -- to be deleted
BRITISH HONDURAS :
Phyla strigulosa (Mart. & Gal.) Moldenke
Fhyla yucatane Moldenke -- to be deleted
HONDURAS :
Phyla strigulosa (Mart. & Gal.) Moldenke
Phyla yucatana Moldenke -- to be deleted
COSTA RICA:
Citharexylum Cooperi Standl. (Guanacaste)
Citharexylum Schottii Greenm. (Alajuela)
Cornutia grandifolia var. normalis (Kuntze) Moldenke (Ala-
juela & Guanacaste )
Lantana canescens H.B.K. (Guanacaste )
Lantana costaricensis Hayek (Heredia)
Lantana glandulosissima Hayek (San José)
Lantana hirta Grah. (Heredia)
Lantana montevidensis (Spreng.) Briq. (San Jose)
Lantana trifolia L. (Puntarenas & San José)
Lippia alba (Mill.) Ne. E. Br. (Guanacaste & San José)
Lippia Brenesii Standl. -- to be deleted
Lippia cardiostegia Benth. (Heredia & San José)
Lippia controversa Moldenke (Puntarenas)
Lippia graveolens H.B.K. (Guanacaste )
Lippia liberiensis Moldenke (Guanacaste )*
Petrea volubilis var. pubescens Moldenke (Ala juela)
Phyle nodiflora var. longifolia Moldenke (Limon)

332 PHYTOLOGIA Vol. 2, no. 8

Phyla scaberrima (A. L. Juss.) Moldenke (Ale juela)
Priva aspera H.BeK. (Alajuela)
Priva lappulecea (L.) Pers. (Alajuela & Guanacaste )
Rehdera trinervis (Blake) Moldenke (Ala juela)
Stachytarpheta Calderonii Moldenke (Guanacaste )
Verbena litoralis H.B.K. (Alajuela, Heredia, & Puntarenas)
Verbena rigida Spreng. (Cartago)
PANAMA:
‘Clerodendrum molle H.B.K. (Panama)
Lantana Camara var. aculeata (L.) Moldenke (Taboga Island)
Phyle betulaefolia (H.B.K.) Greene (Panama )
Phyla nodiflora var. longifolia Moldenke (Manzanillo Is-
land
Phyle strigulosa (Mart. & Gal.) Moldenke (Panamé )
BAHAMAS:
Fhyla strigulosa var. parvifolia (Moldenke) Moldenke
Phyla yucatana var. parvifolia Moldenke-- to be deleted
CUBA : !
Duranta Fletcheriana Moldenke (Pinar del Rfo)
Phyle strigulosa (Mart. & Gal.) Moldenke (Havane, Las Vil-
las, Matanzas, & Oriente)
Phyla strigulosa var. parvifolia (Moldenke ) Moldenke (Hav-
an a, Matanzas, & Oriente
Phyla yucatana Moldenke -- to be deleted
Phyla yucatana var. parvifolia Moldenke -- to be deleted
JAMAICA:
Phyla nodiflora var. reptans (H-BeK.) Moldenke -- to be
deleted
Phyla strigulosa var. parvifolia (Moldenke) Moldenke
Phyla yucatana var. parvifolia Moldenke -- to be deleted
Verbena brasiliensis Vell.
TORTUE:
Lantana montevidensis (Spreng.) Briq.
HISPANIOLA :
Lantana Ehrenbergiana Moldenke (Dominican Republic )*
Lantana exarata Urb. & Ekm. (Dominican Republic)
Lantana montevidensis (Spreng.) Briq. (Dominican Republic }
delete Haiti
Phyla strigulosa (Mart. & Gal.) Moldenke (Dominican Repub-
lic
Phyla strigulosa var. parvifolia (Moldenke) Moldenke (Dom-
inican Republic) |
Phyla yucatana Moldenke -~ to be deleted
| PUERTO RICO:
Phyla strigulosa var. parvifolia (Moldenke) Moldenke
Phyla yucatana var. parvifolia Moldenke -- to be deleted
ST. THOMAS:

Stachytarpheta cayennensis (L. C. Rich.) Vehl
ST. JOHN:

| 1947 Moldenke, Geographic Distribution | 333

Lantana arida Britton
ST. CROIX:
Lentana aride Britton
Phyla nodiflora var. reptana (H.BeK.) Moldenke -- to be
deleted
Phyla strigulosa (Mart. & Gal.) Moldenke
Phyla strigulosa var. parvifolia (Moldenke) Moldenke
Phyla yucatana var. parvifolia Moldenke -- to be deleted
ST. MARTIN:
Lantana arida Britton
Phyla nodiflora var. reptans (HeB.K.) Moldenke -- to be
deleted
Phyla strigulosa var. parvifolia (Moldenke) Moldenke
Phylea yucatana var. parvifolia Moldenke -- to be deleted
GUADELOUPE:
Durante repens L.
Phyle 8° strigulosa var. parvifolia (Moldenke) Moldenke
Phyla yucatana var. parvifolia Moldenke -- to be deleted
DOMINICA:
Lantana radula Sw.
Phyla nodiflora var. reptans (H.eBeK.) Moldenke -- to be
deleted
Phyla strigulosa var. parvifolia (Moldenke ) Moldenke
Phyla yucatana var. pervifolia Moldenke -- to be deleted
MARTINIQUE:
Lantana involucrata var. odorata (L.) Moldenke

Phyla strigulosa var. parvifolia (Moldenke) Moldenke
Phyla yucatane var. parvifolia Moldenke -- to be deleted
BARBADOS :
Phyla nodiflora var. reptans (H.B.K.) Moldenke -- to be
deleted
Phyle strigulosa var. parvifolia (Moldenke) Moldenke
Phyle yucatana var. parvifolia Moldenke -- to be deleted
ST. VINCENT:
Lantana radula Sw.:
Phyla nodiflorea var. reptans (H.BeK.) Moldenke -- to be
deleted

Phyle strigulosa var. parvifolia (Moldenke) Moldenke

Phyla yucetana var. parvifolia Moldenke -- to be deleted
GRENADA:

Lantana Camara var. mista (L.) Le He Bailey
Lantana involucrata var, odorata (L.) Moldenke
TRINIDAD:
Lantana redula Sw.
Phyla nodiflore ver. rosea (D. Don) Moldenke -- to be
deleted
Phyla strigulosa var. parvifolia (Moldenke) Moldenke
COLOMBIA :
Aegiphila bogotensis (Spreng.) Moldenke (El Valle)

/
534 PE YyhO L0GrA Vol. 2, noe &
‘Aegiphila farinoea Moldenke (El Valle)* r
Amasonia lasiocaulos Mart. & Schau. (Vaupes)
Citharexylum subflavescens Blake (El Valle)
Lantana foetida Rusby -- to be deleted
Lantana glutinosa Poepp. (El Valle & Tolima)
Lantana salicifolia H.eBeK. (El Valle)
Lippia Briquetii Moldenke (Cundinamarca )
Lippia hirsuta L. f. (El Valle)
Stachytarpheta straminea Moldenke (El Valle)
VENEZUELA:
Aegiphila bogotensis (Spreng.) Moldenke (Lara)
Aegiphila Fendleri Moldenke (Federal District)
Aegiphila laxiflora Benth. (Monagas)
Aegipnila membrenacea Turcz. (Merida )
Aegiphila mollis HeB.eK. (Merida)
Aegiphila perplexa Moldenke (Monagas)
Aegiphila quinduensis (H.B.K.) Moldenke (Monagas)
Aegiphila roraimensis Moldenke (Bolfvar)
Aegiphila Steyermarkii Moldenke (Bolfvar)*
Aegiphila Steyermarkii var. obtusifolia Moldenke (Bolfvar}
Aegiphila venezuelensis Moldenke (Bol{var)*
Aegiphile verrucosa Schau. (Federal District)
Callicarpa acuminata H.B.K. (Trujillo)
Citharexylum spinosum L. (Bolfvar)
Citharexylum subflavescens Blake (Mérida)
Cornutia microcalycina Pavon & Moldenke (Mérida)
Duranta Steyermarkii Moldenke (Monagas)*
Lantana canescens H.B.K. (Monagas)
Lantana foetida Rusby -- to be deleted
Lantana glutinosa Poepp. (Aragua, Bolfvar, & Cojedes)
Lantana. maxima Hayek (Federal District)
Lantana velutina Mart. & Gal. (Carabobo)
Lippia hirsuta L. f. (Tachira)
Petrea arborea H.B.K. (Lara)
Stachytarpheta australis Moldenke (Cojedes)
Vitex capitata Vahl (Apure & Monagas)
Vitex triflora Vahl (Amazonas)
BRITISH GUIANA:
Aegiphila roraimensis Moldenke -- delete the "*"
ECUADOR:
Aegiphila alba Moldenke (Azuay)
Aegiphila Hoehnei var. puyensis Moldenke (Oriente )*
Clerodendrum fragrans var. pleniflorum Schau. (Loja)
Clerodendrum Ulei Moldenke Gaitiacotzansa)
Cornutia microcalycina Pavon & Moldenke (£1 Oro)
Duranta Sprucei Briq. (El Oro, loja, & Pichincha)
Lantana Moritziana Otto & Dietr. (El Oro)
Lantana rugulosa H.eB.Ke (El Oro & Loja)
Phyla betulaefolia (H.B.K.) Greene (El Oro)

1947 Moldenke, Geographic Distribution 5355

Phyla nodiflora var. reptans (HeBeK.) Moldenke (Loja)
Phyla strigulosa (Mart. & Gal.) Moldenke (Guayas)
Stachytarpheta cayennensis (L. C. Rich.) Vahl (Guayas)
Stachytarpheta jamaicensis (L.) Vahl (Guayas)

Stachytarpheta Steyermarkii Moldenke (Loja)*
Stachytarpheta straminea Moldenke (Loja)

Verbena bonariensis L. (Pichincha)

Verbena crithmifolia Gill. & Hook. (Guayas)

Verbena laciniata (L.) Briq. (Azuey)

Verbena litoralis H.B.K. (Loja)

Verbena microphylla H.B.K. (Azuay)

GALAPAGOS ISLANDS:

Lantana peduncularis Anderss. (Abingdon, Albemarle, Barr-
ington, Charles, Chatham, Gardner, North Indefati-
gable, & Tower)

Lippia.rosmarinifolia Anderss. (Abingdon & Albemarle)

Lippia salicifolia Anderss. (Charles)

Phyla strigulosa (Mart. & Gal.) Moldenke (Duncan & Hood)

Friva leppulacea (L.) Pers. (Charles)

PERU :

Aloysia scorodonioides (H.B.K.) Cham. (Lima)

Clerodendrum molle H.BeK.

Junellia juniperina var. grisea (I. M. Johnst.) Moldenke

Speen

Junellia minima (Meyen) Moldenke (Moquegua )

Lantana Camara var. mista (L.) L. H. Bailey (Lima)

Lantana foetida Rusby -- to be deleted

Lantana glutinosa Poepp. (Apurimac, Cuzco, Libertad, &
Tima

Lantana Haughtii Moldenke (Jun{n)

Lantana scabiosaeflord H.BeK. (Libertad)

Lippia alba (Mill. ) N. E. Br. (Libertad )

Phyla nodiflora (L.) Greene

Verbena bonariensis L.

Verbena clavata Ruiz & Pav. -- delete the "*!

Verbena litoralis H.B.eK. (Libertad )

Verbena Matthewsii Briq. (Loreto)

BRAZIL:

Aegiphila casseliaeformis Schau. (Rio de Janeiro)

Aegiphila integrifolia (Jacq.) Jacks. -- delete Goyaz

Aegiphila Sellowiana Cham. -- delete Mattogrosso

Aegiphila Surfaceana Moldenke (Para)

Aloysia chamaedryfolia Cham. (Rio Grande do Sul)

Aloysia ligustrina var. paraguariensis (Briq.) Moldenke
(Parana )
Aloysia virgata var. elliptica (Briq.) Moldenke (Minas

Geraes

Aloysia virgata var. pletyphylla (Briq.) Moldenke (Santa
Catherina )

\

5336 PHYTOL OGTs Vol. 2, no. 8

Amasonia lssiocaulos Mart. & Scheu. -- delete the "*"

Durante Benthami Briq. (Minas Gerees)

Durante repens L. (Minas Geraes)

Lantana aristeta (Schau.) Briq. (Goyaz & Paiuhy)

Lantana aristata ver. glebrescens Pilger (Mattogrosso)*

Lantana brasiliensis Link (Mattogrosso)

Lantana Camara var. aculeata (L.) Moldenke (Rio de Jarsiro)

Lantana Camara var. nivea (Vent.) L. H. Bailey (Bahia)

Lantana canescens var. integrifolia Moldenke (Rio de Jan-
eiro

Lantana foetida Rusby -- to be deleted

Lantana glutinosa Poepp. (Bahia, Minas Geraes, Para, Rio
Grande do Sul, & S&o Paulo)

Lantana hypoleuca Briq. (Rio Grande do Sul & Rio de Jan-
eiro)

Lantana macrophylla (Cham.) Schau. is the correct form
for this name

Lantana minasensis Moldenke (Rio de Janeiro)

Lantana Pohliana Schau. (Bahia, Minas Geraes, & Rio de
Janeiro

Lantana radula Sw. (Pernambuco, Santa Catharina, & Sao
Paulo); delete the "*"

Lantana Riedeliana Schau. (Rio de Janeiro)

Tantana trifolie L. (Farana)

Lippie Arechavaletae Moldenke (Parana)

Lippie attenuata Mart. -- to be deleted

Lippia hermannioides Cham. -- to be deleted

Lippia iodophylla Schau. (Rio de Janeiro)

Lippia microcephala Cham. (Goyaz & Rio Grande do Sul)

_ Tippia trachyphylle Briq. (Farand)

Fetrea macrostachya Benth. (Pard)

Petrea maynensis Huber (Amazonas )

Petrea rugosa H.BeK. -- to be deleted

Stachytarpheta australis Moldenke (Maranhao & Rio Grande
do Sul)

“Stachytarpheta confertifolia Moldenke (Minas Geraes )*
Stachytarpheta gesnerioides var. cuneata Schau. (Goyaz )
Stachytarpheta glauca Cham. (Minas Geraes)

Stachytarpheta lactea Schau. (Minas Geraes & Rio de Jan-
eiro)

Stachytarpheta Maximiliani var. glabrata Schau. (Rio
Grande do Sul
Verbena alata Cham. (Parana)
Verbena ephedroides Cham. (Minas Geraes)
Verbena hispida Ruiz & Pav. (Sao Paulo)
BOLIVIA:
Acantholippia deserticola (R. A. Phil.) Moldenke (Fotos{)
Aloysia Fiebrigii (Hayek) Moldenke (Sucre)
Aloysia ligustrina var. paraguariensis (Briq.) Moldenke

-

1947 Moldenke, Geographic Distribution 337

(Tarija)

Aloysia scorodonioides var. detonsa (Briq.) Moldenke (Chu-
quisaca

Aloysia virgata var. elliptica (Briq.) Moldenke (Chaco &
Santa Cruz)

Aloysia virgata var. pletyphylle (Briq.) Moldenke (La Faz)
Dureanta Benthami Briq. -- delete the "*"

Junellia bisulcata (Hayek) Moldenke (Potos{)

Junellia uniflora (R. A. Phil.) Moldenke (Potosf)

Lampaya Castellani Moldenke (Potosf)

Lantana Fiebrigii Hayek (Cochabamba & Le Paz)

Lantana foetida Rusby -- to be deleted

Lantana glutinosa Poepp. (Cochabamba, El Beni, & La Paz)

Lantana ovata Hayek -- delete the "*" .
PARAGUAY:

Aloysia virgate var. platyphylla (Briq.) Moldenke

Lantana foetida Rusby -- to be deleted

Lantana glutinosa Poepp.

Lantana montevidensis (Spreng.) Briq.

Lippia bothrioura Briq. -- delete the "*"

Lippia trachyphylla Briq. -- delete the "*"

fs rigida var. obovate (Hayek) Moldenke delete the

Verbena storeoclada Briq. -- delete the "*"
URUGUAY:

Aloysia ligustrina var. paraguariensis (Briq.) Moldenke
Aloysia virgata var. elliptica (Briq.) Moldenke

Lantana Camara ver. aculeata (L.) Moldenke
Lantana foetida Rusby -- to be deleted
Lantana glutinosa FPoepp.
Lantana ovata Hayek
Lippia bothrioura Briq.
Phyla nodiflora (L.) Greene
Verbena corymbosa Rufz & Pav.
Verbena megapotamica Spreng.
x Verbena Osteni Moldenke*
_ Verbena peruviana f. rosea Moldenke*
Verbena rigida var. obovata (Hayek) Moldenke
Verbena storeoclada Brig.
CHILE:
Aloysia Fonckii (R. A. Phil.) Moldenke (Atacama )
Aloysia Reichii var. trilobata Moldenke (Coquimbo)*
Diostea scoparia (Gill. & Hook.) Miers (Aconcagua & Col-
chague
Junellia spathulata (Gill. & Hook.) Moldenke (Aconcagua &
Curico |
Junellie uniflora (R. A. Phil.) Moldenke (Santiago)
lantana Cummingiana Hayek -- to be deleted
lantana glutinosa Poepp.

338 PHYTOLOGIA Vol. 2, no. 8

Phyla nodiflora var. rosea (D. Don) Moldenke (Coquimbo )
Verbena clavate Ruiz & Pav.
Verbena concepcionis Moldenke (Concepcion)*

Verbena hispida Ruiz & Pav. (Colchague)

Verbena litoralis H.BeK. (Coquimbo)

ARGENTINA:

Acantholippia deserticola (R. A. Phil.) Moldenke (Jujuy)

Acantholippia hastulata Griseb. (Salta)

Aloygia chameedryfolia Cham. (Misiones)

Aloysia pulchra (Briq.) Moldenke (Misiones) |

ARogty soars toss var. detonsa (Briq.) Moldenke (Ju-
juy

Aloysia ternifolia Moldenke (Sante Cruz)*

Aloysia virgata var. elliptica (Briq.) Moldenke (Chaco,
Corrientes, Formosa, Santiago del Estero, & Tucumaén)

Aloysia virgata var. platyphylle (Briq.) Moldenke (Chaco,
Jujuy, Misiones, Santiago del Estero, & Tucumén)

Junellia digitata (R. A. Phil.) Moldenke (Salta)

Junellia juniperina (Lag.) Moldenke (Jujuy & La Rioja)

Junellia longidentata Moldenke (Salta)

Junellia minima (Meyen) Moldenke (Jujuy)

Junellia patagonica (Speg.) Moldenke (Sante Cruz)

Junellia seriphioides (Gill. & Hook.) Moldenke (Jujuy)

Junellia serpyllifolia (Speg.) Moldenke (Chubut)

Junellia Silvestrii (Speg.) Moldenke (Santa Cruz)

Junellia Struthionum (Speg.) Moldenke (Neuquen)

Junellia thymifolia (Lag.) Moldenke (Chubut & Santa Cruz}

Junellia tridactylites (Lag.) Moldenke (Chubut)

Junellia Wilczekii (Briq.) Moldenke (Santa Cruz)

Lampaya Castellani Moldenke -- delete the "*"

Lantana achyranthifolia Desf. (Buenos Aires)

Lantana cordobensis Moldenke (Buenos Aires)

Lantana foetida Rusby -- to be deleted

Lantana glutinosa Poepp. (Buenos Aires, Formosa, Mision-
es, Salte, & Tucuman)

Lantana Junelliana Moldenke (Buenos Aires)

Lantana montevidensis (Spreng.) Briq. (Buenos Aires)

Lippia turbinata Griseb. (Buenos Aires)

Stachytarpheta australis Moldenke (Misiones)

Stachytarpheta cayennensis f. albiflora Moldenke (Misio-
nes) ”

Verbena dissecta Willd. (La Pampa)

Verbena ephedroides Cham. (Misiones)

Verbena flava Gill. & Hook. (Chubut)

Verbena gracilescens (Cham.) Herter (Mendoza)

Verbena Hookeriana (Covas & Schnack) Moldenke (Buenos
Aires, Catamarca, Cordoba, La Pampa, La Rioja, Rfo
Negro, & San Luis)

Verbena Hunzikeri Moldenke (Tucuman)*

a Oe at

1947 Moldenke, Geographic Distribution 339

Geebene. Neoi Moldenke (Buenos Aires )*

Verbena scrobiculata Griseb. (Cordoba & San Luis)

Verbena tenera Spreng. (La Rioja)

Verbena tristachya Troncoso & Burkart (Corrientes & Entre

Rios )*
MADEIRA:
Verbena rigida Spreng.
CANARY ISLANDS:
Phyle nodiflora (L.) Greene (Tenerife )
Verbena rigida Spreng. (Tenerife)

BELGIUM :
Fhyle nodiflora (L.) Greene
FRANCE :

Verbena officinalis var. prostrate Gren. & Godr.
PORTUGAL:
Verbena supine L.
SPAIN:
Lantana Camara var. aculeata (L.) Moldenke
Verbena supina L.
BALEARIC ISLANDS:
Phyle nodiflora var. rosea (D. Don) Moldenke (Majorca &
Minorca
Vitex Agnus-castus L. (Majorca & Minorca)
GREECE:
Verbena officinalis L.
CYPRUS:
Vitex Agnus-castus L.
SICILY:
Phyla nodiflora var. reptans (HeBeK.) Moldenke
ALBANIA:
Verbena officinalis L.
ALGERIA:
Phyla nodiflora var. suborbicularis (L. Chev.) Moldenke
Verbena officinalis var. macrostachya Benth.
Verbena supina f. erecta Moldenke
CYRENAICA:?
Verbena supina L.
TRIPOLITANIA:
Verbena officinalis L.
EGYPT:
Lantana montevidensis (Spreng. ) Briq.
Phyle nodiflora var. canescens (H.B.K.).Moldenke
Phyla nodiflora var. suborbicularis (L. Chev.) Moldenke
Verbena supine - supine f. erecta Moldenke
FRENCH WEST AFRICA:
Chascanum marrubiifolium Fenzl (Tibesti)
Lippia adoénsis Hochst. -- to be deleted
Lippia Chevalierii Moldenke (French Soudan & Senegambia)
Verbena officinalis L. (Mauritania)

340 PH Fh OL O Gila Vol. 2, no. 8

ANGLO-EGYPTIAN SUDAN:
Verbena supina f. erecta Moldenke (Nubia)
SRITREA:
Lantana kisi A. Rich.
Phyla nodiflora var. suborbiculeris (L. Chev.) Moldenke
FATMAH ISLAND:
Phyla nodiflora (L.) -Greene
FRENCH GUINEA:
Lippia adoSnsis Hochst. -- to be deleted
Lippia Chevalierii Moldenke
SIERRA LEONS:
Lantana Camara var. aculeata (L.) Moldenke
Stachytarpheta jamaicensis (Le) Vahl
GOLD COAST:
Lippia ado8nsis Hochst. -~ to be deleted
Lippie rugosa A. Chev.
TOGOLAND:
Lantana Camara Le
Lippia adosénsis Hochst. -- to be deleted
Lippia rugosa A. Chev.
FRENCH EQUATORIAL AFRICA:
Lantana Camara var. aculeata (Le) Moldenke (Gabun)
Lippie rugosa A. Chev. (Ubangi-chari)
Phyla nodiflora (L.) Greene (Chad Territory)
Stachytarpheta angustifolia (Mill.) Vahl (Ubangi-chari)
BELGIAN CONGO:
Lantana Camara var. aculeata (L.) Moldenke
Lantana Mearnsii var. congolensis Moldenke
Lantana Mearnsii var. latibracteolata Moldenke
Lantana viburnoides Vahl
Lippia adoensis Hochst. -- to be deleted
Lippie grandifolia Hochst.
Lippia javanica (Burm. f.) Spreng.
Lippie rugosa A. Chev.
Lippia Whytei Moldenke
Phyla nodiflora (L.) Greene
Phyla nodiflora var. reptans (H.B.K.) Moldenke
Verbene officinalis L.
UGANDA PROT=CTORATE:
Lantana Mearnsii var. latibracteolata Moldenke
Lippia javanica (Burm. f.) Spreng.
Lippia Whytei Moldenke
TANGANYIKA TERRITORY:
Gmelina arborea Roxb.
Lantana Camara var. aculeata (L.) Moldenke
Lantana Mearnsii var. congolensis Moldenke
Lantana Mearnsii var. latibrecteolata Moldenke
Lantana scabrifolia Moldenke -- delete the "*"
Lippia Baumii Gurke

/

Lo Moldenke, Geographic Distribution 541

Lippia javanica (Burm. f.) Spreng.
Lippia scabra Hochst. -- to be deleted

Lippia Schliebeni Moldenke
Lippia strobiliformis Moldenke
Lippia Whytei Moldenke
ZANZIBAR PROTECTORATE:
Lippie javanica (Burm. f.) Spreng.
Lippia scabra Hochst. -- to be deleted
KENYA:
Lantana kisi A. Rich.
Lantana scabrifolia Moldenke
Lippia Baumii Lippia Baumii Gurke
Lippia javanica (Burm. f.) Spreng.
Lippia scabra Hochst. -- to be deleted
Lippia Schliebeni Moldenke
Lippia Whytei Moldenke
ANGOLA :
Lantana Mearnsii var. congolensis Moldenke (Loanda )
Lippie Lippia strobiliformis var. acuminata Moldenke
Vitex golungensis J. Ge Baker*
SOUTHERN RHODESIA:
Lantana Mearnsii var. congolensis Moldenke
Lantansa Mearnsii var. latibracteolata Moldenke
BRITISH NYASALAND PROTECTORATE:
Lippia Whytei Moldenke -- delete the "*"
Lippia Woodii Moldenke
PORTUGUES® EAST AFRICA: |
Holmskioldia tettensis (Klotzsch) Vatke (Mozambique)
Lantana salvifolia Jacq. (Mozambique )
Lippia javanica (Burm. f.) Spreng. (Mozambique)
Phyle nodiflora var. reptans (H.BeK.) Moldenke (Lourengo
Marques
SOUTHWEST AFRICA:
Lippia Baumii Gtirke
Phyla nodiflora (Le) Greene
UNION OF SOUTH AFRICA:
Lantana salvifolia Jacq. Biren
Lippia javanica (Burm. f.) Spreng. (Cape of Good Hope,
Natal, & RE ae
Lippia scabra Hochst. -- to be deleted
Lippia Woodii Moldenke (Transvaal )
Phyla nodiflora (L.) Greene (Transvaal )
Phyla nodiflora var. reptans (H.»B.K.) Moldenke (Natal)
Verbena bonarie: bonariensis L. (Cape of Good Hope)
Verbena officinalis L. (Transvaal)
Verbena rigida Spreng. (Basutoland)
SEYCHELLES ISLANDS:
x Stachytarpheta abortive Dans.
MASCARENE ISLANDS: .

342 PHYTOLOGIA Vol. 2, noe 8

Gmelina elliptica J. E. Sm. (Mauritius)
Holmskioldia sanguinea Retz. (Mauritius )
Phyla nodiflora var. canescens (HeBeKe) Moldenke (Maurit-
ius
ARABIA:
Lantana viburnoides Vahl (Yemen)
Phyle nodiflora (L.) Greene (Yemen)
Phyle nodiflora var. reptans (HeB.K.) Moldenke
REUNION?
Verbena bonariensis Le
UNION OF SOCIALIST SOVIET REFUBLICS:
Verbena officinalis L. (Adzharian)
Verbena supina L. (Stalingrad)
TURKEY: .
Fhyle nodiflora (L.) Greene
Verbena supina L.
Verbena supina f. erecta Moldenke
SYRIA: ‘
‘Phyle nodiflora (Le) Greene
Verbena officinalis L.

IRAN?

Fhyle nodiflora (L.) Greene
NEFAL :

Fhyla nodiflora (L.) Greene
INDIA:

Callicarpa cana L. -- to be deleted
Callicarpa candicans (Burm. f.) Hochre
Gmelina arborea var. canescens Haines (Bihar & Orissa)*
Gmelina arborea var. glaucescens C. B. Clarke (Bihar & Or-
issa )*

Gmelina elliptice J. B. Sm.
Holmskioldia sanguinea Retz. (Chamba )
Lantana Camara var. nivea (Vent.) L. H. Bailey (Madras)
Lantana dubia Wall. (Medras)
Lantana indica Roxb. (United Provinces)
lantena salvifolia Jacq. (Punjab & United Provinces)
Tectona grandis var. glabrifolia Moldenke (Madras)
Verbena officinalis L. (Chamba)

FRINCH INDIA:
Gmelina arborea Roxb.

CEYLON: ,
Fhyla nodiflora (L.) Greene
MANCHUKUO:
Vitex Negundo var. heterophylla (Franch.) Rehd.
Vitex Negundo var. incisa (Lame) Ce Be. Clarke -- to be de-
leted
CHINA :

Callicarpa Bodinieri Léeveillé (Sikang)
Callicarpa cana L. -- to be deleted

1947 Moldenke, Geographic Distribution 543

Callicarpa candicans (Burm. f.) Hochr. (Hupeh & Kwangtung )
Callicarpa Dielsii (Léveillé) P'ei -- to be deleted
Callicarpa integerrima ver. serruleta Li (Kwengtung)*
Callicarpa longifolia Lem. (Szechuan) .
Callicarpa rubella var. Dielsii (Leveillé) Li (Chekiang,
Kwangsi, Kwangtung, & Kweichow)*
Caryopteris Forrestii Diels is the correct form for this
name
Caryopteris incana (Thunb.) Miq. (Sikeang)
Caryopteris incana var. brachypoda (Hand.-Mazz.) Moldenke
-- to be deleted r
Caryopteris terniflora Maxim. (Sikang)
Caryopteris trichosphaera W. W. Sm. (Sikang)
Clerodendrum Bungei Steud. (Sikang)
Clerodendrum elachistanthum Merr. (Kwangei )
Clerodendrum kiangsiense Merr. (Chekieng & Kiangsi)*
Clerodendrum kwangtungense var. puberulum Li (Kwangtung )*
Clerodendrum trichotomum var. Fargesii (Dode) Rehd. (Ki-
angesu & Sikang
Congea chinensis Moldenke (Ytinnan)*
Congea tomentosa Roxb. -- to be deleted
Premna subcapitata Rehd. (Sikang)
Tsoongia axillariflora var. trifoliolata Li (Kwangtung)
Verbena officinalis L. (Sikeng
Vitex canescens Kurz (Szechuan)
Vitex Negundo L. (Sikang)
Vitex Negundo var. cannabifolia (Sieb. & Zucce) Hand.-
Mazz. (Sikeang)
Vitex Negundo var. heterophylla (Franch.) Rehd. (Honan, |
Hopeh, Shanei, Shantung, Shensi, Sikang, & Szechuan)
Vitex Negundo var. heterophylla f. multifida (Carr.) Rehd.
(Hopeh )*
Vitex Negundo var. incisa (Lam.) C. B. Clarke -- to be
. deleted
Vitex Negundo var. inciea f. multifida (Carr.) Rehd. --
to be deleted
Vitex yunnenensis W. W. Sm. (Sikang)
JAPAN:
Caryopteris divaricata (Sieb. & Zucc.) Maxim. (Hokkaido &
Pema yy.
Caryopteris incana (Thunb.) Miq. (Kiughiu)
Verbena officinalis L. (Kiushiu)
Mate,
Phyla nodiflora (L.) Greene
HAINAN ISLAND:
Callicarpaecana L. -- to be deleted
Callicarpa candicans (Burm. f.) Hochr.
FRENCH INDO-CHINA:
Gallicarpa cana L. -- to be deleted

544 PHYTOLOGIA Vol. 2, no. 8

Callicarpe cana var. Perryana Dop -- to be deleted
Gallicarpa candicans (Burm. f.) Hochr. (Annam, Cambodia, &

Cochin-china )
Callicarpa candicans var. Perryana (Dop) Moldenke (Cochin-
chins )*

Sphenodesme Jackiana (Wall.) Schau. (Cochin-china)
Tsoongia axilleriflora var. trifoliolata Li (Tonkin )
Vitex canescens Kurz (Cochin-chine
Vitex leptobotrys H. Hallier (Cambodie)
Vitex pinnata L. (Annem)
THAILAND :
Lantana Camara var. aculeata (L.) Moldenke
FEDERATED MALAY STATES:
Callicarpa cana L. -- to be deleted
Callicarpa candicane (Burm. f.) Hochr.
STRAITS SETTLEMENTS:
Callicarps cana L. -- to be deleted
soli tseres cendicens (Burm. f.) Hochr. (Malacca & Penang
Island
Gmelina asiatica L. (Fenang Island)
Gmelina elliptica J. E. Sm. (Malacca)
Lantana Camara var. aculeatea (L.) Moldenke (Singapore)
Sphenodesme barbata (fall.) Schau. (Malacca)
Sphenodesme Jackiana (Wall.) Schau. (Malacce )
Tectona grandis L. f. (Malacca)
Vitex trifolia var. simplicifolia Cham. (Singapore)
FHILIPPINE ISLANDS:
Callicarpe cana L. -- to be deleted
Callicarpa candicans (Burm. f.) Hochr. (Bohol, Cebu, Culi-
on, Luzon, Mindanao, Mindoro, Negros, Palawan, & Panay)
Lantana Camara var. aculeata (L.) Moldenke (Luzon)
Phyla nodiflora var. reptans (H.B.sK.) Moldenke
Vitex Negundo var. heterophylle (Franch.) Rehd. (Luzon)
Vitex Negundo var. incisa (Lam.) C. Be Clarke -- to be
deleted
MARIANNA ISLANDS:
Callicarpa cana L. -- to be deleted
Oallicarpa candicans (Burm. f.) Hochr. (Saipan & Tinian)
CAROLINE ISLANDS:
Callicarpa cana Ls. -- to be deleted
Callicarpa candicans (Burm. f.) Hochr. (Pelew Islands)
SUMATRA :
Callicarpa cana L. -- to be deleted
Callicarpa candicans (Burm. f.) Hochr.
JAVA?!
Callicarpa cana L. -- to be deleted °
Callicarps candicans (Burm. f.) Hochr.
Holmskioldia sanguinea Retz.
KAMBANGAN:

1947 Moldenke, Geographic Distribution 345

Callicarpa cana L. -- to be deleted
Callicarpa candicans (Burm. f.) Hochr.
BRITISH NORTH BORNEO:
Callicarpa cana L. -- to be deleted
Callicarpa candicans (Burm. f.) Hochr.
Lantana Camara var. aculeata (L.) Moldenke
CeLEBES :
Callicerpa cana L. -- to be deleted
Callicarpa candicans (Burm. f.) Hochr.
Duranta repens L.
LESSER SUNDA ISLANDS:
Callicarpa cana L. -- to be deleted
Callicarpa candicans (Burm. f.) Hochr. (Banka, Lombok,
Salajer Islands, Sumbawa, & Timor)
N2W GUINEA:
Callicarpa cana L. -- to be deleted
Callicarpa candicans (Burm. f.) Hochr. (Dutch New Guinea)
Faradaya chrysoc ada Ke. Schum. -- to be deleted
Feradaya splendida F. Muell. (Northeastern New Guinea)
Gmelina Ledermanni H. J. Lam (Northeastern New Guinea)
HAWAIIAN ISLANDS:
Duranta repens Le (Oshu)
Vitex trifolia L. (Oahu)
BISMARK ARCHIPZLAGO:
Callicarpa cana L. -- to be deleted
Callicarpa candicans (Burm. f.) Hochr. (Hermit Islands &
New Ireland |
SOLOMON ISLANDS:
Avicennia marine var. resinifera- (Forst.) Bakh. (Malaita)
NEW CALEDONIA:
Lantana tiliaefolia Cham.
Oxera Morierii Vieill. is the correct form for this name
Premna foetida Reinw.
Stachytarpheta australis Moldenke
TONGA ISLANDS:
Premna taitensis var. rimaterensis F. He Br. (Vavan Isl.)
FIJI ISLANDS:
Lantana Camara var. aculeata (L.) Moldenke (Viti Levu)
Tectona grandis var. “glabrifolia Moldenke (OQvalau)
AUSTRALIA:
Chloanthes grandiflora Moldenke (Western Australia )*
Clerodendrum costatum R. Br. (Queensland )*
Durenta repens L. (New South Wales)
Verbena officinalis L. (South Australia)
LIZARD ISLAND:
Callicarpa pedunculata R. Br.
NEW ZEALAND:
Avicennia marina var. resinifera (Forst.) Bakh. (Rangito-
to Island)

‘

(3546 PHYTOLOGIA Vole 2, noe 8

Teucridium parvifolium Hook. f. (North Island & South Is-
land

CULTIVATED :
Aloysia chamaedryfolia Cham. (France)
Aloysia polystach Griseb.) Moldenke (Argentina )
Aloysia triphylla Lier. ) Britton (Ecuador) - |
Bouchea prismatica (L.) Kuntze (Spain)
Callicarpa americana L. (Alabama & Spain)
Caryopteris incana (Thunb. ) Miqe (Belgium & Sweden)
Citharexylum montevidense (Spreng.) Moldenke (Uruguay)
Clerodendrum nutans Wall. (Florida)
Clerodendrum splendens Ge Don (Queensland )
Congea tomentosa Roxb. (Trinidad & Venezuela)
Congea velutina Wight (Belgian Congo & Java)
Congea villosa Wight (India & Straits Settlements)
Cornutia grandifolia (Schlecht. & Cham.) Schau. (Florida)
Durante repens L. (Egypt, Sicily, & Spain)
Duranta repens var. alba (Masters) Le He Bailey (Bermuda &

Mexico
Duranta repens var. microphylla (Willd.) Moldenke (Trini-
dad)

Gnelina elliptica J. E. Sm. (Belgium)
Holmskioldia sanguinea Retz. (England )
Holmskioldia speciosa Hutch. & Corbish. (Hawaiian Islands)

es ee a ee

Lantana achyranthifolia Desf. (Germany, New York, & Spain)
| Lantana annua L. (Spain)

lantana antidotalis Schum. & Thonn. (Belgian Congo)

lentana arida Britton (Belgium & Spain)

Lantana boyacana Moldenke (plein

Lantana Camara Le (Belgium, France, Germany, Spain, & Tex-
as

Lantana Camara var. aculeata (L.) Moldenke (Belgium, Ger-
many, & Spain)

Lantana Camara ver. mista (L.) Le He Bailey (Spain)

Lantana Camara var. mutabilis (Hook.) Le He Bailey (New
York)

Lantane Camara var. nivea (Vent.) L. He Bailey (Belgium &
Scotland)

Lantana Chamissonis (D. Dietr.) Benth. (France)

Lantana dubia Wall. (Germany & Spain)

Lantana foetida Rusby -- to be deleted

Lantana fucata Lindl. (France)

Lantana glandulosissima Hayek (Scotland )

Lantana glutinosa Poepp. (France & Uruguay)

Lantane Heughtii Moldenke (Spain)

. Lantana hirte Grah. (Belgium)

Lantana involucrata L. (France)

Lantana involucrata var. odorata (L.) Moldenke (Belgium)

Lantana montevidensis (Spreng.) Briq. (Belgium, France,

1947 Moldenke, Geographic Distribution 347

Mexico, & Texas)

Lantane reticulata Pers (France)

Lantana salvifolia Jacq. (Belgium & Germeny)

Lantana tiliaefolia Cham. (Germany )

Lantana trifolia L. (Belgium, France, Germany, New York,
Scotland, Spain, & Trinidad )

Lantana undulata Schrank (Germany )

‘Lantana urticaefolie Mill. (France)

Lantana velutina Mart. & Gel. (Belgium, France, & Scot-
. land

Lippie alba (Mill.) N. EB. Br. (Belgium, Costa Rica, Germ-
any, & Maryland )

Lippia callicarpaefolia H.B.K. (France)
Lippia graveolens HeBeK. (Costa Rica)
Lippia javanica (Burm. f.) Spreng. (France)
Lippia micromera Schau. (Hawaiian Islands)
Lippia scabra Hochst. -- to be deleted
Petraeovitex multiflora (J. &. Sm.) Merr. (Java)
Phyla lanceolata (Michx.) Greene (California)
Phyla nodiflora (L.) Greene (Italy)
Phyle nodiflora var. reptans (H»BeK.) Moldenke (Germany )
Phyla nodiflora var. rosea (De. Don) Moldenke (Belgium)
Phyla nodiflora var. suborbicularis (L. Chev.) Moldenke
France
Phyle scaberrima (A. Le Juss.) Moldenke (France & Germmy)
Fremna corymbosa var. obtusifolia (R. Br.) Fletcher
(abe)
Premna corymbosa var. sambucina (Wall.) Moldenke (Belgium)
x Stachytarpheta adulterina Urb. & Ekm. (Java)
Stachytarpheta angustifolia (Mill.) Vahl (Germany & Spain)
Stachytarpheta cayennensis (Ls C. Rich.) Vehl (Belgium &
Germany )

Stachytarpheta elatior Schrad. (Belgium)

eee

Stachytarpheta jamaicensis (L.) Vahl (Belgium, Java, &
Ey

Stachytarpheta mutabilis (Jacq.) Vahl (Belgium & France)

Stachytarpheta urticaefolia (Salisb.) Sims (Belgium &
ery
Tectona grandis L. f. (Belgtan Congo)

Tectona grandis var. glabrifolia Moldenke (Ceylon)

Verbena bipinnatifida Nutt. (Belgium)

Verbena bipinnatifida var. latilobata Perry (Mexico)

Verbena bonariensis L. (Belgium & Peru)

Verbena bracteata Lag. & Rodr. (Belgium & France)

Verbena canadensis (L.) Britton (Belgium, Germany, Italy, &
Spain ’

Verbena carolina L. (Spain)

Verbena elegans H.BeK. (New York)

x Verbena Engelmannii Moldenke (Spain)

348 PHYTOLOGIA . Vol. 2, now &

Verbena hastata L. (Belgium, England, France, Germany, &
Spain

Verbena hispida Rufz & Pav. (France & Germany)

x Verbena hybride Voss (Brazil )

Verbena incisa Hook. (Germany & Spein)

Verbena lasiostachys Link (France, Germany, & Spain)

Verbena litorelis H.eB.eK. (France & Spain)

x Verbena matritensis Moldenke (Spain)*

Verbena mendocina R. A. Phil. (New York)

Verbena menthsefolia Benth. (Spain)

Verbena officinalis L. (Belgium, Quebec, & Spain)

Verbena peruviana (L.) Britton (Belgium)

Verbena phlogiflora Cham. (Belgium & Germany)

Verbena rigida Spreng. (Belgium, Peru, & Spain)

Verbena santiaguensis (Covas & Schnack) Moldenke (New York)

Verbena simplex Lehm. (Belgium, Czechoslovakia, France, &

Spain)

Verbena stricta Vent. (France)

Verbena supina L. (Belgium & Frence)

Verbena tenera Spreng. (Belgium & Texas)

Verbena tenuisecta Briq. (Kenya)

Verbena urticifolia L. (Germany, Pennsylvania, & Spain)

Verbena xutha Lehm. (Belgium)

Vitex Agnus-castus f. roses Rehd. (Texas)

Vitex altissima var. alata (Willd.) Moldenke (Florida)

Vitex Negundo var. heterophylla (Franch.) Rehd. (Arizona,

Austria, Barbados, Belgium, Brazil, British Guiana,
Californig, Denmark, District of Columbia, Englend,
France, Germany, Illinois, Italy, Kansas, Martinique,
Massachusetts, Missouri; Mongolia, New Jersey, New
York, Oklahoma, Pennsylvania, Russia, Spain, Sweden,
Switzerland, Trinidad, & Virginia)

Vitex Negundo ver. heterophylla f. miltifida (Carr.) Rehd.

Vitex Negundo var. incisa (Lam.) C. B. Clarke -- to be

deleted
Vitex parviflora A. L. Juss. (Florida)

FOSSILIZED:
Clerodendrum robustum Klotzsch (Pleistocene of Cameroons *
Clerodendrum Thomasii Moldenke (Pleistocene of Cameroons)
Vitex Doniana Sweet (Pleistocene of Cameroons)

a

THE KNOWN GEOGRAPHIC DISTRIBUTION OF THE MEMBERS OF THE

ERIOCAULACEAE. SUPPLEMENT 1
Harold N. Moldenke

Several thousand additional specimens of Eriocaulaceae
have been examined by the writer since the publication in
1946 of his "The known geographic distribution of. the menm-
bers of the Eriocaulaceae". These specimens have brought to
light 92 new country or island records, 139 new state, prov-
ince, or department records, and 61 new county or parish re-
cords not previously recorded. Also 121 additional scientif-
ic names must be added to the alphabetic list of scientific
names proposed in this group, including mis-spellings and
mis-accreditions. The addition of these names brings the to-
tal of scientific names now accounted for to 2181; the total
accepted genera to 11 and species and varieties 1202. The
actual specimens on which these records are based will even-
tually be published by the writer in others of his series of
papers on this family. They are deposited in the herbaria of
the New York Botanical Garden, New York State Museum, Univ-
ersity of Texas, L. H. Bailey Hortorium, Barnard College,
Jardin Botanique de 1'Etat at Brussels, Butler University,
University of Miami, Columbia University, University of ~
California at Berkeley, Carnegie Museum, University of Cin-
cinnati, DeFauw University, Leland Stanford University,
Earlham College, Botaniska Trédgard at Gdteborg, Duke Uni-
versity, Cornell University, University of California at Los
Angeles, Universidad Nacional de Mexico at Mexico City, In-
stituto Miguel Lillo, Frinceton University, Jardin Botanico
at Madrid, Museo Nacional de Historia Natural at Buenos
Aires, Naturhistoriska Riksmuseum at Stockholm, Instituto
Darwinion at San Isidro, Vanderbilt University, and the Uni-
versity of Vermont.

CANADA :
Quebsc:
Eriocaulon septangulare With. (Fontiac County & Mistass-
ini Region)
UNITED STATES OF AMERICA:
Maine: '
Eriocaulon septangulare With. (Androscoggin County)
Vermont :
Eriocaulon septangulare With. (Grand Isle, Rutland,
Windham, & Windsor Counties)

349

5350 PHYTOLOGIA Vol. 2, no. &

Massachusetts:
Eriocaulon septangulere With. (Bristol County)
Rhode Island: :
Eriocaulon septengulare With. (Washington County)
New York:
Eriocaulon Farkeri Be. Le Robinson (Albany, Orange, & Ul-
ster Counties
Eriocaulon septangulare With. (Albany, Columbia, Oneida,
Onondaga, Rockland, & Schenectady Counties’)
New Jersey:
Eriocaulon decangulare L. (Hunterdon County)
Pennsylvania :
Eriocaulon decangulere Le
Virginia:
Eriocaulon decangulare Le
Eriocaulon septangulare With.
Lachnoceulon anceps (Walt.) Morong (Sussex County)
North Carolina:
Eriocaulon compressum Lam. (Buncombe & Rowan Counties)
Eriocaulon decangulare L. (Jackson County)
Lachnocaulon anceps (Walt.) Morong (Martin County )
South Carolina: ;
Eriocaulon compressum Lam. (Darlington County )
Eriocaulon decangulare L. (Darlington & Greenville
Counties)
Lachnocaulon anceps (Walt.) Morong (Darlington County )

Syngonanthus flavidulus (Michx.) Ruhl. (Lexington
County )
Georgia:

Syngonanthus flavidulus (Michx.) Ruhl. (Lowndes County)
Florida:
Eriocaulon compressum Lam. (Baker County )
Eriocaulon decangulare L. (Escambia County )
Eriocaulon lineare Small (Volusia County )
Briocaulon Ravenelii Chapm. (Hillsborough & Lee Count-
ies
‘Lachriocaulon anceps (Walt.) Morong (Escambia & Saint
Johns Counties
Syngonenthus flavidulus (Michx.) Ruhl. (Marion & Semin-
ole Counties
Alabama :
Syngonanthus flavidulus (Michx.) Ruhl. (Baldwin County )
Mississippi:
Briocaulon compressum Lam. (Hancock County )
Louisiana:
Eriocaulon compressum Lam. (Orleans Parish)
Texas:
Eriocaulon compressum Lam. (Hardin & Jefferson Counties)
Eriocaulon decangulare L. (Anderson, Austin, Freestone,

1947 Moldenke, Briocaulacese 351

Jasper, Jefferson, Newton, Robertson, Rusk, & Tyler
Counties )
Eriocaulon Kérnickianum Van Heurck & Muell.-Arg. (Polk
County )
Eriocaulon texense Korn. (Austin, Leon, Milam, & Robertson
Counties
Lachnocaulon anceps (Walt.) Morong (Jefferson, Newton, &
Tyler Counties)
MEXICO:
Eriocaulon Benthami Kunth (Veracruz)
Eriocaulon capitutatum Moldenke*
Eriocaulon Ehrenbergianum Klotzsch (Federal District)
Eriocaulon microcephalum H.BeK. (Federal District)
Eriocaulon paradoxum Moldenke*
Eriocaulon Pringlei S. Wats. (Mexico)
CUBA :
The records given for "Santa Clara" should be changed to
read "Las Villas" /
COLOMBIA:
Paepalanthus columbiensis Ruhl. (£1 Valle)
Paepalanthus crassicaulis Korn. (Santander Norte)
Paepalanthus ensifolius (H.B.K.) Kunth (Magdalene)
Paepalanthus Lindenii Ruhl. (Santander Norte)
Faepalanthus muscosus Korn. (El Cauca)
Paepalanthus pilosus (H.B.K.) Kunth (Santander Norte )
Syngonanthus ceulescens (Poir.) Ruhl. (Santander Norte)
VENEZUELA:
Carptotepala insolita Moldenke (Bolfvar)*
Eriocaulon guianense Kérn. (Amazonas)
Eriocaulon dimorphopetalum Moldenke (Bol{fvar)*
Eriocaulon melanocephalum Kunth (Bolfvar)
Eriocaulon Steyermarkii Moldenke (Bol{fvar)*
Eriocaulon tenuifolium Klotzsch (Bolfvar)
Leiothrix Steyermarkii Moldenke (Bolfvar)*
Leiothrix umbratilis Moldenke (Bolfvar)*
Paepalanthus capilleceus Klotzsch (Bolfvar)
Paepalanthus capilleceus var. proliferus Gleason (Ama-
zona s
Paepalanthus columbiensis Ruhl. (Mérida)
Paepalanthus convexus Gleason (Bolfvar)
Paepalanthus crassicaulis Kérn. (Merida)
Paepelanthus dendroides (H»BeK.) Kunth (Meride )
Paepalanthus dichotomus Klotzsch (Bol{var)
Paepalanthus fraternus N. &. Br. (Bolfvar)
Paepalanthus Gleasonii Moldenke (Amazonas & Bolfvar)
Paepalanthus Karstenii Ruhl. (Mérida & Téohira)
Paepalanthus Maguirei Moldenke (Amazonas)
Paepalanthus meridensis Klotzsch (Tachira)
Paepalanthus miscosus Kérn. (Merida)

352 PHYTOLOGIA Vol. 2, no. 8

Paepalanthus pauper Moldenke (Amazonas ) ©
Paepalanthus perplexans Moldenke (Bolfvar )*
Paepalanthus pilosus (H-B.K.) Kunth (Mérida)
Faepalanthus roraimensis Moldenke (Bolfvar )*
Paepalanthus Schomburgkii Klotzsch (Bolivar)
Paepalanthus squamuliferus Moldenke (Bolf{var)*
Paepalanthus Steyermarkii Moldenke (Bolfvar )*
Paepalanthus subacaulescens N. &. Br. (Bol{var)
Paepalanthus subsessilis Moldenke (Lara )*
Paepalenthus subtilis Miq. (Amazonas & Bol{var)
Faepalanthus truxillensis Korn. (Tachira)
Philodice Hoffmannseggii Mart. (Amazonas)
Rondonanthus micropetalus Moldenke (Bol{fvar)*#*
Rondonantnus roraimae (Oliv.) Herzog (Bolfvar)
Syngonanthus biformis (N. E. Br.) Gleason (Amazonas & Bol-
Tvar)
Syngonanthus caulescens (Poir.) Ruhl. (Amazonas)
Syngonanthus duidae Moldenke (Amazonas )*

Syngonanthus eriophyllus var. glanduliferus Ruhl. (Bol-
{var)

Syngonanthus fertilis (Kérn.) Ruhl. (Bol{fvar)
Syngonanthus glandulosus Gleason (Bolfvar)
racilis (

Syngonanthus Korn.) Ruhl. (Amazonas, Bolfvar, &
Sucre )

Syngonanthus gracilis var. glabriusculus Ruhl. (Amazonas)
Syngonanthus gracilis var. hirtellus (Steud.) Ruhl. (Ama-
zonas )

Syngonanthus heteropeploides Berens (Ama zonas )

Syngonanthus longipes Gleason (Bolivar)
Syngonanthus simplex (Miqe) Ruhl. (Amazonas)

Syngonanthus tricostatus Gleason (Bol{var)

Syngonanthus umbellatus (Lam.) Ruhl. (Amazones)

Syngonanthus venezuelensis Moldenke (Bol{ver)*

Syngonanthus verticillatus (Bong.) Ruhl. (Bolfvar)
BRITISH GUIANA:

Eriocaulon guianense Korn.

Eriocaulon heterodoxum Moldenke*

Paepalanthus brunneus Moldenke*

Paepalanthus capillaceus var. proliferus Gleason -- delete
the **

Paepalanthus filipes Moldenke*
Paepalanthus Gleasonii Moldenke -- delete the "*"
Paepalanthus griseus Moldenke*

Paepa lanthus pauper Moldenke
Philodice Hoffmannseggii Mart.

Syngonanthus glandulosus Gleason -- delete the Wat

Syngonanthus guianensis Moldenke*

Syngonanthus longipes Gleason -- delete the ***
Syngonanthus savannarum Moldenke *

i
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4 - ys -— :
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PHYTOLOGIA

‘Designed to expedite botanical RECON C: ae
November, 1947 No. 9
CONTENTS

7AN MELLE, P. J., From texts to plants—or from
| . MATES RO) ORES Eco. vcrbaceescnsrnah’ sind spudty Mil stds ge Sasa 353

bee

3 | Mote H.N., The known geographic distribution of the
Biot. Verbenaceae, Avicenniaceae, Stilbaceae, and

Sym phoremaceae. Supplement 7 ............+4. -acnaapeega an

yy,

Published by H. A. Gleason and Harold N. Moldenke —
The New York Botanical Garden,
Bronx Park, New York 58, N. Y.

2

Price of this number, 50 cents; per volume, $5.00 in advance

Vol. 2, No. 8 was issued June 30, 1947

LIBRARY
NEW YORK
BOTANICAL

GARDEN

/

FROM TEXTS TO PLANTS - OR FROM PLANTS TO TEXTS?

P.J.- van Melle

A Difference of Approach, [llustrated by Juniperus chin-
ensis var. globosa Hornibrook, and var. plumosa Hornibr.,
f. & Slow-growing Conif., 62 and 66 (1923) sensu Cornman
and
x J. media var. globosa (Hornibr.) van Melle, and var. plum-
osa (Hornibr.) van Melle in Phytologia 2 : 191 (1946).

z

In a thesis: "Studies in the Genus Juniperus" (Cornell
University, March 1947) on the taxonomy of junipers cultiva-
ted in the United States, John F. Cornman excoriates a paper
published by me in Phytologia (as cited), "The Junipers Com-
monly Included in Juniperus chinensis". His strictures seem
excessively voluminous and immoderate, almost obsessive, in-
appropriate to the usually objective nature of a thesis, and
poorly substantiated. He washes my face for me, quite behind
the ears, for instance, for my treatment of the above-
mentioned Hornibrookian epithets, which, says he, I inter-
changed, in misunderstanding of taxonomic principle and pro-=
cedure.

Actually I did no such thing. On the basis of evidence
reviewed below I interpret Hornibrook's var. globosa as pis=
tillate. This conclusion is employed by me as a fixed point
for the treatment of the wobbly globosa-plumosa complex of
Hornibrook, whose 1923 texts appear to me inapt, inconclus-
ive and somewhat confused, while his 1938 texts are more
definitely erroneous in important parts. Cornman describes
the var. globosa Hornibr. as staminate, and as equalling my
var. plumosa (Hornibr.). We cannot both be right.

Of my designation of a Ching No. 53 sheet as the type of
my var. plumosa he says that any novice would at once note
the discrepancy. I trust that, if he would compare with my
designated type sheet (U. S. Nat'l Herb. No. 1245122) unmis-
takable material of my var. plumosa, the result would fall
out more favorable to me. There is no more unmistakable in-
dication of what my var. plumosa is than to describe it as
equal (including leaf glands) to Cornman's var. aureo-
variegata or my f. albo-variegata - minus the variegations.
It equals reverted, green parts of these variants. I reject
as inconstant and unreliable the gland character noted by
Cornman in his var. aureo-variegata.

If from his point of view I seem to have mistakenly iden-
tified my plumosa material with the Ching sheet cited by him
he is, of course, free, and in his light bound, to disagree

453

554 Pel tT Ob Oe 2 2 Vol. 2, no. 9

with me. Such disagreements are no rare occurrence between
workers. But it is hardly customary for one to call another
practically a nitwit over them or to allege "fundamental er-—
ror", "misunderstanding of the type method as employed in
orthodox taxonomic procedure", etc. It may well be that
Cornman, by failing to review my designated type sheet, by
reviewing, instead, a sheet not cited in my 1946 paper, and
by attaching Van Melle type labels to sheets not so designa-
ted by me (pp. 280, 298) permitted himself a taxonomic pec=
cadillo or two.

On pe. 291 he says: "Upon inquiry as to why he apparently
interchanged Hornibrook's names and descriptions, van Melle
writes: 'My treatment of the two varieties represents, in
the main, the traditional horticultural grouping. While I
base my varietal names on Hornibrook, I did so without any
particular deference to Hornibrook's texts'. This violates
the basic premise of taxonomy and plant nomenclature, and
the conclusions cannot be considered seriously."

I am duly mortified at this public quotation from my cor-
respondence, ungroomed as it was for publication, and look-
ing all the more vulnerable away from its context. My inten-
tion was, of course, "the letter of Hornibrook's texts".

From the literature of cultivated plants I gather an im-

pression that a worker's veneration of the letter of texts
' is proportioned approximately to his own knowledge of his
materials. In many cases only that knowledge can provide a
proper basis for the evaluation of texts. Without it one be-
comes easily a slave to the letter of texts = a sort of tax-
onomic automaton; and there is constant danger of arriving
at concepts which, however orthodoxly derived from texts,
may bear only a sketchy likeness to existing kinds of plants.

If then, in Cornman's eyes, I seem a little disrespectful
of Hornibrook's letter (but does not he, himself, play rath-
er freely with that author's foliage descriptions?), my only
defense is that I have had these junipers under observation
for over 55 years. My experience with them dates back to
1911, when, at Bobbink & Atkins, Rutherford, N. J. = then
large importers - it was my job to check against invoices
incoming shipments from Europe and Japan. The junipers under
discussion here then constituted a considerable item. The
var. plumosa, sensu van Melle, was then coming in from Bos-
koop mainly as J. japonica, to a lesser extent also as J.
chinensis procumbens; the var. globosa, sensu van Melle, as
J. japonica globosa, rather rarely as J. japonica nana, and
more rarely as J. chinensis procumbens globosa. The name
nena was then not in use in Boskoop to an appreciable extent.
It was more particularly an English term for the var. globo-
sa, sensu Van Melle.

IT am well aware, then, that Hornibrook, in 1923, under

-

1947 van Melle, From Texts to Plants 355

his vars. globosa and plumosa, was not describing thereto-
fore unknowm junipers, but was only endeavoring, more or
less successfully, to describe kinds that had been long and
widely known. Thus, in working over his texts, my principal
concern was not: "How can I best preserve the letter of his
texts?", but rather: "How can I best manage to present the
several elements in this complex correctly and in an orderly
grouping?"

Generally, in studies of this sort, my procedure is ap-
proximatelythis: Make an "index plantarum" - an inventory of
the elements which require to be accounted for. Enumerate
them, even if, at first, only by numbers or letters. Study
them; group them as you come to think they should be. Do
your own, initial job of taxonomy on them, and give it time
to age. Keep correcting, revising, refining. Then compare
your work with past treatments. In turn, study this litera-
ture; accept from it what revisions or corrections seem in
order. But insofar as your own insights continue to appear
preferable to you, hold on to them, and do not hesitate to
re-interpret, emend or reject past treatments. Trust your
own eyes. Finally, express your findings in terms of the a-
vailable nomenclature. And all this within a not too strict
interpretation of the laws.

This sort of thing, says Cornman (pp. 242, 243), is not
taxonomy. Unanointed logic! Blasphemy! One can have no truck
with it.

Indeed, it is not the way of much of what passes among us
for the taxonomy of cultivated conifers. Not all of it, nor
even, one hopes, the greater part, but much of it, consists
of wholly or largely literary, unrealistic concepts of kinds,
derived perhaps through the most meticulous taxonomic proce-
dures from texts and specimens which cannot be identified
with known kinds of plants without the aid of extraneous,
circumstantial evidence of a chronological or other purely
logical nature. It consists on the one hand of ignorance of
the living plants involved and on the other hand of a morbid
veneration of decrepit texts and specimens.

Some of Cornman's work appears to me to be of this kind.
For instance, his J. chinensis, which may be said to occupy
a pivotal place in his Thesis, and which ought to represent
a fairly concise concept, does not, in my opinion, represent
any one definable kind, discerned and then named. It repre-
sents, rather, an effort to account for. a J. chinensis in
terms of the literature of that species - than which there
is not a more befuddled chapter in all the taxonomy of juni-
pers. Cornman's var. typica comprises a galaxy of junipers,
and a number of obscure literary entities. For instance, of
the J. virginica of Thunberg he says: "There is no reason to
suppose that he had anything but J. chinensis." One asks:

556 PHYTOLOGIA Vol. 2, noe 9

what grounds are there to suppose that he did have it? And
that the J. japonica cernua and dimorpha of Roxburgh repre-
sent it?

On pp. 242 and 243 Cornman says of my inclusion in J.
chinensis L. of the "variety" oblonga: "In general, van
Melle's papers now published must be rejected because of
their fundamental errors. He deprecates herbarium specimens
as inadequate, as indeed they usually are, but from only a
poor photograph of the type of Linnaeus’ J. chinensis he at-
taches Linnaeus! name to what we know as var. oblonga. Thus
he rejects customary logic without a trial and depends sole-
ly upon circumstantial evidence and 'the resources of dis-
ciplined intuition'."

That is a mouthful. In my "Review of J. chinensis et al."
(New York’ Botanical Garden, April, 1947) I admit my inabili-
ty to identify satisfactorily with the Linnaean type and
texts, on a basis of their intrinsic value only, any one
known kind of juniper. I admit frankly that my interpreta-
tion of the Linnaean record is based for a large part upon
the best of chronological and circumstantial evidence at my
command. I doubt that Cornman is in a position to do better
with it. I regard as the most telling aspect of the specimen
the denseness of its (exclusively juvenile) foliage; end of
the texts: "Folia.....magis quam in reliquis conferta" and
"distinctissima densitate foliorum." All this checks very
well with the one, distinct, juniper associated by me with
J. chinensis: the almost exclusively pistillate J. chinensis
foemina of nurseries, originally, and still widely, known as
J. chinensis. To its name was added, in the 1850's, the epi-
thet "foemina" to distinguish it from seedlings then being
distributed in England by Standish & Noble of the monoecious
J. sphaerica Lindl. -- which then, as they do today, bore
only or almost exclusively staminate flowers in their youth.
These seedlings were at the time erroneously believed to
represent the staminate counterpart of the earlier-cultivat-
ed, pistillate J. chinensis; and they were called J. chinen-
sis mas or mascula.

Cornman's J. chinensis L. agrees with mine to the extent
that he includes the "foemina" material in his var. typica.
Only, while I limit my citations to that entity plus its ob-
vious synonyme (including the oblonga name), he presente the
var. oblonga as highly distinct. In addition, he cites under
his var. typica all manner of other junipers, such as my J.
sphaerica vars. dioica and neaboriensis, which are conspicu-
ously different things from the "foemina" material.

This is my view of the oblonga name: It was given in 1914
by Bobbink & Atkins to an importation from Europe, while I
was in their employ. It was my judgement then, and has been
ever since, that the material so named differed in no manner

1947 van Melle, From Texts to Plants 357

worth recognition from that which had been theretofore known
at Bobbink & Atkins and elsewhere as J. chinensis and J.
chinensis foemina. It is very well possible that, at the
time, it appeared to somebody as a little different in gen-
eral aspect - in the way that many a batch of the "foemina"
material, perhaps pruned a little differently, or grow on a
different soil or under different conditions, looks a little
different from another. Comparisons at the time, and since -
many of them based upon large plants distributed under the
two names - convince me that the oblonga listing represented
nothing but a new name for a very old thing. Probably no

' juniper has been sent into the world under more different
names than the old "foemina" material.

Is there anything about my disposition of this name to
warrant Cornman's allegation of "fundamental error" or "re-
jection of customary logic without trial"?

But, to return to the vars. globoss and plumosa Hornibr.
- my first move toward a treatment of this complex was the
making of an inventory of the elements in it, known to be in
cultivation. This inventory revealed the existence of the
following two groups, a and b: ’

Group a, consisting of Group b, consisting of
2 elements, 4 elements,
both pistillate: all staminate:

a, the typical green form
al, a yellow-suffused form

» the typical green form

1, a yellow-suffused form
2, a yellow-variegated form
3, a white-variegated form

My index did not include the globosa cineria element des-
cribed by Hornibrook in 1923 from a small plant, now ex-
tinct. A corresponding element was known in England about
1910 as nana glauca. It does not appear to have become wide-
ly grown in Europe. Again, in 1940, Grootendorst described a
J. chinensis Blaauw's Variety (possibly the same element) as
a novelty from Japan, about to be introduced in Europe. The
illustration of this plant suggests the habit of my stamin-
ate group b.

The idea of these two, apparently closely related groups,
one pistillate, the other staminate, may look artificial.
Indeed, the fact that no staminate seedling of my pistillate
group appears to have been know is noteworthy. It is match-
ed in other kinds of juniper. None is known of my x J. media
var. arbuscula, of J. procumbens Sieb. & Zucc., or of Je
squamata var. Meyeri Rehd. The last recorded, cultivated
material of the J. chinensis foemina of nurseries (J. chin-
ensis L. sensu van Melle) is represented by the f. aurea
(Young), which originated in England before 1872. Yet pis-
tillate plants of these several kinds fruit freely in nure-
eries.

Iv jo lo jo"

358 PHYTOLOGIA . Vol. 2, no. 9

I do not propose to explain the phenomenon. I merely. re-
port it. It seems probable that apomixy is involved. The
matter merits investigation. At any rate, I need not hesita-
te to present my groups a and b, respectively, as pistillate
and staminate. In each of them the constituent elements are
mutually identical except as to color, and for a not consis-
tent tendency in a 1 and in b 1 and 2 toward protracted juv-
enility. Neither do I hesitate to reject Cornman's presenta-
tion of his var. aureo-variegata as intermediate between my
two groups.

My pistillate group a equale the materials shipped into
the United States from Boskoop in large quantities as Je
japonica globosa and globosa aurea; my staminate group b,
those shipped in from Boskoop mainly as J. japonica, japoni-
ca aurea, aureo-variegata and albo-variegata, and to a less-~
er extent by corresponding chinensis procumbens listings.

It seems that Hornibrook's globosa group (including the
misplaced f. cineria) was matched in England by a group of
three nana listings. An R. H. S. Award of Merit was given on
December 10, 1908 "To J, chinensis nana aurea from Messrs.
Waterer, Bagshot. Three new dwarf forms of the Caan juni-
per were shown - nana, nana glauca and nana aurea."

It seems certain that both the globosa al and plu plumosa groups
were included in Beissner's J. chinensis procumbens of 1891,
and in the notoriously inclusive japonica listings as far
back as the early 1870's - at which time, also, the Pfitzer
Juniper made its debut, in France, as J. japonica pendula.

The first clear separation of the two groups discernible
to me is that of the Boskoop listings in the early 1900's of
J. japonica, with 4 variants, and J. japonica globosa, with
1 variant. However, it is on the more or less formal litera-
ture of their names that one must base one's choice of
names. Therefore, Hornibrook's texts merit consideration.
Yet, if it were not that his names have become established
in horticulture I might well have rejected them as nomina
dubia. For I see in them very little literal, descriptive,
distinguishing value. I preferred, however, to explore what
sense might be discovered below their textual surface, and
from the related nana listings of 1923. Thus I discovered
implications more compelling than the letter of the texts.

Except for the aureo-variegata and albo-variegata elem
ents, my treatment of the complex is based, nomenclaturally,
upon Hornibrook's 1923 edition. I need not accept the 1938
treatment, even though my grouping of the variants and my
reduction to synonymy of the var. decumbens with var. plu-
mosa, and my citation, under the latter, of the J. japonica
and . J. chinensis procumbens of Boskoop concur with the 1938
edition.

While Hornibrook nowhere states outright the sex of the

1947 van Melle, From Texts to Plants 359

materials except that of his globosa cineria, he says in
1925 that his var. globosa and f. aurea are of the same sex;
that they have not yet borne fruit, while "form 3 (cineria)
is a male plant." Again, in 1938: "Var. globosa cineria is a
different form; it is a male plant." Does not this imply
that his var. globosa and its aurea form were pistillate?

In 1923 he lists a var. nana (Hochst.) as follows: "A
form received from the Arnold Arboretum without description.
Extremely slow-growing.....it would seem to be the erect
form of the var. globosa, its sprays being similar to that
variety in size and general appearance." Living plants at
the Arnold Arboretum harking back to material received there
as var. nana, from Waterer in 1909, and from Farquhar in
1917, are all pistillate, identical with mya element - with
my var. globosa. They do not differ from it by any* supposed
erect character, by which, only, Hornibrook distinguished
this material from his var. globosa. The type specimen of my
var. globosa was taken from the Farquhar material, even
though this is labelled var. plumosa in the Arboretum.

I think that I am on safe enough ground in identifying
Hornibrook's globosa group, consisting (exclusive of the
misplaced cineria form) of 2 elements, with my pistillate
group a, which consists of 2 elements. Having done this, -
then, if Hornibrook's plumosa group (consisting of 4 ele-
ments) is to be interpreted at all in terms of the known
junipers within the complex, I am bound to identify it with
my staminate group b which consists of 4 elements. In the.
case of two of Hornibrook's four plumosa elements - his
aureo-varierata and albo-variegata - anyone may ascertain
for himself that these (the only known variegated elements
in the complex) are both staminate. Since nothing in the
1923 edition militates otherwise, I assume, as a matter of
simple logic, that Hornibrook's plumosa aureo-varisgata and
albo-variegata, minus their variegations, equal his var.
plumosa. Such is the composition of my plumosa group; b 2
minus 2 equals b; b’4 minus 5 equals b.

This is no over-simplification. Thus, my treatment of the
globosa-plumosa complex is reduced to the grouping of its
several elements under a pistillate and a staminate type, as
shown in my a and b schedule. Each group has its consistent,
distinct characters. All that is needed to place any of the
elements under its proper heading is to know either its sex
or its habit. My Latin diagnoses attribute to the pistillate
and to the staminate groups the characters that go with
them. This involves no undue violence to Hornibrook's 1923
texts. But it is at variance with the habit ascribed by him
in the 1938 edition to the J. japonica and J. chinensis pro-
cumbens of Boskoop — names which he associates with his var.
plumosa umosa. I quote: "As grown in large quantities in Holland,

360 PATE Ob 0 Gus Vol. 2, noe 9

this form (plumosa) makes eventually a somewhat loose shrub
about twice as broad as high.....-Hitherto this form has
been sent out by well-known Dutch nurseries under the names
J. japonica and J. chinensis procumbens." This is not the
habit of these two Boskoop listings, but that of the J. jap-
onica globosa of Boskoop - my var. globosa (Plate X in my
WReview" ). A true, youthful habit phase of the J. japonica
of Boskoop is shown by Hornibrook under his var. globosa, in
the Report, Conif. Conference (1932) and in his 1938 edi-
tion. This is the habit of the staminate material - not that
of the var. nana, not of his var. globosa of 1923.

I conclude that Hornibrook at no time appeared to know
clearly, one from the other, the green types of his vars.
globosa and plumosa. Had he understood them well, he would
not have been as uncertain as he appears to be in 1938 about
where to place his variegated forms. I quote: "On the whole
they approach nearest in habit and foliage to Je Ce Vare
plumosa Hornibr.......and possibly the best way out of the
existing confusion ie to classify them as forms of var. plu-
mosa; the suggestion to do so originates with Mr. Herman J.
Grootendorst of Boskoop, whose firm has grown and observed
these forms for a great many years, and on consideration I
agree with him." Actually, these staminate forms which he
places with his var. plumosa are variants of the material
depicted by him under his var. globosa. No careful worker,
aware of these defects in Hornibrook's post-1923 treatment
would wish to perpetuate them; and it is because of them
that I exercise my right to reject all or part of the post~-
1925 treatments.

Cornman follows me in associating the habit “ultimately
twice asyhichfas \wide" with the pistillate sex; but, unaware
of Hornibrook's mis-description of the J. japonica of Bos=
koop and of the implied pistillate nature of the 19235 var.
globosa, he reverses my names. To his variegated material he
ascribes, erroneously, the habit of the pistillate and the
sex of the staminate group, as well as a) gland character
different from that which he notes under his vars. globosa
and plumosa.

In terms of the known elements in the complex his var.
aureo-variegats is neither flesh nor fish. There are no such
intermediate elements; and the glands in the variegated mat-
erial are as variable as they are throughout the complex.
Sornman sidesteps the impasse of his aureo-variegata mater-
ial by saying: "There is no compulsion to classify it nomen-
claturally as a variant of either" variety.

I believe that in retaining the grouping of my a and b
schedule, in line with the usage of Boskoop and with Horni-
prook's nomenclatural grouping, I have preserved a helpful
perspective, which is lost in Cornman's treatment. I believe

1947 van Melle, From Texts to Plants 361

that my investigation of Hornibrook's treatments is at least
as thorough as Cornman's, and more closely related to the
living materials involved; and that Cornman, having failed
to discover the implications and defects noted by me, was at
a disadvantage, and in no position to spank anybody.

He errs, also, in lumping with his var. aureo-variegata
the albo-variegata element, on a basis of preserved speci-
mens. I should not dare do such a thing, knowing that white
variegations of junipers often turn to various pale to deep
shades of yellow on herbarium sheets, in a short or longer
period of time. Actually, the albo-varicgata material dif-
fers from the aureo-variegata both in the color and in the
size of the variegated parts. The effect (as Hornibrook notes
correctly) is a speckled one; - like that of J. Sabina f.
variegata.

Then, having thrown out Hornibrook's albo-variegata list-
ing, he proceeds to transfer its synonym, J. chinensis pro-
cumbens albo-variegata of Beissner (1891), to the var. alba
Rehd. But I shall not go into a detailed criticism of Corn-
man's dispositions. I have limited myself to two of the num-
erous instances in which his sweeping condemation of my
work appears wholly out of order. It would have been un-~
reasonable to expect his thesis on cultivated junipers to
be a work of mature skill. It is an initial effort. If he
is to do effective work in this 1ield, he may well arrange
to swap certain apparent mental attitudes for new ones.

For instance, he may well cultivate a more critically
inquisitive mind in relation to problems of the origins of
meny of these materials. To dismiss them, with a sort of
finality, as "garden forms" and "clons" seems a facetious
and unscientific way of disengaging oneself from one of the
most important and difficuv.ys inquiries concerning them. It
is a charming, but unrealistic thought that such things as
"J. chinensis" vars. pyramidalis, Sheppardii, Parsonsii,
Pfitzeriana, etc., are found in horticultural cabbage
patches.

On p. 244 he says that all the cultivated varieties of
J. chinensis, except var. Sargentii, "so far as has yet
been show, are apparently variants selected from cultivat-
ed plants; most of them are clons." I object particularly to
the word "apparently". For instance, in the case of his var.
Sheppardii (my J. Sheppardii), of which we have in cultiva-
tion monvuscious material and dioecious, in both sexes, and a
diversity of color forms, I do not believe that it can be
"apparent" to any able observer that this aggregate repre-
sents a variant selected from any cultivated kind, let alone,
directly or indirectly from the strictly dioecious "foemina"
material. This goes as well for J. sphaerica Lindl., of
which clearly and obscurely monoecious as well as dioecious

362 Praie eT Ok! Oar lk Vol. 2, no. 9

phases and varieties, in both sexes, ere cultivated. The
Pfitzer Juniper - a selected variant? Of what?

Cornman criticizes me sharply for identifying some of my
junipers with wild materials. I can see nothing but preju-
dice in that attitude. In the case of Japanese specimens
cited by me for my J. Sheppardii var. torulosa (Eastw.), he
does not review these sheets, but passes them by with the
brief, erroneous observation that all are from Honshu, and
none unquestionably wild. Yet they include Wilson's collec-
tion from Yakushima, reported by him as wild, and another of
his collections from the Idzu Peninsula, which he reports as
"seid to be wild". Inasmuch as herbarium specimens of this
variety are mostly very clearly recognizable, I suppose that
Cornman would have to admit them as identical with the cul-
tivated material of torulosa, if he were to account for
them. In the cases of the Sheppardii and sphaerica entities,
he ignores the occurrence in cultivation of monoecious mat-
erials. I believe that I could show him cultivated plants of
J. sphaerica of every degree of monoecism. The thought seems
to be repulsive to him that among the so-called varieties of
J. chinensis there should be perfectly good wild species and
varieties; and he leans over so far in this prejudice that
in a number of instances his judgement seems seriously im-
paired. While he accounts faithfully for every monoecious
species published as such, for instance, by Martinez and
Florin, he jabbers about the monoecism of J. sphaerica Lindl.
representing, perhaps, only an abnormality of the type spec-
imen, while it is abundantly manifest in cultivation.

It seems to me that in relation to the origins of culti-
vated junipers we must either bury all thought under plati-
tudes such as that cited from Cornman's thesis (p. 244), or
else explore every avenue of inquiry. I can see no valid ob-
jection to the traditional practice of checking cultivated
materials of unknown origin against herbarium records of
wild plants, which is what I have done. That, at least, rep-
resents a mode of inquiry into the origins of cultivated
plants. The opposite of it is Cornman's refusal to admit the
wild nature of Faurie's specimens of J. procumbens Sieb. &
Zucc. Bound to diagnose it as a cultivated variety of J.
chinensis, he suggests that Faurie may have had his notes
mixed up.

There is, I think, no more justification, in the absence
of positive evidence, for ascribing to any of these junipers
a garden origin than for declaring them to be wild. The one
assumption is as prejudiced as the other. Yet, I think it is
perfectly legitimate to speculate and conjecture about their
origins, as long as conjectures are clearly presented as
such.

If Cornman is to do effective and intelligible work on

1947 van Melle, From Texts to Plants 363

our cultivated junipers I believe that he will have to doa
sorting-out job on his J. chinensis, which is, as yet, in my
opinion, a loosely inclusive, undefinable concept, based not
upon observation of living plants, but upon the literature
of what he probably believes to be that of J. chinensis, but
which, from the 1850's on, is largely that of J. sphaerica.
He is more likely to arrive at clarity in the matter from
the study of living plants than from the literature.

I regret that his initial contribution becomes for me the
occasion of this rebuttal. I congratulate him upon his or-
dination, and trust that he will increase in wisdom and in
stature; that he will come to contribute much to the know-
ledge of cultivated junipers. He is as yet over~dependent
upon texts. I hope that he may come to work increasingly
from living plants toward texts, and bring with him into
this field a refreshing breeze of first-hand knowledge of
the living plants. That is what is mostly needed.

NOTES CN NEW AND NOTEWORTHY PLANTS. IIT

Harold N. Moldenke

ALOYSIA VIRGATA var. ELLIPTICA (Briq.) Moldenke, comb. nov.

Lippia virgata var. elliptica Briq., Ann. Tonserv. & Jard.
Bot. Genev. 7-6: 304. 1904.

This was published as "var. pie hmete through typo-
graphic error in Phytologia 2: 310. 1947.

BUDDLEIA MEGALOCEPHALA f. ALBILANATA Moldenke, f. nov.

Haec forma a forma typica speciei torentis albis recedit.
-- This form differs from the typical form of the species in
its very dense tomentose pubescence on the branches, branch-
lets, petioles, lower leaf-surfaces, peduncles, and calyxes
being white.

The type was collected by Jacob F. Brenckle (no. 47-283)
at high altitudes in the pine barrens east of Lake Atitlan,
Guatemala, on February 21, 1947, and is deposited in the
Britton Herbarium at the New York Botanical Garden. The col-
lector describes it as a thick and short tree, the old
trees leaning over and supporting on their trunks epiphytic
ferns, orchids, mosses, etc.

CITHAREXYLUM ULEI var. CALVESCENS Moldenke, var. nov.
Haec varietas a forma typica speciei foliis subtus non
hirtellis recedit. -- This variety differs from the typical

364 PEE TRO Llane Dus Vol. 2, noe 9

form of the species in not having its leaf-blades hirtellous
along the larger venation beneath.

The type was collected by R. Froes (no. 1719) near the
river, Candido Mendes, Maracassumé River region, Maranhaéo,
Brazil, on May 2, 1932, and is deposited in the Britton Her-
barium at the New York Botanical Garden. The collector de-
scribes it as a tree 20 feet tall, with whitish flowers, and
records the common name of "tarumd branco".

NURANTA VESTITA var. GLABRESCENS Moldenke, var. nov.

Haec varietas a forma typica speciei ramis remulisque
foliisque inflorescentiisque sparsissime puberulis vel glab-=
ratis recedit. -- This variety differs from the typical form
of the species in having its branches, branchlets, leaves,
and inflorescences only very sparsely puberulent or even
glabrous.

The type was collected by J. Kiehl and A. S. Costa Serra
[Herb. Inst. Agron. do Estado S#o Paulo, Secc. Bot. 4027;
Herb. Inst. Bot. Sao Paulo 42081] at Cascata, Sa&o Paulo,
Brazil, on December 14, 1938, and is deposited in the Brit-
ton Herbarium at the New York Botanical Garden.

ERIOCAULON SCHIMPERI var. GIGAS Moldenke, var. nov.

Haec varietas a forma typica speciei recedit: floribus
masculis tubo sepalorum ca. 2 mm. longo, lobis 4.5 mm. long-
is, et tubo petalorum 5 mm. longo, lobis 2.5 mm. longis;
floribus foeminis sepalis 5.5 mm. longis et petalis Es mmm «
longis. -- This variety differs from the typical form of the
species in having the sepal-tube about 2 mm. long, the lobes
about 4.5 mm. long, and the petal-tube 5 mm. long and its
lobes 2.5 mm. long in the staminate flowers. In the pistill-
ate florets the sepals are 5.5 mm. long and the petals 4.5
mm. long. The heads and leaves also average larger than in
the typical form of the species, and the involucral bract-
lets are more sharply acuminate.

The type was collected by I. R. Dale (no. 3397) in the
Marakwet Hills, Kenya, in Jure of 1935, and is deposited in
the herbarium of the Jardin Botanique de 1'Etat at Brussels.

ERICCAULGN STEINBACHII (Moldenke) Moldenke, comb. nov.

(SO Sees Steinbachii Moldenke, Phytologia 2: 231--232.
1947.

My good friend and colleague, Dr. Alberto Castellanos,

has very kindly pointed out an error in my diagnosis of
this species. The flowers which I had dissected were appar-
ently abnormal or so far past anthesis that the stamen char-
acters were obscure. The normal staminate flowers have six
(not three) stamens. Also, the peduncle is 3-costate (not 2-
costate. The plant is obviously an Eriocaulon, rather than a

1947 Moldenke, New and Noteworthy Plants 365

Paepalanthus. It has been suggested that it may be E. lepto-
hyllum Kunth. In Kunth's species, however, the leaves are
5-85 cm. long, 0.5--1 mm. wide; the peduncles are soli-
tary or sometimes 2 or 3 per plant, S- or 6-costate, and 12=

16 cm. rall; the sheaths are 4 cm. long; the heads are sub-
globose, slightly compressed in drying, white-villous at the
apex; the involucral bractlets are obovate, brunneous-fus-
cous, obtuse; the receptacular bractlets are narrowly obova-
te, subacuminate, puberulent on the back, ciliolate toward
the apex; the staminate florets have the sepals connate to
the middle or beyond, oblong, very obtuse, puberulent on the
back at the top, the petals united half way up, obtuse, pil-
ose, and the anthers black; and the pistillate florets have
the sepals obovate, rather obtuse, puberulent at the top,
and the petals subspatulate, white, pilosulous, and black-
glanduliferous. In E. Steinbachii, on the other hand, the
leaves are 4--5 cm. long, about 1.5 mm. wide at the mid-
point; the peduncles are numerous, about 20 per plant, 3=-
costate, 6--1l1 cm. tall; the sheaths are 3 cm. long; the
heads are hemispheric, not compressed, brown or black, not
white-villous on the top; the involucral bractlets are lan-
ceolate, acute, hyaline or gray; the receptacular bractlets
are narrowly oblong, acute, glabrous throughout; the stamin-
ate florets have the sepals connate at the base only, the
free part elliptic, acute, glabrous throughout, the petals
united 2/3 to 4/5 their length, acute, glabrous throughout,
the anthers yellow; and the pistillate florets have the se-
pals narrow-elliptic or oblong, acute, glabrous throughout,
and the petals narrowly-oblong, brownish, glabrous through-
out, not glanduliferous.

LANTANA MONTEVIDENSIS f. ALBIFLORA Moldenke, f. nov.

Haec forma a forma typica speciei corollis albis recedit.
-- This form differs from the typical form of the species in
having white corollas.

The type was collected by Robert W. Schery (no. 584) in
sandy soil in the treeless “subalpine” area, with Vellozie~
cease associates, at 1000 m. altitude, Municipality of Morro
do ¢ Chapeu, Bahia, Brazil, in April of 1944, and is deposited
in the Britton Herbarium at the New York Botanical Garden.
The plant is described by the collector as 5 dm. tall, with
white flowers.

LIPPIA EKMANI Moldenke, sp. nov.

Herba perennis; caulibuse ut videtur simplicibus rectis
subtetragonis stramineis sparsissime asperulis glabrescenti-
bus; internodis elongatis; foliis ternatis; petiolis subob-
soletis; laminis chartaceis ellipticis acutis vel submucro-.
natis subintegris vel supra mediam paucidentatis, ad basin

366 Po HeY!TiObcO'G: Ia Vol. 2, noe 9

rotundatis vel subacutis, utrinque dense resinoso-punctatis,
supra asperis, subtus laevibus glabratisque; inflorescentiis
terminalibus racemiformibus.

Perennial herb; stems apparently simple and erect, sub-
tetragonal, to about 80 om. tall, stramineous, very sparsely
and obscurely asperulous above, becoming smooth in age; in-
ternodes elongated, 7--8.5 cm. long; leaves ternate; petio-
les inconspicuous or subobsolete, to 2 mm. long, subglabrous
or with a few scattered hairs; blades chartaceous, uniformly
bright-green on both surfaces, elliptic, 2.3--4.5 cm. long,
1.3--2.5 cm. wide, acute or submucronate at the apex, suben=
tire or with a few very much appressed teeth above the mid-
dle, rounded or subacute at the base, densely resinous=punc-
tate on both surfaces, asperous above, smooth and glabrous
(or with a very few scattered microscopic hairs) beneath;
midrib plane or subimpressed above, prominulous beneath;
secondaries very slender, arcuate-ascending, 3 or 4 per
side, terminating in the sinuses between the marginal teeth;
inflorescence terminal, racemiform, 15--20 cm. long, the 4—
6 straight erect sympodia 2--4 cm. long, tetragonal, rather
densely resinous—glandular and puberulent; peduncles similar
to the stems, 4--6 cm. long, tetragonal, asperous~puberulent
and resinous-glandular; heads numerous, the uppermost in
pairs on the rachis, the lower in whorls of 3—=5, the upper
sessile or on slender pilosulous and resinous stalks 2--6
mm. long, the lower on stalks to 2 cm. long; the individual
heads hemispheric, 1--1.5 cm. wide, eventually to 1 cm.
long, many-flowered; bracts 3 at each node of the rachis,
narrowly elliptic, to 1 cm. long and 4 m. wide, attenuate-
subacuminate at the apex, attenuate-acute at the base, ses-
sile, asperulous-puberulent and resinous-glandulose on both
surfaces; bractlets subtending the individual flowers broad-
ly elliptic-subovate, about 5 mm. long and 3 mm. wide, acum-
inate at the apex, asperulous-strigillose and resinous-
glandular on the back; corolla exserted, its tube about 5
mm. long, densely resinous<glandular and puberulous on the
outside, the limb 4—-5 mm. wide, glabrous on both surfaces
or slightly resinous at the base on the outside.

The type of this handsome and very distinct species was
collected by Erik Leonard Ekman (no. 1974) at Posadas, Mis-
iones, Argentina, in 1907 or 1908, and is deposited in the
herbarium of the Naturhistoriska Riksmuseum at Stockholm.

OXERA OBLONGIFOLIA var. ARTENSIS (Dubard) Moldenke, comb.
nov. ?
Oxera neriifolia var. artensis Dubard, Bull. Soc. France

53: 712. 1906.
OXERA OBLONGIFOLIA var. SINUATA (Dubard) Moldenke, comb.nov.

1947 Moldenke, New and Noteworthy Plants 367

Oxera neriifolia var. sinuata Dubard, Bull. Soc. France
Bor 712s 1906.

PAEPALANTHUS WILLIAMSII Moldenke, sp. nov.

Herba perennis caulescens; caulibus elongatis firmis, ju-
ventute villosulo-tomentosis, senectute glabrescentibus
atro-brunneis; foliis graminaceis ad apicem caulis rosulat-
is erecto-patentibus, ad basin dense albo-lanatis; vaginis
multistriatis oblique fissis; pedunculis 8 elongatis 5- vel
6-costatis glabris; capitulis hemisphaericis duris rigidis.

Caulescent perennial herb, apparently at least 7 dm.
tall, probably taller; stems elongate, thin, firm, more or
less villosulous-tomentose, especially on the youngest
parts, the tomentum wearing off in age, exposing a smoothish
deep-brown stem; stem-leaves numerous, rather densely over-
lapping, more or less appressed or the lower ones variously
spreading, grass-like, about 4 cm. long, 3--4 mm. wide at
the mid-point, sharply attenuate at the apex, sessile and
but very slightly narrowed at the base, not clasping, micro-
scopically puberulent or glabrous on both surfaces, often
villosulous at the base with whitish hairs like the stems;
stems terminated by a dense tuft of often somewhat longer,
erecto-spreading, less attenuate or merely acute leaves,
densely whitish-lanate at the base and between the leaves,
the longest leaves to 8 cm. long and 6 mm. wide at the mid-
point, the innermost smallest and only 2.5 cm. long; sheaths
5-—6 cm. long, many-striate, only very slightly twisted,
closely appressed, the rim obliquely split; peduncles about
8 per plant, arising from the center of the terminal tuft of
leaves, 30--32 cm. long, 5- or 6-costate, slightly twisted,
glabrous throughout or microscopically puberulent just be-
neath the head; heads hemispheric, tough, rigid, about 10
mm. in diameter; involucral bractlets very numerous, flaves-
cent-brunneous, in about 5 imbricate series, increasing in
size inwards, scarious, tough, convex on the outer and con-
cave on the inner surface, thicker at the base, lanceolate,
2——-53.5 mm. long, 1.2--2 mm. wide, acute at the apex, glab-
rous and very shiny throughout on both surfaces; receptacle
densely white-villous; receptacular bractlets white, navicu-
lar, narrow-elliptic, about 3.2 mm. long and 1 mm. wide at
the middle, attenuate or subacuminate at the apex, glabrous
on both surfaces; staminate florets on a pedicel about 1 mm.
long: sepals 3, white, separate, navicular, elliptic, about
2mm. long and 0.6 mm. wide, attenuate at apex, puberulous
on the back; petals 3, equal, free almost to the base, hya~
line, elliptic, erect, the free part about 1.2 mm. long and
0.4 mm. wide, somewhat navicular, closely adnate before an-
thesis, acute at apex, glabrous on both surfaces; stamens 35
plainly opposite the petals, inserted at the very base of the

368 PRY TOL 0 Gt A Vol. 2, noe 9

free portion of the petals; filaments 0.4 mm. long, white,
glabrous; anthers oblong, about 0.3 mm. long, dorsifixed
just below the apex, the 2 thecae slightly spreading at the
base; rudimentary 3=-parted pistil yellowish, about 0.2 mm.
long; pistillate florets sessile: sepals 4, separate to the
base, firm, erect, whitish or slightly flavescent, chaffy,
oblong-oblanceolate, about 3 mm. long and 0.6 m. wide,
somewhat navicular, enfolding the rest of the flower, snap-
ping off easily at the base, acute at apex, densely pilose-
pubescent on both surfaces; petals 3, whitish or faintly
flavescent, erect, firm, oblanceolate-elliptic, 1.5--2.1 mm.
long, 0.5--0.6 mm. wide, acuminate at apex (often 3-lacini-
ate in age), long-villous on both surfaces with white hairs,
sometimes less so in age, not bearded, not glanduliferous;
stigmas 3, O.1--C.2 mm. long; style-appendages 3, arising at
the same level as the stigmas and longer than they, 0.3--0.6
mm. long, erecto-spreading, glabrous; style stout, about 0.4
mm. long, glabrous, brown at base; ovary subglobose, about
0.8 mm. long and wide, glabrous, 3-angled, 3-celled, 3-
ovulate.

The type of this species was collected by Llewelyn Willi-
ams (no. 15051) -- in whose honor it is named -- in low
places near the palm grove "Carandé" in the savanna of San
Antonio, Rfo Orinoco, Amazonas, Venezuela, at an altitude of
121 m., on April 27, 1942, and is deposited in the Britton
Herbarium at the New York Botanical Garden. It is a great
pleasure to dedicate this species to this noted collector
and botanist, who has done such wonderful work on the flora
of Venezuela and Peru.

POLYGALA CURTISSII f. ALBA Moldenke, f. nov.

Haec forma a forma typica speciei racemis densis et flor-
ibus albis recedit. -- This form differs from the typical
form of the species in its densely congested inflorescences
and white flowers.

The type was collected by me (no. 19269) on road embank-
ments and shoulders along the Blue Ridge Mountains Parkway
near Galax, Carroll 0o., Virginia, on August 28, 1947, and
is deposited in the herbarium of Oregon State College.

PORTULACA GRANDIFLORA f. PLENA Moldenke, f. nov.

Haec forma a forma typica speciei corollis plenis reced-
it. -- This form differs from the typical form of the spe-
cies in its "doubled" corollas.

The type was collected by me (no. 4118) from cultivated
material at Watchung, Somerset Co., New Jersey, on September
3, 1928, and is deposited in the Britton Herbarium at the
New York Botanical Garden. I cannot find that this form has
hitherto been validly named in botanical literature.

1947 Moldenke, New and Noteworthy Plants 369

STACHYTARPHETA LAEVIS Moldenke, sp. nov.

Herba robusta; ramis tetragonis ubique glabris nitidis
brunnescentibus; foliis oppositis; petiolis gracilibus alat-
is ubique glabris; laminis leviter chartaceis ovatis brun-
nescentibus subacutis vel obtusis, ad basin rotundatis, reg-
ulariter serratis utrinque glabris nitidisque; spicis ter-
minalibus solitariis elongatis ubique glabris.

Coarse herb; branches tetragonal, completely glabrous and
shiny, brunnescent; principal internodes 3--5 cm. long;
leaves decussate-opposite, usually with a cluster of very
small ones on much abbreviated twigs in their axils; petio-
les slender, 2--10 m. long, winged, completely glabrous;
blades thin-chartaceous, ovate, brunnescent in drying, 2.5--
5 cm. long, 1--2.8 cm. wide, subacute or blunt at apex,
rounded into the broadly winged petiole at base, uniformly
serrate from base to apex with acute or submucronate teeth,
completely glabrous and shiny on both surfaces; midrib and
the 4 or 5 pairs of slender secondaries plane above, promin-
ulous beneath; vein and veinlet reticulation obscure, usual-
ly only the larger tertiaries discernible; spikes terminal,
solitary, elongate, to about 36 cm. long, about 1 cm. wide
(including the corollas); peduncle very short or obsolete;
rachis slender, glabrous, sculptured after anthesis; bract~
lets narrow-lanceolate, about & mm. long and 1 mm.” wide,
long-attenuate at apex, somewhat scarious-margined, glabrous
or subglabrate, subappressed, slightly shorter than the cal-
yx; corolla about 1 cm. long, pale=blue, the limb 5 m. wide.

The type of this species was collected by Carl Axel Mag-
nus Lindman (no. A.607) in shady places along roadsides,
Porto Alegre, Rio Grande do Sul, Brazil, on November 3,
1892, and is deposited in the herbarium of the Naturhistor-
iska Riksmuseum at Stockholm. The collector states that the
plant is used medicinally as a stomachic.

STACHYTARPHETA LOEFGRENI Moldenke, sp. nov.

Frutex; ramis gracilibus tetragonis submarginatis densis-
sime pubescentibus vel subvillosis; foliis oppositis; petio-
lis alatis dense villoso-pubescentibus; laminis chartaceis
late ellipticis acutis vel subacuminatis, ad basin longe ac-
uminatis supra mediam serratis supra scabridis pilosulisque
subtus dense velutino-tomentellis; spicis terminalibus ut
videtur 3 dense mltifloris; bracteolis anguste lanceolatis
longe attenuatis vel caudatis valde divergentibus, densiss~-
ime longeque ciliatis.

Shrub; branches slender, tetragonal, slightly margined at
the angles, very densely pubescent or subvillous, less
densely so in age, the hairs yellowish when ycung, grayish
in age; principal internodes 2--8 cm. long, more abbreviated
on the younger parts; leaves decussate-opposite; petioles 5=

370 Poo ET Orb Oo ast £ Vol. 2, noe 9

10 mm. long, winged, densely villous-pubescent with yellow-
ish or white hairs, flattened above; blades chartaceous,
grayish-green on both surfaces, broadly elliptic, to about
9.5 cme long and 5 cm. wide, acute or subacuminate at the a-
pex, long-acuminate into the winged petiole at the base,
serrate from the middle to the apex with broad=based, round-
ed, and rather appressed teeth, scabridous and more or less
pilosulous above, densely velvety-tomentellous beneath; mid=
rib plane or very slightly subimpressed above, prominent be-
neath; secondaries slender, about 5 per side, plane or very
slightly subimpressed above, prominulous beneath; vein and
veinlet reticulation fine, the larger parts plainly visible;
spikes terminal, apparently 3, the two lower ones shorter,
all densely many-flowered, to about 8 cm. long, to 2.5 cm.
wide; peduncle obsolete or very short; rachis completely
hidden by the closely imbricate flowers; bractlets narrow-
lanceolate, 11~15 mm. long, about 2 mm. wide at the base,
long-attenuate or caudate at apex, widely divergent from the
rachis during anthesis, very densely long-ciliate on the
margins, otherwise glabrate or obscurely pilosulous; corolla
very large and showy, about 3 cm. long.

The type of this most distinctive species was collected
by Alberto Léfgren (no. 692) in "caatinga" at Ingazeiro,
Ceard, Brazil, on April 26, 1910, and is deposited in the
herbarium of the Naturhistoriska Riksmuseum at Stockholm.

STACHYTARPHETA MOLLIS Moldenke, sp. nov.

Frutex; ramis obsolete tetragonis densissime tomentosis;
foliis firme chartaceis oppositis sessilibus, laminis ellip-
ticis acutis, ad basin rotundatis, supra mediam serratis,
utrinque dense villosis; costa venis majoribusque supra im-
pressis subtus valde prominentibus; spicis solitariis ter-
minalibus brevibus ubique dense villosis.

Shrub; branches obsoletely tetragonal, very densely to-
mentose with sordid-gray or brownish hair; principal inter-
nodes 4~-7 cm. long; leaves firmly chartaceous, opposite,
sessile, the blades elliptic, 3.5--5 cm. long, 2--3 cm.
wide, acute at apex, rounded at base, serrate from the mid-
dle to the apex with rather coarse and rounded more or less
appressed teeth, densely villous on both surfaces with sord-
id-gray hairs; midrib, the 3--5 slender secondaries, and the
larger veinlets impressed above and prominent beneath; spike
solitary, terminal, short, about 7 cme long, densely villous
throughout, about 2 cm. in diameter (exclusive of corollas);
peduncle very short, densely villous; rachis densely villous
but completely hidden by the closely imbricate flowers;
bractlets lanceolate, 11--12 mm. long, 2--3 mm. wide at the
base, long-attenuate or acuminate at apex, densely villous
on the back, equaling or slightly exceeding the villous cal=-

1947 Moldenke, New and Noteworthy Plants 371

yx; corolla brick-red, showy.

The type of this distinct species was collected by Aug-
uste Frangois Marie Glaziou (no. 21906) between Sobradinho
and Lagoa do Mestre d'Armas, Goyaz, Brazil, in November or
December, and is deposited in the herbarium of the Naturhis-
toriska Riksmseum at Stockholm.

STACHYTARPHETA SESSILIS Moldenke, sp. nov.

Herba pumila; caule simplice vel l-ramuloso gracile
stramineo tetragono laxe patenti-hirsuto; foliis oppositis
sessilibus membranaceis oblongis argute acutis, ad basin ro-
tundatis vel subtruncatis, serratis utrinque sparse hirsutu-
lis; spicis solitariis terminalibus; rhachide crasso albo~
hirsutulo post anthesin profunde excavato; bracteolis magnis
lanceolatis acuminatis argute adpressis stramineis subglab-
ratis vel minutissime ciliolatis.

Low herb, to about 35 cm. tall; stems simple or with one
erect branch, slender, stramineous, tetragonal, loosely hir-
sute with soft white hairs 1--2 mm. long standing at right
angles to the stem, normally equally hirsute from apex to
base; principal internodes 2--4 cm. long; leaves decussate-
opposite, sessile, membranous, oblong, 2.5--3.5 cm. long,
1.1--1.2 cm. wide, sharply acute at the apex, rounded or
subtruncate at the base, serrate with rather coarse and short
teeth from the base to the apex, sparsely hirsutulous on
both surfaces, more densely so along the midrib beneath;
midrib very slender, plane above, prominulous beneath; sec-
ondaries very slender, about 5 per side, practically indis-
cernible above, faint beneath; veinlet reticulation indis-
cernible; spikes solitary, terminal, to about 16 cm. long,
about 5 mm. wide (exclusive of the corollas); peduncles ob-
solete; rachis stout, whitish-hirsutulous, deeply excavated
after anthesis, plainly visible between the bractlets;
bractlets large, lanceolate, 8--9 mm. long, 2--2.5 mm. wide,
acuminate at the apex, barely contiguous and not at all im-
bricate at maturity, closely appressed to the rachis, sub-
glabrous and stramineous or very minutely ciliolate toward
the apex and strigillose-pilose toward the base; calyx equal
to or slightly shorter than the bractlets, microscopically
strigillose; corolla-tube 10--12 mm. long, glabrous, the
limb about 1 cm. wide.

The type of this unmistakable species was collected by
Alberto Léfgren (no. 160) in "caatinga" at Salvacéo, Ceard,
Brazil, on March 6, 1910, and is deposited in the herbarium
of the Naturhistoriska Riksmuseum at Stockholm.

SYNGONANTHUS AKURIMENSIS Moldenke, sp. nov.
Herba perpumila acaulescens; foliis rosulatis numerosis
linearibus recurvatis densiuscule patenti-~pubescentibus, ad

ae PHYTOLOGIA : Vol. 2, no. 9

apicem obtusis saepe recurvatis; pedunculis numerosis tri-
costatis gracillimis paulo tortis glabris stramineis; vagin-
is arcte adpressis dense incanis, pilis arctissime adpressis
inflatis minutis bulboideis; capitulis hemisphaericis albid-
is vel niveis, parvis 2--4 mm. latis.

Very dwarf acaulescent herb; leaves rosulate, numerous,
linear, recurved, 5--15 mm. long, 0.5--1 mm. wide, rather
densely spreading=-pubescent with short white hairs, less so
in age, blunt and often subuncinately recurved at the apex;
peduncles several to 8 per plant, 2--5 cm. long, 3-costate,
very slender, slightly twisted, glabrous, stramineous;
sheaths 5-10 mm. long, closely appressed, densely incanous
with very closely appressed inflated minute bulb-like hairs
which are usually without any capillary appendage, but some-
times appendaged ones are interspersed, obliquely split at
the apex; heads hemispheric, white or whitish, 2--4 mm. in

diameter; involucral bractlets oblanceolate, white or stram-

ineous, about 1.5 mm. long and 0.8 mm. wide, rounded in out-
line but irregularly erose-laciniate at the apex, glabrous
and shiny on both surfaces; receptacle densely long-pilose
with white hairs; staminate florets: borne on a capillary
stalk about 0.6 mm. long; sepals 3, hyaline, separate, ell-
iptic, about 0.7 mm. long and 0.3 mm. wide, glabrous, not
glanduliferous; petals 3, hyaline, apparently separate (7),
of the same size, shape, and texture as the sepals, glabrous
and not glanduliferous; pistillate florets: sepals 3, separ-
ate, hyaline, oblong, about 1.9 mm. long and 0.8 mm. wide,
acute at the apex, glabrous; petals 3, linear, hyaline, con-
nate at the middle, but easily separating in age, about 1.1
mm. long and 0.2 mm. wide, long=-pilose near the middle on
the inner surface with hyaline hairs reaching about to the
apex, not glanduliferous; style subobsolete or to 0.1 m.
long, glabrous, its appendages 3, about O.4 mm. long; stig-
mas 3, about 0.2 mm. long; ovary 3-celled. .

The type was collected by Francisco Tamayo (no. 3234) in
sandy soil on Cerro Akurimd, Bolfvar, Venezuela, in March of
1946, and is deposited in the United States National Herbar-
ium at Washington. The collector says that the plants grow
from 5 to 8 cm. tall. The remarkable hairs of the sheaths
are most characteristic.

THE KNOWN GEOGRAPHIC DISTRIBUTION OF THE MEMBERS OF THE
ERIOCAULACEAE. SUPPLEMENT 2

Harold N. Moldenke

The following records are a continuation of the series

.

1947
begun by me in

Moldenke, Eriocaulaceasae 212

Phytologia 2: 349--352 (1947).

UNITED STATES OF AMERICA:

New York:
Eriocaulon

Delaware:
Eriocaulon
Eriocaulon

septangulare With. (Cayuga County )

compressum Lam. (Sussex County )
septangulare With. (New Castle County )

North Carolina:
Lachnocaulon anceps (Walt.) Morong (Pender County )
Lachnocaulon minus (Chapm.) Small (Bladen County )

Michigan:
Eriocaulon
Texas:
Eriocaulon
California:
Eriocaulon
COLOMBIA:
Paepalanthus
VEN EZUELA:
Paepalanthus
Paepalanthus
Paepalanthus
Syngonanthus
SUR IN AM:
Paepalanthus
Paepalanthus
Paepalanthus
Syngonanthus
the =<"
Syngonanthus

Syngonanthus
FRENCH GUTANA:

septangulare With. (Baraga County )

texense Kérn. (Jefferson County )

cinereum R. Br. (Stanislaus County)
ensifolius (H.B.K.) Kunth (Santander Norte)

Tatei Moldenke (Ler)
truxillensis Kérn. (Larg)
Williamsii Moldenke (Amazonas )*
akurimensis Moldenke (Bolfvar)*

Maguirei Moldenke

polytrichoides Kunth
tafelbergensis Moldenke*

eriophyllus var. glandulosus Ruhl. -- delete

gracilis (Kérn.) Ruhl.
surinamensis Moldenke*

Eriocaulon guianense Kérn. -- delete the acid

Syngonanthus
ECUADOR:

Paepalanthus

Paepalanthus
Loja)
Paepalanthus
Paepalanthus
Paepalanthus
Syngonanthus
PERU:
Paepalanthus

Syngonanthus
BRAZIL:

caulescens (Poir.) Ruhl.

andicola Kérn. (Loja)
ensifolius (H.B.K.) Kunth (Azuay, Carchi, &

Espinosianus Moldenke (Santiago-Zamora)*
Karstenii Ruhl. (Loja)

loxensis Moldenke (Loja)*

macrocaulon Ruhl.

pilosus (H.B-K.) Kunth (Cuzco)
caulescens (Poir.) Ruhl. (San Mart{fn)

Ericcaulon Beauverdi Moldenke (Sao Paulo)*
Briocaulon cipoense Alv. Silv. (Minas Geraes )*

374 PHYTOLOGIA Vol. 2, no. 9

Eriocaulon crassiscapum Bong. (Rio de Janeiro & Rio Grande
do Sul) -- delete the "*!

Briocaulon dictyophyllum Ktrn. (Parang)

Eriocaulon gibbosum Kérn. (Goyaz & Rio de Janeiro)

Eriocaulon giganteum (Eeauverd) Beauverd -~ to be deleted

Eriocaulon heteropeplon Alv. Silv. (Minas Geraes )*

Eriocaulon Humboldtii Kunth (Amazonas )

Eriocaulon leptophyllum Kunth -- delete the "*"

Eriocaulon longepedunculatum Alv. Silv. -- to be deleted

Eriocaulon melanocephalum Kunth (Amazonas )

Eriocaulon modestum Kunth -~ delete the "*"

Eriocaulon paludicola Alv. Silv. is the correct form for
this name

Eriocaulon Silveirae Moldenke (Minas Geraes)*

Leiothrix. cuscutoides Alv. Silv. (Minas Geraes )*

Leiothrix Edwallii Alv. Silv. (Sao Paulo) *

Leiothrix hirsuta var. Magalha@esii Alv. Silv. (Minas Gera-
es )*

Leiothrix obtusifolia Alv. Silv. (Minas Geraes)*

Leiothrix sclerophylla Alv. Silv. (Minas Geraes)*

Paepalanthus Dusenii Ruhl. (Sao Paulo)

Paepalanthus fasciculatus (Rottb.) Kérn. (Pard)

Paepalanthus myocephalus (Mart.) Kérn. (Pernambuco)

Paepalanthus neopulvinatus Moldenke (Minas Geraes )*

Paepalanthus paucifolius Alv. Silv. (Minas Geraes)*

Paepalanthus pulvinatus Alv. Silv. -- to be deleted

Paepalanthus Warmingianus Kérn. (Minas Geraes)
Syngonanthus anthemiflorus (Bong.) Ruhl. -- delete the "*"

Syngonanthus caulescens (Poir.) Ruhl. (Parané, Pernambuco,
& Rio Grande do Sul)

Syngonanthus centauroides var. subappressus Ruhl. (Rio
Grande do Sul

Syngonanthus Fischerianus (Eong.) Ruhl. (Amazonas & Para)

Syngonanthus gracilis var. glabriusculus Ruhl. -- delete

the "*
Syngonanthus pracilis var. hirtellus (Steud.) Ruhl. -- de-
lete the "*

Syngonanthus heteropeploides Herzog —- delete the "*"
Syngonanthus rufo-albus Alv. Silv. (Minas Geraes)*

Syngonanthus Widgrenianus (Kérn.) Ruhl. (Sao Paulo)
BOLIVIA:

Eriocaulon Steinbachii (Moldenke) Moldenke (Santa Cruz)*

Leiothrix flavescens (Bong.) Ruhl. (El Beni)

Paepalanthus muscosus Kérn. (La Paz) ;

Paepalanthus speciosus (Bong.) Ruhl. (Santa Cruz)

Syngonanthus caulescens (Poir.) Ruhl. (Santa Cruz)

Syngonanthus Fischerianus (Bong.) Ruhl. (Santa Cruz)
PARAGUAY:

Eriocaulon magnum Abbiatti

1947 Moldenke, Eriocaulaceae 315

URUGUAY:
Eriocaulon Arechavaletae Herter is the correct form for
this name
Eriocaulon modestum Kunth
ARGENTINA:

Eriocaulon Arechavaletae Moldenke -- to be deleted
Eriocaulon crassiscapum Bong. (Misiones)
Eriocaulon leptophyllum Kunth (Corrientes )
Eriocaulon magnum Abbiatti (Chaco)
Eriocaulon missionum Castell. (Misiones )*
Eriocaulon Sellowianum Kunth (Corrientes )
Eriocaulon sp. indet. — to be deleted
Syngonanthus anthemiflorus (Bong.) Ruhl. (Misiones)
Syngonanthus caulescens (Poir.) Ruhl. (Misiones)
UNION OF SOCIALIST SCVIET REPUBLICS:
Eriocaulon Fuergerianum Kérn. (Buryato-Mongolskaya)
Eriocaulon chinorossicum Lom (Far Eastern Territory )*
Eriocaulon nipponicum Maxim. (Buryato-Mongolskaya)
Eriocaulon robustius (Maxim.) Mak. (Far Zastern Territory }*
ABYSSINIA:
Eriocaulon Schimperi Kérn. -- delete the "*"
SIERRA LEONE:
Paepalanthus Lamarckii Kunth
FRENCH WEST AFRICA:
Eriocaulon bifistulosum Van Heurck & Muell.-Arg. (French
Soudan )
FRENCH EQUATORIAL AFRICA:
Paepalanthus Lamarckii Kunth (Gabun)
BELGIAN CONGO:
Eriocaulon Schimperi Kérn.
RUANDA & URUNDI:
Eriocaulon Schimperi Kérn.
UGANDA:
Eriocaulon Volkensii Engl.
TANGANYIKA TERRITORY:
Eriocaulon mesanthemoides Ruhl. — delete the "*"
Eriocaulon Schimperi Kérn.
Eriocaulon Volkensii Engl. -- delete the "*"
MAFIA ISLAND:
Paepalanthus Lamarckii Kunth
KENYA:
Eriocaulon Schimperi var. gigas Moldenke*
BRITISH NYASALAND PROTECTCRATE:
Eriocaulon mesanthemoides Ruhl.
Eriocaulon Schimperi Kérn.
INDIA:
Eriocaulon Dalzellii Kérn. (Bengal) -- delete the "*"
Eriocaulon Dianae Fyson -- delete the "*"
Eriocaulon luzulaefolium Mart. (Madras )

376 PERSE Ob OG a Vol. 2, noe 9

Eriocaulon Vanheurckii Muell.-Arg. -- delete the "*"
SALSETTE ISLAND:

Eriocaulon Dianae Fyson

Eriocaulon Vanheurckii Muell.—Arg.

BURMA?
Eriocaulon cinereum R. Br.
CEYLON:
Eriocaulon Dalzellii Kérn.
CHINA:

Eriocaulon alpestre Hook. f. & Thoms. (Fukien)
Eriocaulon Puergerianum Kérn. (Kwangtung & YUnnan)
Eriocaulon cristatum Mart.
Eriocaulon Rockii Moldenke (Ytinnan)*
Eriocaulon truncatum Hamilt. (Kiangsi & Kweichow)
Eriocaulon yunnanense Moldenke (Ytinnan)*
JAPAN :
Eriocaulon atrum Nakai (Honshiu)
Eriocaulon cinereum Re Br. (Musashi)
Eriocaulon nipponicum Maxim. (Honshiu)
Eriocaulon robustius (Maxim.) Mak. is the correct form for
this name
Eriocaulon sikokianum Maxim. (Hilachi & Kiushiu)
HAINAN ISLAND:
Eriocaulon Buergerianum Ko6rn.
FRENCH INDO-CHINAs
Eriocaulon Robinsonii Moldenke (Annam)*
Eriocaulon truncatum Hamilt. (Annam)
THAILAND:
Eriocaulon glabriflorum Ridl.
Eriocaulon ubonense H. Lecomte
TERUTAU ISLAND:
Eriocaulon glabriflorum Ridl.
FEDERATED MALAY STATES:
Eriocaulon glabriflorum Ridl. -- delete the "*"
STRAITS SETTLEMENTS:
Eriocaulon cristatum Mart. (Malacca)
Eriocaulon glabriflorum Ridl. (Langkawi Islands)
LIUKIU ISLANDS:
Eriocaulon australe R. Br. (Iriomote Island)
BISMARK ARCHIPELAGO:
Eriocaulon brachypeplon Kérn. (New Ireland)*
AUSTRALIA:
Eriocaulon graphitinum F. Muell. & Tate (South Austral-
ia)*; this is the correct form for this name
ea pe spectabile F. Muell. (Northern Territory &
Queensland )*
Eriocaulon Tatei Ruhl. (Northern Territory )*
Eriocaulon tortuosum F. Muell. (Northern Territory )*
FOSSILIZED:

1947 Moldenke, Eriocaulaceae 377

Eriocaulon porosum Lesq. (Eocene of Colorado)*

Addenda and errata to the alphabetic list of scientific
names proposed in the Eriocaulaceae, including mis-spell-
ings and mis-eccreditions

Carptotepala Moldenke
Carptotepala insolita Moldenke

Dupatya caulescens (Poir.) Kuntze = Syngonanthus caulescens
(Poir.) Ruhl.

Dupatya elegans (Bong.) Kuntze = Syngonanthus elegans (Bong.)
Ruhl.

Dupotya Kuntze = Paepalanthus Mart.

Dupotya flavidula (Michx.) Kuntze = Syngonanthus flavidulus
(Michx.) Ruhl.

Eriaucolon L. = Eriocaulon L.

Eriaucolon elongatum Bong. = Paepalanthus elongatus (Bong.)
Kérn.

Eriaucolon gnaphalodes Michx. = Eriocaulon compressum Lam.

Eriocaulon alatum H. Lecomte

Eriocaulon alpestre var. perpusillum Nakai

Eriocaulon alpestre var. robustius Maxim. = Eriocaulon rob-
ustius (Maxim.) Mak.

Sriocaulon annamense H. Lecomte

Eriocaulon Arechavaletae Castell. = Eriocaulon magnum Abbi-
atti

Eriocaulon Arechavaletae Herter

Eriocaulon Arechavaletae Moldenke = Eriocaulon Arechavaletae
Herter

Eriocaulon argenteum Heyne = Eriocaulon quinguangulare L.

Eriocaulon argentinum Castell. = Eriocaulon leptophyllum

Kunth
Eriocaulon articulatum (Huds.) Morong = Eriocaulon septangu-
lare With.

Eriocaulon articulatum var. submersumn Haberer = Eriocaulon
septangulare With.

Eriocaulon atratum Thwaites = Eriocaulon subglaucum Ruhl.
not E. subcaulescens Hook. f.

Eriocaulon atrum Nakai

Eriocaulon barba-caprae Fyson

Eriocaulon Beauverdi Moldenke

Eriocaulon Boni H. Lecomte

Eriocaulon brevipedunculatum Merr.

Eriocaulon brizoides (Kunth) Steud. = Syngonanthus gracilis
var. Koernickeanus Ruhl.

Eriocaulon bromelloideum H. Lecomte

Eriocaulon Brownianum var. nilagirense Fyson = Eriocaulon

nilagirense Steud.

378 PHYTOLOGIA Vol. 2, no. 9

Eriocaulon capitulatum Moldenke

Eriocaulon caricifoliur Gardn. = Syngonanthus laricifolius
Gardn.) Ruhl.

Eriocaulon caulescens Kunth = Syngonanthus caulescens
Poir.) Rual.

Eriocaulon cauliferum Mak.

Eriocaulon chinorossicum Lom

Eriocaulon Christopheri Fyson

Eriocaulon cipoense Alv. Silv.

Eriocaulon Comptonii Rendle

Eriocaulon congense Moldenke = Eriocaulon Schimperi Kérn.

Eriocaulon conicum (Fyson) C. &. 0. Fischer

Eriocaulon coreanum H. Lecomte

Eriocaulon cristatum Heyne = Eriocaulon cristatum Mart.

Eriocaulon Cuatrecasasi Moldenke = Dichromena monostachya
TBSckl.) C. B. Clarke, Cyperaceae

Eriocaulon cubralense Alv. Silv. = Eriocaulon cabralense
Alv. Silv.

Eriocaulon decemangulare L. = Eriocaulon decangulare L.

Eriocaulon dimorphopetalum Moldenke

Eriocaulon evoideum Britton & Small = Eriocaulon ovoideum
Britton & Small

Eriocaulon falcatum Bong.

Eriocaulon gibbosum var. brevifolium Kérn. = Eriocaulon
gibbosum K6érn.

Eriocaulon gibbosum var. longifolium K6rn. = Eriocaulon
gibbosum Kérn.

Eriocaulon gnaphaloides Michx. = Eriocaulon compressum Lam.

Eriocaulon gnapholoides Michx. = Eriocaulon compressum Lam.

Eriocaulon graphiticum Tate = Eriocaulon graphitinum F.
Muell. & Tate

Eriocaulon graphitinum F. Muell. & Tate

Eriocaulon heterodoxum Moldenke

Eriocaulon heteropeplon Alv. Silv.

Briocaulon hexangulare L. = Eriocaulon sexangulare L.

Eriocaulon leptodictyon A. Gray = Eupatorium leptodictyon
A. Gray, Carduaceae

Eriocaulon ligulatum Bon. = Paepalanthus lingulatus (Bong.)
Kunth

Eriocaulon ligulatus Bong. = Paepalanthus lingulatus
Bong.) Kunth

Eriocaulon lingulatus Bong. = Paepalanthus lingulatus
Bong.) Kunth

Eriocaulon missionum Castell.

Eriocaulon palludicola Alv. Silv. = Eriocaulon paludicola
Alv. Silv.

Eriocaulon paradoxum Moldenke

Eriocaulon quadriangulare Lour. = Eriocaulon sexangulare L.

Eriocaulon quinquangulare Heyne = Eriocaulon cristatum Mart.

1947 Moldenke, Eriocaulaceae 379

Eriocaulon Robinsonii Moldenke _

Eriocaulon robustium (Maxim.) Mak. = Eriocaulon robustius
Maxim.) Mak.

Eriocaulon robustius (“axim.) Mak.

Eriocaulon robustum var. caulescens Fyson = Eriocaulon atra-
tum var. major Thwaites

Eriocaulon Rockii Moldenke

Eriocaulon Schimperi var. gigas Moldenke

Eriocaulon septangulare Kunth = Eriocaulon septangulare With.

‘Eriocaulon septentrionalis (Huds.) Morong = Eriocaulon sep-
tangulare With.

Eriocaulon serotinum Walt. = Eriocaulon decangulare L.

Eriocaulon setaceum Hook. f. = Eriocaulon intermedium K@érn.

Eriocaulon sexangulare Fyson = Eriocaulon longifolium Nees

Eriocaulon Sieboldianum Steud. = Eriocaulon cinereum R. Br.

Eriocaulon Steinbachii (Moldenke) Moldenke

Eriocaulon Steyermarkii Moldenke

Eriocaulon subacaulescens Hook. f. = Eriocaulon subcaules-
cens Hook. f.

Eriocaulon Volkensii var. Mildbraedii Ruhl.

Eriocaulon Wightianum Hook. f. = Eriocaulon robusto-Brownia-
num Ruhl. & E. Brownianum Mart.

Eriocaulon yunnanense Moldenke

Eriocaulon zeylanicumKérn. = Eriocaulon ceylanicum K6érn.

Eriocaulon zeylanicum var. subcaulescens Fyson = Eriocaulon
subglaucum Ruhl.

Eriocaulon /-angulare With. = Eriocaulon septangulare With.

Hyphydra amplexicaulis Vahl = Tonina fluviatilis Aubl.

Lachnenthes Michauxii Kunth = Lachnocaulon anceps (Walt.)
Morong >

Leiothrix.cuscutoides Alv. Silv.

Leiothrix Edwallii Alv. Silv.

Leiothrix hirsuta var. Magalha&esii Alv. Silv.

Leiothrix obtusifolia Alv. Silv.

Leiothrix sclerophylla Alv. Silv.

Leiothrix Steyermarkii Moldenke

Leiothrix umbratilis Moldenke

Peepalanthus aerens Alv. Silv. = Paspalanthus aereus Alv.
Silv.

Paepalanthus aereus Alv. Silv.

Paepalanthus angustus Alv. Silv. = Paepalanthus augustus

Bigs Silv.
Paepalanthus anreus Alv. Silv. = Paepalanthus aureus Alv.
Silv. .

Paepalanthus arborecens Alv. Silv. = Paepalanthus arbores~
cens Aly. Silv.

Paepalanthus bifidus Kunth = Paepalanthus bifidus (Schrad.)
Kunth

Peepalanthus brunneus Moldenke

380 PHYTOLOGIA Vol. 2, noe 9

Paepalanthus caulescens (Poir.) Kunth = Syngonanthus caules-
cens (Poir.) Ruhl.

Paepalanthus desinfolius Alv. Silv. = Paepalanthus densi fol-
ius Alv. Silv.

Paepalanthus diversiflius Alv. Silv. = Paepalanthus diversi-~
folius Alv. Silv.

Paepalanthus elegans (Bong.) Kunth = Syngonanthus elegans
(Kérn.) Ruhl.

Paepalanthus elegans Mart. = Syngonanthus elegans (Kérn.)
Ruhl.

Paepalanthus Espinosianus Moldenke

Paepalanthus filipes Moldenke

Paspalanthus glancopodus Alv. Silv. = Paepalanthus glaucopo-
dus Alv. Silv.

Peepalanthus griseus Moldenke

Paepalanthus Gustarvii Alv. Silv. = Paepalanthus Gustavii
Alv. Silv.

Paepalanthus jordadensis Alv. Silv. = Paepalanthus jordanen-
sis’ Alv. Silv.

Paepalanthus loxensis Moldenke

Paepalanthus Maguirei Moldenke
Paepalanthus myocephalus Mart. = Paepalanthus myocephalus
(Mart.

) Korn.
Paepalanthus myrlocephalus Mart. = Paepalanthus myocephalus
(Mart.) Kérn.

Paepalanthus myriophylus Alv. Silv. = Paepalanthus myrio-
phyllus Alv. Silv.

Paepalanthus oereus Alv. Silv. = Paepalanthus eaereus Alv.
Silv.

Paepalanthus orthogonolis Alv. Silv. = Paepalanthus Ae pore
nalis Alv. Silv.

Paepalanthus paucifolius Alv. Silv.

Paepalanthus pauper Moldenke

Paepalanthus perplexans Moldenke

Paepalanthus pisrophorus Alv. Silv. = Paepalanthus spiro-
phorus Alv. Silv.

Paepalanthus polyclados Alv. Silv. = Paepalanthus polycladus
Alv. Silv.

Paepalanthus ramosissimos Alv. Silv. = Paepalanthus ramosis=-
simus Alv. Silv.

Paepalanthus rhyzocephalus Alv. Silv. = Paepalanthus rhizo-

cephalus Alv. Silv.

Paepalanthus robustns Alv. Silv. = Paopalanthus robustus
Alv. Silv.

Paepalanthus roraimensis Moldenke

Paepalanthus scopulorum Moldenke
Paepalanthus serralapensis Moldenke

Paepalanthus squamuliferus Moldenke
Paspalanthus Steinbachii Moldenke = Eriocaulon Steinbachii

1947 Moldenke, Eriocaulaceae 381

(Moldenke) Moldenke

Paepalanthus Steyermarkii Moldenke
Paepalanthus subsessilis Moldenke

Paepalanthus syngonan-thoides Alv. Silv. = Paepalanthus syn-
gonanthoides Alv. Silv.

Paepalanthus tafelbergensis Moldenke
Paepalanthus tenuicculis Alv. Silv. = Paepalanthus tenuicau-
lis Alv. Silv.

Paepalanthus tortilis Mart. = Paepalanthus tortilis (Bong.)

Marte

Paepalanthus umbillatus Kunth = Syngonenthus umbellatus
(Lam. ) Ruhl.

Paepalanthus vehetinus Alv. Silv. = Paepalanthus velutinus
Alv. Si lv e

Paepalanthus viridifolius Alv. Silv. = Paepalanthus rigidi-
folius Alv. Silv.

Paepalanthus Warmingii Kérn. = Paepalanthus Warmingianus
Kérn.

Paepalanthus Williamsii Moldenke

Paepalanthus sp. Niederlein = Eriocaulon argentinum Castell.

Rhondonanthus Herzog = Rondonanthus Herzog

Rondonanthus micropetalus Moldenke

Syngonanthus akurimensis Moldenke

Syngonanthus caulescens Ruhl. = Syngonanthus caulescens
Poir.) Ruhl.

Syngonanthus duidae Moldenke
ngonanthus elegans (Bong.) Ruhl. = Syngonenthus elegans
Kérn.) Ruhl.

Syngonanthus elegans var. rufescens Ruhl. = Syngonanthus
elegans (Kérn.) Ruhl.

Syngonenthus ‘gracilis Molfino = Eriocaulon argentinum
Castell.

Syngonanthus guianensis Moldenke

Syngonanthus lagopodioides (Michx.) Ruhl. = Syngonanthus
lagopodioides (Griseb.) Ruhl.

Syngonanthus minutulus (Steud.) Moldenke

Syngonanthus rufo-albus Alv. Silv.

Syngonanthus savannarum Moldenke

Syngonanthus surinamensis Moldenke

Syngonanthus venezuelensis Moldenke

Syngonenthus Wilsoni Moldenke = Syngonanthus Wilsonii Mol-
denke

The 161 names in the preceding list are supplementary to

the 2060 names listed by me on pages 28 to 60 of my booklet

entitled "The known geographic distribution of the members

of the Eriocaulacesae", published in 1946.

THE KNOWN GEOGRAPHIC DISTRIBUTION OF THE MEMBERS OF THE _
VERBENACEAE, AVICENNIACEAE, STILBACEAE, AND SYMPHOREMACEAE.
SUPPLEMENT 7

Harold N. Moldenke

CANADA:
Ontario:
Verbena hastata L. (Wentworth County )
Verbena urticifolia L. (Essex County )
British Columbia:
Verbena hastata L. (York County)
Vancouver Island:
Verbena bracteata Lag. & Rodr.
UNITED STATES OF AMERICA:
Maine:. '
Verbena hastata L. (Cumberland & Hancock Counties)
New Hampshire:
Verbena hastata L. (Hillsboro County)
Rhode Island:
Verbena hastata L. (Kent County )
Connecticut:
Verbena urticifolia var. lelocarpa Perry & Fernald

(Hartford County )

Delaware:

Verbena simplex Lehm. (Sussex County )
Maryland:

Verbena hastata L. (Wicomico County )
Virginias

Verbena canadensis (L.) Britton (Princess Anne County)
Verbena hastata L. (Roanoke County )
Verbena officinalis L. (Dinwiddie & Henrico Counties )
Verbena urticifolia L. (Albemarle, Rockingham, & York
Counties )
Verbena urticifolia var. leiocarpa Perry & Fernald
(Carroll & Princess Anne Counties )
South Carolina:
Lantana Camara L.
Verbena urticifolia var. leiocarpa Perry & Fernald
(Williamsburg County)
Ohio:
Verbena bracteata Lag. & Rodr. (Scioto County)
Verbena canadensis (L.) Britton (Pike County )
xVerbena hybrida Voss (Ashtabula County)
Verbena simplex Lehm. (Franklin County)
Vitex Negundo var. heterophylla (Franch.) Rehd.
(Preble. County )
382

1947 Moldenke, Geographic Distribution 383

Tllinois: .
Verbena hastata L. (Pope County )
Verbena stricta f. albiflora Wadmond (Winnebago County)
Michigans
Verbena canadensis (L.) Britton (Berrien County)
Wisconsin:
Verbena stricta f. albiflora Wadmond
Minnesota:
Verbena hastata L. (Mower County )
Verbena urticifolia var. leiocarpa Perry & Fernald (Mow-
er County
Missouri:
Verbena canadensis (L.) Britton (Saline County )
Verbena stricta Vent. (Pemiscot County)
Colorado:
Verbena bipinnatifida Nutt. (Boulder County )
Texas:
Phyla cuneifolia (Torr.) Greene (Floyd County )
Phyla incisa Small (Erath County )
Phyla strigulosa var. parvifolia (Moldenke) Moldenke
Cameron County )
Verbena bipinnatifida Nutt. (Castro & Hockley Counties )
Verbena bracteata Lag. & Rodr. (Erath County)
Verbena pumila Rydb. (Hockley & Willacy Counties )
Verbena quadrangulata Heller (Cherokee County )
Verbena tenuisecta Briq. (Shelby County)
Arizona:
Verbena bracteata Lag. & Rodr. (Cochise & Pima Counties )

Verbena Gooddingii var. nepetifolia Tidestr. (Yuma

County

California:
Phyla nodiflora var. reptans (H.B.K.) Moldenke (Lake

County

Phyla nodiflora var. rosea (D. Don) Moldenke (Santa Bar-
bara County )
Verbena Abramsi Moldenke (Lake, Orange, & Trinity

Counties
Verbena lasiostachys var. scabrida Moldenke (Ventura
County )
MEXICO:

Aloysia triphylla (L'Hér.) Britton (Tlaxcala)
Citharexylum affine D. Don (Aguascaliontes )

Lantana Camara L. (Aguascalientes)

Lantana hispida ekg tae

Lippia Pringlei Briq. (A uascalientes

Phyla strigulosa (lant. ‘ Gal.) Moldenke (Coahuila & Oaxaca)

Phyla strigulosa var. parvifolia (Moldenke) Moldenke
(Guanajuato & saleetaie)

Verbena Andrieuxii Schau. (San Luis Potosf)

384 PY ? OL: O07 Vol. 2, noe 9

Verbena perennis var. Johnstoni Moldenke (Zacatecas )
GUATEMALA: 4

Lippia hypoleia Briq. (Alta Verapaz)

Lippia nodiflora var. reptans (H.B.K.) Moldenke (Guatemala)
HONDURAS:

Lippia cardiostegia Benth. (Gracias)

Lippia lucens Standl. (Comayagua)
COSTA RICAs

Citharexylum Standleyi Moldenke (Cartago)

Lippia graveolens H.B.K. (San José)

Lippia Torresii Standl. (Cartago)

Stachytarpheta angustifolia (Mill.) Vahl (Limén)
PAN AMA:

Lippia americana L. (Herrera)

Lippia hemisphaerica Jacq. -- to be deleted
CUBAS

Lippia acuminata C. Wright (Las Villas)

Nashia nipensis (Urb.) Moldenke (Oriente )*
JAMAICA: .

Priva mexicana (L.) Pers.
HISPANIOLA:

Citharexylum Schulzii Urb. & Ekm. (Dominican Republic)
ST. JOHN:

Clerodendrum aculeatum (L.) Schlecht.
ST. KITTS:

Lantana involucrata L.
COLOMEBTAs

Citharexylum Poeppigii Walp. (Méta)

Congea tomentosa Roxb. (Antioquia)

Lippia americana L. (Atldntico, Bolfvar, Cundinamarca, Go-

| ajira, Magdalena, & Santander Norte)

Lippia hemisphaerica Jacq. -- to be deleted

Stachytarpheta canescens H.B.K. (Cundinamarca)

Stachytarpheta straminea Moldenke (£1 Cauca)

Vitex orinocensis var. multiflora (Miq.) Huber (Caqueta)
VENEZUELA!

Aegiphila grandis Moldenke (Mérida)

Aegiphila inteprifolia (Jacq.) Jacks. (Bolfvar)

Aegiphila racemosa Vell. (Mérida)

Ghinia spicata (Aubl.) Moldenke (Bolfvar)

Lantana armata Schau. (Bolfvar)

Lippia americana L. (Zulia)

Lippia hemisphaerica Jacq. -- to be deleted

Phyla betulaefolia (H.B.K.) Greene (Amazonas)

Stachytarpheta elatior var. Jenmani Moldenke (Aragua &
Guaricd)

Stachytarpheta Sprucei Moldenke (Bolfvar)
Vitex compressa Turcz. (Lard)

Vitex Negundo var. heterophylla (Franch.) Rehd. (Aragua)

1947 Moldenke, Geographic Distribution 385

ECUADOR:
Aloysia scorodonioides (H.B.K.) Cham. (Loja)
Cormutie mierocelyeine var. pulverulenta Moldenke (Chimbo-
razo
Lantana scabiosaeflora H.B.K. (Loja)
Lantana Sprucei Hayek (Loja)
Lantana Svensonii Moldenke (Loja)
Lantana trifolia L. (Loja)
Lippia americana L. (Guayaes)
Lippia hemisphaerica Jacq. -- to be deleted
Lippia hyptoides Benth. -- to be deleted
Petrea Andrei Moldenke (Loja)
4 Phyla betulaefolia (H.B.K.) Greene (Loja)
ERUs
Junellia Hayekii Moldenke (Arequipa)
Lantana Moritziana Otto & Dietr. (Madre de Dfos)
Lippia alba (Mill.) N. E. Br. (Lima)
Petrea pubescens Turcz. (Madre de Dfos)
BRAZIL:
Citharexylum Glaziovii Moldenke -- delete Rio de Janeiro
Citharexylum laetum Hiern is the correct form for this
name; delete Maranhao
Citharexylum Ulei var. calvescens Moldenke (Maranhfo)*
Duranta vestita Cham. -- delete Sao Paulo
Lantana canescens H.B.K. -= Sao Paulo
Lantana montevidensis f. albiflora Moldenke (Bahia)*
Lippia asperrima Cham. (Paranda
Lippia lacunosa Mart. & Schau. (Mattogrosso)
Lippia Morongii Kuntze (Mattogrosso)
Lippia nana Schau. (Minas Geraes)

Lippia pedunculosa Hayek (Bahia)

Lippia phryxocalyx Briq. (Nattogrosso)

Lippia pumila Cham. (Parang)

Lippia turnerifolia Cham. (Santa Catharina)
Stachytarpheta azurea Moldenke --to be deleted
Stachytarpheta laevis Moldenke (Rio Grande do Sul) *

Stachytarpheta Loefgreni Moldenke (Ceard)*
Stachytarpheta lythrophylla Schau. (Cearg)

Stachytarpheta Maximiliani Schau. (Minas Geraes & Rio de
Janeiro)

Stachytarpheta mollis Moldenke (Goyaz)*
Stachytarpheta polyura Schau. — delete the
Stachytarpheta sessilis Moldenke (Ceard)*

Stachytarpheta simplex Hayek (Mattogrosso)
Vitex orinocensis var. multiflora (Miq.) Huber (Amazonas)

BOLIVIA:

Citharexylum Poeppigii var. margaritaceum Poepp. & Molden-
ke (La Paz)

Lippia alba (Mill.) N. E. Br. (El Peni)

et

386 Fo BY sT.0 b O0h sk Vol. 2, noe 9

cippia boliviana Rusby (Tarija)
Lippia trachyphy lla Briq. (Cochabamba)
PARAGUAY:
ee imbricata Kuntze
Lippia phryxocalyx Briq. -- delete the "*"
Lippia tegulifera Briq. -- delete the "*"
Lippia tegulifera var. parvifolia Briq. -- to be deleted
Lippia turnerifolia Cham.
Lippia xerophylla B: Briq. ~~ to be deleted
Stachytarpheta polyura Schau.

URUGUAY s
Lippia asperrima var. longipedunculata Moldenke -- to be
deleted
Verbena tenera var. albiflora Kuntze
CHILE: .
Lippia disepala R. A.» Phil. -- to be deleted
ARGENTINA:

Aloysia triphylla (L'Hér.) Britton (Buenos Aires)
Aloysia virgata var. elliptica (Briq.) Moldenke (Jujuy)
~enkara micrantha Briq. (Jujuy)

Lippia alba (Mill.) N. E. Br. (Corrientes)

Lippia asperrima Cham. (Jujuy)

Lippia asperrima var. longipedunculata Moldenke -- to be
deleted

“22 Ekmani Moldenke (Misiones )*
Lippia Crisebachiana Moldenke (Jujuy)
Lippia lupulina Cham. (Migiones)
Lippia Morongii Kuntze (Misiones )
Lippia tegulifera var. ovata Briq. (Misiones)
Lippia turnerifolia Cham. (Misiones)
Verbena dissecta Willd. (Catamarca)
Verbena storeoclada Eriq. (Santiago del Estero)
ALGERTA:
Chascanum marrubiifolium Fenzl
UGANDA PROTECTORATE:
Clerodendrum volubile P. Beauv.
KENYA:
.Premna Holstii Glirke
Vitex Volkensii Gtirke
INDIAs
Clerodendrum Phlomidis L. f. (Bengal)
Holmskioldia s sanguinea Retz. (Madras)
CHINA:
Clerodendrum Leveillei Fedde (Ytinnan)*
BELEP ISLANDS: =
Oxera oblongifolia var. artensis (Dubard) Moldenke (Art)*
NEW CALEDONIA:
Oxera oblongifolia var. sinuata (Dubard) Moldenke*

Oxera pulchella var. Dep lancheana Dubard*

1947 Moldenke, Geographic Distribution 387

’ Oxera pulchella var. microcalyx Dubard*
Oxera subverticillata var. candelabrum Beauvie .*

AUSTRALIA:
Chloanthes Stoechadis R. Br. —= delete the "*"
NEW ZEALAND:
Chloanthes Stoechadis R. Br.
CULTIVATED:
Bouchea fluminensis (Vell.) Moldenke (Brazil)
Callicarpa dichotoma (Lour.) K. Koch (Florida)
Caryopteris incana var. nana Moldenke (Cregon)*
Caryopteris incana var. superba (Dreer) Bobbink & Atkins
is the correct form for this name; known from New Jer=
sey, New York, & Pennsylvania.
Citharexylum ellipticum Sessé & Moc. (California)
Citharexylum Glaziovii Moldenke (Brazil)
Clerodendrum speciosissimum Van Geert (Massachusetts )
Clerodendrum trichotomum var. tomentosum Moldenke (New
York
Clerodendrum ugandense Prain (California)
Clerodendrum viscosum Vent. (Cuba)
Lippia alba (Mill.) N. E. Br. (Hispaniola)
Lippia Pringlei Briq. (Germany )
Monochilus gloxinifolius Fisch. & Mey. (Russia)
Verbena platensis Spreng. (New York)
Vitex parviflora A. L. Juss. (Maryland)

ADDITIONAL NOTES ON THE GENUS AEGIPHILA. VIII

Harold N. Moldenke

Many hundreds of additional specimens of this genus have
been examined by me since the publication of the seventh
installment of these notes in 1941, and much additional in-
formation has come to light. The material of the group from
nine additional herbaria has been studied. The abbrevia-
tions employed to designate these herbaria hereinafter are
as follows: Bt = Butler University, Indianapolis; Om = Car-
negie Museum, Pittsburgh; Du = Dudley Herbarium, Stanford
University; Io = [Lowa State College, Ames; Me = Instituto
de Biologia, Universidad Nacional de México; Si = Instituto
Darwinion, San Isidro; Ug = Museo de Historia Natural, Mon-
tevideo; Ur = University of Illinois, Urbana; and Vt = Uni-
versity of Vermont, Burlington. All other abbreviations
herein employed have been explained in previous install-
ments of this series or in my original monograph.

388 PHYTOLOGIA Vol. 2, no. 9

References: Reichenb., Conspect. Reg. Veg. 1: 117. 1828;
Steud., Nom. Bot., ed. 2, 1: 29. 1840; Le Cointe, A Amazonia
Brasileira III, Arvores e Plantas Uteis 127. 1934; Leén,
Revista de la Sociedad Geogrdfica de Cuba 2: 44. 1942; Samp-
aio & Peckolt, Arquiv. Mus. Nac. Rio Jan. 37: 334. 1943;Le
Cointe, O Estado do Para 232. 1945; Wynne, Tax. Index 8:
entry 226. 1945; Reko, Bol. Soc. Bot. Mex. 4: 35. 1946;
Irm&io Augusto, Flora do Rio Grande do Sul 230. 1946.

Wynne, in the reference cited above, misspells this gen-
eric name "Aegiphala"; Steudel, in the reference cited
above, spells it "Aegiphyla", and this spelling is recorded
in synonymy in the "Index Kewensis". Reichenbach, in the
reference cited above, writes it "Aegiphila L." and gives
it as a valid genus in the Labiatae, section Verbeneae; he
also gives Manabea Aubl. as a valid genus in the same sec-
tion. Le Cointe in the 1934 reference cited above records
the conmon name "cipo pitomba"” for an unidentified species
of this genus from Brazil, and in his 1945 work (cited a-
bove) he records the common name "uruerana" for an unidenti-
fied species of this genus from Pard. Sampaio and Peckolt,
in their paper entitled "A nomenclatura das espécies na
‘Flora Fluminensis' de Conceig&o Veloso e sua correspondén-
cia atual" (cited above) state that A. inflexa Vell., Ae

stipulata Vell., and A. umbellata Vell. are actual ly rubia-
ceous. In this they follow Schauer. In my booklet "An alpha-
betic list of invalid and incorrect scientific names propos-
ed in the Verbenaceae and Avicenniaceae", pages 2 & 3
(1942), I reduced A. inflexa to Paychotria, subgenus Mapour-
ea, and A. umbellata to Feramea.

AEGIPHILA ACULEIFERA Moldenke

Steere describes the flowers of this species as pale yel-
low, blooming in January; Skutch calls it a small tree. It
grows on mountainsides in the cloud forest in Colombia, at
an altitude of 2500 m. The Steere collection does not exhib-
it the glandular apiculations normally found on the leaf-
blades of this species. It has been identified by Killip as
"Aegiphila aff. A. glandulifera".

Additional citations: COSTA RICA: Alajuela: Skutch 3255
(S). COLOMBIA: Métas Steere 7091 (W--1833960).

AEGIPHTLA ALBA Moldenke

Holdridge, Teesdale, Myer, Little, Horn, & Marrero, For-
ests West. & Cent. Ecuador 46 (1947) record the common name
"margarita". Little also records the names "lulu", "margar-
ita", "masamoro", and "savaluca de montana" on herbarium
labels. He describes the plant as a tree 26 to 66 feet tall,
with a trunk diameter of 6 to 12 inches at breast height,
the bark light-gray, smooth, with slight cracks and with li-

1947 Moldenke, Notes on Aegiphila 389

chen patches. He states that the white flowers are borne in
clusters along the stem or in “axillary clusters", the flow-
er buds green. On his no, 6439 he states that the flowers
themselves are greenish. It has been collected in flower in
April, May, and June; immature fruit was collected in April
and mature fruit in May. He describes the species as common
and dominant in wet tropical forests, common in cut-over
woods, and very common in cacao plantations. His no. 6439
was found at an altitude of only 150 feet, while Steyermark’
plant was growing at an elevation of 850 meters! This latter
collector describes the plant as a shrub 10 feet tall, with
subcoriaceous erect leaves, inhabiting steep slopes in rich
rain-forest jungles.

Additional citations: ECUADOR: Azuay: Steyermark 52756
(F--1205652, N). #1 Oro: BE. L. Little 6675 [U. 9. Forest
Serv. 98564] (N). Eemeraldas: E. L. Little 6331 [U. S. For-
est Serv. 98292] (W--1877593). Los Rfos: EB. L. Little 6439
[U. S. Forest Serv. 98266] (N). Pichincha: E. L. Little 6154
[U. 8S. Forest Serv. 96814] (W-~1877632).

AEGIPHILA AMAZONICA Moldenke

The species has been collected by Ducke in non-inundated
forests on terra firma, blooming in January.

Additional citations: BRAZIL: Amazonas: Ducke 864 (N, W--

AEGIPHILA ANOMALA Pittier

Austin Smith states that this species is "usually a shrub"
although it may becove a tree 15 m. tall, the base of the
trunk to 40 em. in diameter, the bark dark-brown, suberose,
and roughened, the leaves membranous, nearly flaccid, and
light-green, the buds "buffy green", and the white flowers
produced in “nearly globose" clusters. He found it in heavy
Clay-loam soil in nearly open exposures in hilltop woodlands
at the upper limit of the tropical zone, blooming in July.
It has been erroneously distributed as A. Valerii Standl.

Additional citations: COSTA RICA: Alajuela: Brenes 6552
[518] (N), 15661 [189] (N); A. Smith N.Y.138 (N). SS

AEGIPHILA BOGOTENSIS (Spreng.) Moldenke

The Dawe 192 previously cited as from "Department unde-
termined", Colombia, is actually from Cundinamarca. Steyer-
mark records the common name "yuco blanco" for this species
and describes the plant as a tree 20 feet tall, with leaves
that are dark-green above and pale-green beneath, the stem,
petiole, calyx, and under surface of the midrib tawny, the
corolla-tube pale-green and waxy, the lobes creamy-white. He
describes the wood as "good", and states that the tree
"grows very tall". in moist cloud forests on south- and

390 PHY.T.OL 06.2 A ) Vol. 2, no. 9

southwest-facing slopes, at altitudes of 2530--2575 m.,
flowering in February. Killip found it at altitudes of 2900
to 3200 m., blooming in August, with "creamy or waxy-white"
corollas. Cuatrecasas found it at altitudes of 2700 to 3100
m., blooming in January and April, fruiting in April and
September. He describes it as a large tree in woods, with
white or ochraceous=-white corollas. Daniel records the com
mon name "saca-ojo", and says the tree is 5--7 m. tall, the
fruit rounded and green in July. Garcia y Barriga records
the common name "queso fresco", and says the tree grows to
10 m. tall, at altitudes of 1900--2100 m., fruiting in Janu-
ary. Tomas found it at 3000 m., flowering in July, and Dry-
ander at 2900 m., fruiting in August. It has been mistaken
for a species of Brunfelsia and thus distributed.

Additional citations: COLOMBIA: Antioquia: Daniel 528
(N); Tomds 1512 (N). Caldas: Dryander 2809 (W--1879534);
Killip 9811 (N); Tomdés 2415 (W--1857909). Cundinamarca: Cu-
atrecasas & Jaramillo 12014 (w--1850860); H. Garcia y Barri-
ga 11032 (W--1852224). El Valle: Cuatrecasas 20812 (N). Nar-
ifio: Cuatrecasas 11963 (W--1799876). VENEZUELA: Lard: Stey-

ermark 55265 (N

AEGIPHILA BRACHIATA Vell.

References: Sampaio & Peckolt, Arquiv. Mus. Nace Rio Jan.
37: 334. 1943; Lombardo, Flora Arb. Arbores. Urug. 18 &
201. 1946; Irm&o Augusto, Flora do Rio Grande do Sul 231 &
236. 1946.

Irm&o Augusto on page 236 of the work cited above spells
the name "Aegiphila brachyata Vell.", which he gives as a
synonym under A. triantha Schau. Lombardo states that the
plant is a shrub 2=--3 m. tall, sparsely branched, found in
the departments of Tacuarembé, Treinta, and Tres of Uruguay.
He also states that Arechavaleta published a photograph of a
flowering branch of this plant in An. Mus. Nac. Montevid. 4:
62, pl. 1 (1902), along with a description.

The Curran specimen cited below has very small flowers
for this species, and is thus anomalous. The Sellow specimen
cited below is perhaps an isotype of A. triantha.

Additional citations: BRAZIL: Rio de Janeiro: Curran 636
(N). State undetermined: Sellow s.n. [Brasilia] (Vt).

AEGIPHILA CHRYSANTHA Hayek

The Poeppig 2314 collection is also the type collection
of A. lutea Poepp.

Additional citations: PERU: Loreto: Poeppig 2314 [Mac-
bride photos 343135] (Kr--photo of logotype).

AEGIPEILA CORDATA Poepp.
Additional citations: PERU: Loreto: Poeppig 2158 [Mac-

1947 Moldenke, Notes on Aegiphila 391
bride photos 34312] (Kr--photo of type).

AEGIPHILA CORDATA var. COLOMEIANA Moldenke

Cuatrecasas describes this plant as a vine with "ramas
sepia verdoso claro", bright-green leaves and calyx, and
yellowish-white corollas, inhabiting woods at elevations of
5 to 20 m., blossoming in February. It has been confused
with A. racemosa Vell.

Additional citations: COLOMBIA: El Valle: Cuatrecasas

13993 (N).

AEGIPHILA CORDIFOLIA (Rufz & Pav.) Moldenke

The type collection of this species, made by Rufz and
Pavon at "Panatahua"” -- a locality which hitherto could not
be accurately located as to department -- actually came
from Hudnuco, Peru, and should be so cited.

AEGIPHILA COSTARICENSIS Moldenke
Additional citations: MEXICO: Chiapas: Matuda 2101 (Dp--
28971). COSTA RICA: Alajuela: A. Smith 1818 (N).

AEGIPHILA CUATRECASASI Moldenke

Cuatrecasas describes this species as a small or large
tree, to 10 m. tall, with soft wood, the leaves subcoriace-
ous, flexible, clear-green or gray-green, or "hoja herbacea
gruesa", the fruit produced in glomerules, fleshy, yellow-
ish-green, 15--18 mm. long, with a firm epicarp, 4-seeded.
He found the tree at altitudes of from 5 to 1750 m., fruit-
ing in April.

Additional citations: COLOMBIA: El Valle: Ouatrecasas

17075 (N), 21007 (N).

AEGIPHILA DEPPEANA Steud.

The Liebmann 11957, previously cited as from "State un-
determined", Mexico, is probably actually from Puebla, ac-
cording to a letter received by me from M. Martfnez, dated
May 5, 1945. The Macbride photograph cited below is a
photograph of the type specimen of A. Eerteriana Schau. The
Dugand & Jaramillo collection cited below was made at an
altitude of 200--250 m., where the species was blooming in
January. Ferris 6259 exhibits leaves which are membranous in
texture and are glabrate on both surfaces -- it obviously
represents the A. pacifica of Greenman, which may, after all,
turn out to be a valid species or, at least, variety.

Additional citations: MEXICO: Oaxaca: Martfnez-Calderén
418 (Me). Tres Marias Islands (Maria Madre): Ferris 625
(Du--145788). COLOMBIA: Atléntico: Dugand & Jaramillo 4056
(N, W--1900073). Magdalena: Bertero s.n. [Herb. DeCandolle
850; Macbride photos 33932] (Kr—-photo); H. H. Smith 881 (Cm,

392 PHYTOLOGIA Vol. 2, noe 9
Vt), 1864 (Cm, Vt).

AEGIPHILA ELATA Sw.

References: Abh. Akad. Berl. 215. 1831; Contrib. Univ.
Mich. Herb. 8: 60. 1942; Roig y Mesa, Plant. Med. Cuba 411 &
770. 1945.

The Hahn s.n. from "Potrero", Mexico, cited previously as
from "State undetermined", is probably from Veracruz, accor-
ding to a letter from my friend, M. Martinez, dated May 5,
1945, and should be so cited. The synonym "Aegiphila corni-
folia Kunth" is recorded by the "Index Kewensis". The spec-
ies is described by Matuda as a woody vine in second growth,
savannas, end advanced forests of Tabasco. Gentle says it is
a woody vine, with yellow flowers and fruit, inhabiting sec-
ondary forests on river banks in Eritish Honduras, where he
found it in flower and fruit in August. Roig y Mesa, in the
work cited above, records the common name “guairo santo de
costa". The British Guiana Forest Department specimen cited
below bears the inscription "5 cm. diam. gray-brown papery-
barked rope from vrown of tree; leaves thinly fleshy; fls.
in terminal compound inflorescences; calyx pale-green, glab-
rous, 3-lobed; corolla tubular, palest cream, lobes erect-
spreading; stamens white."

Additional citations: FLORIDA: Dade Co.: Buswell s.n.
[July 25, 1935] (Bu). CUBA: Las Villas: R. A. Howard 6441
(N). Oriente: Alain & Crisogono 307 (Ha); Hioram 6611 (Ha),
6710 (Ha, Ha, N, N)3 Leén 10113 Ha), 18185 (H re JAMAICA:
Maxon & Killip 747 (Ur). MEXICO: Tabasco: Matuda 3406 (Du--
299395). BRITISH HONDURAS: Gentle 3569 (N), 3578 (N), 3579
(N). HONDURAS: Atléntida: Yuncker, Koepper, & Wagner 8
(3). COLOMBIA: Bolfvar: Moritz 1478 (Wey. Cundinamarcas H.
Garcia y Barriga 12129 (W--1900406). BRITISH GUIANA: Herb.
Forest Dept. Br. Guian. 4027 [F.1291] (N).

AEGIPHILA ELEGANS Moldenke
Additional citations: BRAZIL: Amazonas: Krukoff 8701 (Ss).

AEGIPHILA FALCATA Donn. Sm.

Wedel describes this species as a tree 15 feet tall, with
yellow flowers, blooming in September.

ae citations: PANAMA: Bocas del Toro: Wedel 683
(E).

AEGIPHILA I‘ARINOSA Moldenke
See original description in Phytologia 2: 306--307. 1947.
Specimens examined: COLOMBIA: El Valle: Cuatrecasas 21689
(N--type).

AEGIPHILA FENDLERI Moldenke

1947 Moldenke, Notes on Aegiphila 393

Steyermark describes this species as a woody vine, with
membranous leaves which are deep grass-green above and buff-
green beneath, and the calyx and bracts pale buff-green. He
found it on seaward-facing north mountain slopes, at alti-
tudes of 1830 to 2130 m., blooming in June.

Additional citations: VENEZUELA: Federal District: Stey-

ermark 56959_(N).

AEGIPHILA FERRUGINEA Hayek & Spruce

Diels in his Contrib. Conocim. Veg. Flora Ecuador (trans.
R. Espinosa] 268 (1938) cites Diels 783, from Carchi, as
this species. Wiggins describes it as a shrub to 6 m. tall.
He collected it at an altitude of 9700 feet, blossoming in
August.

Additional citations: ECUADOR: Carchi: Wiggins 10685 (Du-
311630). Pichincha: Spruce 5473 [Macbride photos 34311] (Kr-
photo of type).

AEGIPHILA FILIPES Mart. & Schau.

Smith collected this species at an altitude of 5000 feet
in Magdalena, blossoming in February.

Additional citations: COLOMBIA: Magdalena: H. H. Smith
1831 (Cm, Vt). BRAZIL: Amazonas: Krukoff 8041 (s), 8042 (3).

AEGIPHILA FLORIBUNDA Moritz & Moldenke
Additional citations: VENEZUELA: Aragua: Moritz 1765
[Macbride photos 34310] (Kr-=photo).

AEGIPHILA FLUMINENSIS Vell.

References: Sampaio & Peckolt, Arquiv. Mus. Nac. Rio de
Jan. 37: 334. 1943.

Additional citations: BRAZIL: Rio de Janeiro: Riedel &
Lusehnath 323 (N).

AEGIPHILA GLANDULIFERA Moldenke

Fruiting-calyxes and fruit of this species have now been
seen, so the following information can be appended to the
species description: fruiting-calyx cupuliform, 4--4.5 mm.
long, about 9 mm. wide, minutely pulverulent-puberulent, its
rim truncate, entire or slightly erose; fruit ochraceous,
later black, otlong-elliptic, 8--10 mm. long, 6--9 mm. wide,
glabrous.

Diele in Contrib. Conocim. Veg. Flora Ecuador [translated
by Espinosa] 268 (1938) cites Diels 94 from Tunguragua, and
describes the corolla as greenish-yellow, the anthers clear-
yellow. Haught says it is a small tree, 5m. tall, very
sickening-fetid, with white rather showy flowers, blooming
in November at an altitude of 100 m. He believes that his
no. 2061 is a different species from his no. 1629, but I re-

394 PRY“? O"L OGeh A Vol. 2, no. 9

gard both collections as representing the same species. Klug |
reports the common name "chirapa sacha", and describes the
plant as a shrub 2 m. tall, with cream-colored flowers in A-
pril, gorwing in forests at an altitude of 220 m. Krukoff
found it in old clearings, a shrub 12 feet tall, with a stem
diameter of 2 inches. Cuatrecasas says it is a small tree to
8 m. tall, with thin=-herbaceous gray-green leaves, green
calyx, and yellow corollas, growing at altitudes of 5--50 m.
He found it in flower and fruit in February and March. Ginz-
berger describes it as a shrub with yellow-green flowers in
August. It has been confused with A. filipes Mart. & Schau.

Additional citations: COLOMBIA: El Valle: Cuatrecasas
10958 (N, N). Santander Sur: Haught 2061 ( F--929606, N, W--
1742327 ). PERU: Loreto: Klu 5016 (F--685001). BRAZIL: Ama-
(ee Ee 8290 (F--929898, N). Pard: Ginzberger 902

F--954891 ).

AEGIPHILA GLANDULIFERA var. PYRAMIDATA L. C. Riche & Moldenke
This plant somewhat resembles. A. laevis (Aubl.) Gmel.,
but may be distinguished by its more elongated terminal pan-
icles, its thin-membranous leaf-bliades, which are densely
marked with glandular disks along the midrib beneath, and
its densely strigillose or short-strigose branchlets, pe-

duncles, rachis, pedicels, and petioles.

AEGIPHILA GLEASONII Moldenke

This species differs from all other Guianan species of
the genus in having abbreviated, sessile, glomerate, about
6-flowered cymules in the axils of the extremely large
leaves. The leaf-blades are to 42 cm. long and 16 cm. wide.

AEGIPHILA GLOMERATA Benth.

Little reports the common names "palo flojo", "palo de
cereuchara", and "arritagua" for this plant. He describes it
as a small tree, 16--40 feet tall, with a trunk diameter of
4-6 inches at breast height, gray or light-gray, rough,
fissured, shredding bark, the fissures about 3 mm. deep and
l cm. wide. opposite pubescent leaves, axillary flower clus-
ters, and yellow corollas. He found it in dry forests, flow-
ering and fruiting in June. His no. 6693, cited below, rep-
resents the first fruiting collection known.

Additional citations: ECUADOR: El Oro: 2. L. Little 6693
([U. 9. Forest Serv. 98639] (W--1878649), 6696 [U. S. Forest
Serv. 98613] (W--1878642).

AEGIPHILA GLORIOSA Moldenke
Additional citations: BRAZIL: Bahia: Blanchet 1998 (F--
976379) «

1947 Moldenke, Notes on Aegiphila 395

AEGIPHILA GRANDIS Moldenke

Cuatrecasas describes this species as a small tree, 5 m.
tall. He found it fruiting in March at an altitude of 200 m.
Triana found it flowering in August at an altitude of about
1800 m.

Additional citations: COLOMBIA: Caquetd: Cuatrecasas 8700
(W--1795403 ). Cundinamarca: Mutis 4554 (F--712945, N--photo,
Z--photo); Triana 2080 [Macbride photos 28379] ( F--830241--
photo, Kr=-photo), 3712 [2] (Jc). Tolima: Goudot s.n. [Port-
achuelo, Quindiu] (F--642172--photo of type). VENEZUELA:
Mérida: Steyermark 56458 (F--1221913, N).

AEGIPHILA GRAVEOLENS Mart. & Schau.

This binomial is sometimes inaccurated credited to
"Schau. & Mart." or "Mart, & Schum."

Additional citations: BRAZIL: S&o Paulo: A. Gehrt 30081
( F~-895955), gn. [Herb. Inst. Biol. 3S. Paulo 30081] (F--
895767); Lund 796 [Macbride photos 7880] (F--645500--photo
of type, Kr==photo of type, Ne-photo of type).

AEGIPHILA GUIANENSIS Moldenke

This species may be distinguished quickly from the simi-
lar A. integrifolia (Jacq.) Jacks. in its branches being
densely short-villous with yellowish pubescence and its
young leaf-blades being densely lanate-tomentose beneath.
Killip and Cuatrecasas describe it as a small treem with the
young inflorescence greenish. They found it in dense tidal
forests. Pittier found it at 90 m. elevation, blooming in
June, while Triana found it flowering in January at an ele-
vation of 300 m.

Additional citationss COLOMBIA: Chocé: Killip & Cuatre-
casas 1 (N). Cundinamarca: Triana 2084 [Macbride photos
28380] (F--830245--photo, Kr=-photo). Méta: Triana 3713 [4]
(Jc). VENEZUELA: Bolfvar: H. Pittier 13401 (Kr). BRITISH
GUIANA: M. R. Schomburgk 404, in part (F--642175--photo of

type).

AEGIPHILA HASSLERI Brigq.

This plant is illustrated by Arechavaleta, An. Mus. Nac.
Montevideo 4: 62, pl. 1 (1902), under the name of A. trian-
tha. This is the picture to which Lombardo refers in his
Flora Arb. Arbores. Urug. 185 & 201 (1946) -- see under A.
brachista in these present notes. The specimen on which the
picture was based was collected in Uruguay by Cornelio B.
Cantera. Jérgensen describes A. Hassleri as a small tree,
2--4 m. tall, with sulphur-yellow corolla and yellow fruit,
very common in hedges and thickets, in flower and fruit in
September. Schréder calls it a "large tree".

Additional citations: PARAGUAY: Fiebrig 260 (F--642177--

396 PAY T.0 LL OG 2 A Vol. 2, noe 9

photo); Hassler 3193 [Macbride photos 24613] (F--772047--

photo of cotype, Kr--photo of cotype); Jérgensen 3662 (Du--
185439). URUGUAY: Schréder s.n. (Herb. Osten 16059] (Ug);

Arechavaleta s.n. [Herb. Osten 13002] (Ug). ARGENTINA: Misi-~
ones: D. Rodriguez 566 (Herb. Inst. Miguel Lillo 52532] (N).

AEGIPHILA HAUGHTII Moldenke

Schunke describes this species as a tree, 8m. tall, with
a stem 28 cm. in circumference and white flowers, blooming
in March. He collected it "on rising ground."

Additional citations: ECUADOR: Guayas: Haught 2904 (N--
fragment of type, N=-photo of type, W--1/707582--type, Z--
photo of type). PERU: Loreto: Schunke 338 (W--1459225).

AEGIPHILA. HERZOGII Moldenke

Additional citations: BOLIVIA: Santa Cruz: Herzog 1369
[Macbride photos 22381] (F--642176--photo of isotype, Fe=
830239=-photo of isotype, Kr-—photo of isotype).

AEGIPHILA HIRSUTA var. COLOMBIANA Moldenke

See the original description of this variety in Castanea
10: 44 (1945). The type collection was made in wet woods
along the Rfo San Miguel, at an altitude of 350 m., on the
Ecuador=Colombia boundary.

Specimens examined: COLOMBIA: Putumayo: Cuatrecasas 11032
(W--1798861--type).

AEGIPHILA HIRSUTISSIMA Moldenke
References: Pittier, Supl. Plant. Usual. Venez. 54. 1939.

AEGIPHILA HOEHNEI var. PUYENSIS Moldenke

See the original description of this variety in Phytolo-
gie 2: 214 (1947). It is a woody vine, about 3 m. long, with
off-white flowers, blooming in May at an altitude of 3000
feet.

Specimens examined: ECUADOR: Orientes Steere & Camp 8283
( F--1163157--type, N--photo of type, Si--photo of type, Z--
photo of type). .

AEGIPHILA HOEHNEI var. SPECTABILIS Moldenke

See the original description of this variety in Castanea
10: 44-45 (1945). The collectors describe it as a woody
vine, with white green-tinged corollas, growing in forests
at the edge of mangrove belt, blooming in June.

Specimens oxamined: COLOMBIA: El Valle: Killip & Cuatre-

casas 38978 (N-~type).

AEGIPHILA INSIGNIS Moldenke
Additional citations: PERU: Ancachs: Rufz & Pavon 3/94

1947 Moldenke, Notes on Aegiphila DST

(F--850850), this fragment, collected in 1797, may be part
of the type collection.

AEGIPHILA INTEGRIFOLIA (Jacq.) Jacks.

References: Jacq., Hist. Stirp. Amer. 15, pl. 175, fig.
7. 1780; Jacks., Ind. Kew. 1: 386. 1895; Junell, Symb. Bot.
Upsal. 4: 82 & 83. 1934; Pittier, Supl. Plant. Usual. Venez.
54. 1939; Lanjouw & Uitten, Rec. Trav. Bot. Néerl. 37: 152.
1940; Irm&o Augusto, Flora do Rio Grande do Sul 231 & 236.
1946; Veloso, Mem. Inst. Oswaldo Cruz 44: 267, 282, 292, &
335. 1946.

Lanjouw and Uitten, in the reference cited above, tell of
discovering the actual type specimen of Manabea arborescens
(and therefore of the genus Manabea) of Aublet in Herb. Den-
aiff 3: 109 -- e flowering branch closely resembling Aublet's
plate. Junell, in the reference cited above, discusses the
gynoecium morphology of the species and gives an illustrat-
ion of it in his Fig. 133. Jackson, in the reference cited

above, records this binomial as "Aegiphila integrifolia
Jacq.", while Irm&o Augusto on page 236 of his work cited

above, gives "Aegiphy lla discolor Willd." and "“Aegiphylla
integrifolia Jacq." as synonyms. Veloso, on page 355 of the
work cited above, records the species as "Aegiphila
arboreuceus". He states that the species is a tree about 3
m. tall, the trunk 10 cm. in circumference, with latex,
growing in more or less wet places in climax and subclimax
associations of Lecythis-Sickingia, Virola-Tapirira, and
Tapirira~Simaruba. He reports that the seeds are used by the
natives.

The specimens collected by Ruiz & Pavon at "Pantahua" and
"Chichao", Peru, and recorded in Brittonia 1: 339 (1934) as
from an undetermined department of Peru, are actually from
Hudnuco and should be so cited. The Cuatrecasas 8873 col-
lection exhibits especially small and silky leaves, even
though the plant is in full anthesis. It may represent an as
yet undescribed variety or species, although the inflores-
cence is very typical of A. integrifolia. It is described as
having white flowers, blooming in March. Williams 2052 also
does not seem to be typical material and resembles some of
Ducke's material which represents another species. Lawrence
548 has very small flowers and very obovate leaves, and on
this account is not typical. Schunke 343 has the immature
leaves golden-velutinous, and is described as a bush 4 m.
tall, with a stem 10 cm. in circumference and white flowers,
blooming in March, at altitudes of 100--125 m.

McCarroll describes A. integrifolia as a "large tree", 9
m. tall, its white flowers with a "lively delicate odor",
growing at 1550 m. elevation. Metcalf says it is a bush 2-~
2.5m. tall, with "dirty-brown" fruit in May and June, in

398 PHYTOLOGIA Vol. 25 noe 9

dense growth on moist shaded banks in regions with much fog
and rain, at an altitude of 1900 m. Klug describes it as a
tree 5m. tall, with white flowers, at altitudes of 1200 to
1600 m. Williams records the common name "tabaquillo" and
describes it as a shrub or tree, 3—--12 m. tall, with a
rounded crown, trunk 30 cm. in diameter, straight and with-
out branches for the first 4 m., the outer bark gray and
rough, the inner bark rather thick and clear-chestnut or
dark=-red in color, the wood light in color, and the flowers
white or whitish. He also notes that "la labura y el dira-
men son de color rosado y susceptible a los ataques de in-
sectos", He found it in rocky places and in high secondary
woods on terra firma, at altitudes of 120-125 m., blooming
in August and September. Cuatrecasas describes it as a small
tree, 6m. tall, the stem 10 cm. in diameter, branches
white-tomentose, the leaves herbaceous and clear— or gray-
green on the upper surface, pale or clear-green on the under
surface, the calyx greenish-rhite or pale yellowish-green,
and the corolla white, blooming in May at altitudes of 5 to
80 m. The Britton Herbarium specimen of his no. 17491] in-
cludes a large strip of the bark.

Additional citations: VENEZUELA: Amazonas: Ll. Williams
13174 (Ve), 15854 (W--1876460), 16005 (W--1876541). Bolfvar: :
Steyermark 57675 (F--1221911, N); Tamayo 2982 (W--1906645). |
COLOMBIA: Boyacdé: Lawrence 548 ( F--708632)..Caquetd: Cuatre-
casas 8 (W--1795006). El Valle: Cuatrecasas 17369 (N, N),
17491 (N). BRITISH GUIANA: Maguire & Fanshawe 23060 (N),
23476 (N). PERU: Hudnuco: Rus & Pavon 12 ( F—~7 12587).
Loreto: J. M. Schunke 38. (F--997587, N); Ll. Williams 2052
(F--613150), 2795 (F=--608731). San Mart{n: Klug L668 (F-=
736254). Punot McCarroll 94 (N); R. D. Metcalf 30 (W-=
1876045). BOLIVIA: La Paz: M. Bang 564 (Om, Io—32313).

es es pe ee ee

AEGIPHILA INTERMEDIA Moldenke

The description given in Phytologia ls 397--398 (1940)
under A. salticola Moldenke applies to the Ducke s.n. [Herb.
Rio de Jan. 25593 | collection there cited, but this collec-
tion seems better placed under A. intermedia. It is, in
fact, very possible that A. salticola should be reduced to
synonymy under A. intermedia. Williams records the common
name "tabaquillo”, and collected it in flower in May. Ducke
describes it as a small tree, with white flowers, blooming
in January, growing in secondary non-inundated forests.

Additional citations: VENEZUELA: Amazonas: Ll. Williams
13174, in part (W--1800206). BRAZIL: Amazonas: Ducke 136

F—-901732). Maranh&io: Herb. Gen. Mus. Para. 2270 [Macbride

photos 28382] (F--830240—-photo of isotype, Kr——photo of
isotype).

1947 Moldenke, Notes on Aegiphila 399

AEGIPHILA LAETA H-B.K.

Haught describes this plant as a slender shrub, 2 m.
tall, tending toward a tree-like habit, with very inconspic-
uous flowers, blooming in August, growing in forests at an
altitude of 250 m. Daniel describes the corollas as cream
colored and the fruit red, each with 3 or 4 seeds. He fowd
it in flower and fruit in July.

Additional citations: COLOMBIA: Antioquia: Daniel 2047
(N). Goajira: Haught 4316 (N, W--1709284). Magdalena: Bon-
pland 1664 (F--976536--fragment of type); Daniel 2047, in
part (F--1007465); H. H. Smith 330 (Ca--584595, 38).

AEGIPHILA LAEVIS (Aubl.) Gmel.

An additional synonym is Aegiphila longifolia Willd. ex
Moldenke, Suppl. List Invalid Names 1, in syn. 1941. Lan-
jouw & Uitten in Rec. Trav. Bot. Néerl. 37: 152 (1940) tell
of discovering the type specimen of Aublet's Manabea laevis
in Herb. Denaiffe 3: 109 -- a flowering branch closely re-
sembling Aublet's plate. The name written on the Aublet
photograph cited below is, curiously enough, "“Aegiphila lae-
vis (Jacq-) Gmel." Cuatrecasas describes the species as a
large vine or small tree, with subcoriaceous, rather thick
and flexible leaves, which are dark-green and slightly shiny
on the upper surface and clear~green on the lower surface,
the calyx pale-green or greenish-yellow, the corolla yellow
ish or "clear-green", blooming in February, March, and May,
and fruiting in May. He found it at altitudes of 1--50 m.

Additional citations: COLOMBIA: El Valle: Cuatrecasas
14302 (N), 15946 (W--1853869), 17705 (N). SURINAM: Kappler
5435 (F--588407--fregment); Maguire & Stahel 22782 (N).
FRENCH GUIANA: Aublet s.n. (F--642180--photo of isotype).

AEGIPHILA LANATA Moldenke

The mis-epelling "Aegiphila lanta" is recorded in Molden-
ke, Suppl. List Invalid Names l, in syn. (1941).

Additional citations: BRAZIL: Goyazs Glaziou 21917 [Mac-
bride photos 28383] (F--830246--photo of isotype, Kr-~photo
of isotype).

AEGIPHILA LANCEOLATA Moldenke

The collection number is written "D.1642" on the specimen
cited below.

Additional citations: BRAZIL: State undetermined: J. E.
Pohl 1642 (F--869797--fragment ).

AEGIPHILA LAXICUPULIS Moldenke

The "“Aegiphila martinicensis L." listed by Calderén &
Standley, Lista Preliminar de Plantas de £1 Salvador, Flora
Salvadorefia, ed. 2, 235 (1941) is actually A. laxicupulis,

400 PHYTOLOGIA Vol. 2, noe 9

and the common name "palo de zope" applies to this species.

The mis-spellings "A. lexicaulis" and "A. laxicupula" are

recorded -~ the former in my Suppl. List Invalid Names 1, in

syn. (1941) and the latter in Alph. List Invalid Names 2, in

syn. (1942). Skutch describes the species as a small tree 15

to 30 feet tall, the trunk 7 inches in diameter at breast

height, with cream-colored corollas, blooming in September

in second=growth thickets and woods, at an altitude of 2600

feet. He describes the species as "dioecious" and says his

no. 1280 represents the staminate and his no. 13510 the pis-

tillate form. .
Additional citations: GUATEMALA: Quezaltenango: Skutch :

$280 (F--933625), 1310 (F-—933704). NICARAGUA: Matagalpas

Rothschuh 628 ( F—~-642187--photo). :

AEGIPHILA LAXIFLORA Benth.

This species somewhat resembles A. membranacea Turcz.,
but may be distinguished by its smaller leaves (3--10.5 cm.
long, 1.5--5 cm. wide), its very slender or filiform pedun-
cles, sympodia, and inflorescence—branches, its very light-
gray or almost white stems and larger branches, and its min-
utely puberulent or glabrate branchlets. Steyermark de-
scribes it as a shrub, 15 feet tall, with membranous erect
leaves, which are deep-green above and dull paler-green be-
neath, calyx greenish, and corolla greenish-yellow, blooming
in April at altitudes of 700 to 600 m.

Additional citations: VENEZUELA: Bolfvar: Steyermark
57719 (F--1221900, N). Monagas: Steyermark 62242 (F—
1205704). BRITISH GUIANA: M. R. Schomburgk 772 [Macbride
photos 28384] (F--830242--photo of isotype, F--869788-—~frag-
ment of isotype, Kr--photo of isotype).

AEGIPHILA LEHMANNII Moldenke

Lawrance describes this species as a tree 20 feet tall,
the trunk 6--7 inches in diameter, with white to cream-col-
ored odorous flowers, blooming in June. He found it ina
heavy forest front, at an altitude of 4500 feet.

Additional citations: COLOMBIA: Boyacd: Lawrance 156 (F--
708505). Chocé: Triana 2083, in part [Macbride photos 28385]
( F--830243--photo, Kr--photo).

AEGIPHILA LHOTZKIANA Cham.
Two additional synonyms are Aegiphila glandifera Casar. ex

Moldenke, Prolim. Alph. List Invalid Names 2, in syn. (1940)

and A. glandulifera Casar. ex Moldenke, Suppl. List Invalid
Nemes 1, in syn. (1941). Mello Barreto describes it as a tree
3m. tall or even 4 m. tall, with white flowers in November.
Markgraf collected it an an altitude of 1000 m. It has been
confused with A. Sellowiana and A. verticillata. -

Lithoprinted from Author's cs
EDWARDS BROTHERS,
Lithoprinters

ANN ARBOR, MICH!
1947 ‘

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April, 1948 No. 10

CONTENTS

NACHINO, J. V., A new species of Eupatorium ois
re RIG VY OSE MR TEINES io vg cbc vac hacaobeaesinnseccgreGasijeneess+us OO

[OLDENKE, H. N., Notes on new and noteworthy
PUTDER SDV e825 op aN Ob Let calves an csepuain eden songs onoodds eo S HOO

‘son, H. A, Notes on South American MelastOMES......1.00000+-428

ic NACHINO, J. V., A new species of ean
es): PREP OED eck cues ctu hel hs sotdden deta cnccmeavschioveet RO

Published by H. A. Gleason and Harold N. Moldenke
The New York Botanical Garden,
Bronx Park, New York 58, N. Y.

Price of this number, 75 cents; per volume, $5.00 in advance

Vol. 2, No. 9, was issued December 3, 1947

Beet tan iar he NEW YORE
F chee BOTANICAL
LARCAR I >t» Oe

A NOTE ABOUT AN AI-LEGED DISCUSSION OF JUNIPERUS

John F. Cornman

In the November 1947 issue of this Journal there appeared an
article (1) about investigations I have made in the taxonomy of
the genus Juniperus. Since I have not established a reputation
_ through taxonomic publications, I feel obligated to discuss
briefly the nature of the material reviewed by vanMelle, the
_ validity of a serious allegation, and, in the interest of ob-
_ jectivity, to present a paragraph not quoted by vanMelle.
a Faramount is the fact that the paper under discussion is a
_ thesis submitted to the Oornell University Graduate School. Two
copies are required by the Faculty of the Graduate School and,
after being catalogued, they are deposited in the University
_ archives. While one of the copies is made available to inter-
_ ested research workers, the paper itself has no more signifi-
cance than any other unpublished manuscript.

Since I have not published on Juniperus, no defense of un-
_ published taxonomic details is necessary. The intricacies pre-
: sented by vanMelle in the November paper are not new material,
and thus do not alter my personal opinions on the technicali-
ties. It is intended that my own observations will be published
as ea series of papers on taxonomic and related problems in the
genus.
The thesis man::script contains no comment on vanMelle's com-
pleted book (2), for it was not published until later. We await
a discussion of that work by a competent and disinterested re-
viewer. .
On page 354 of his November paper vanMelle states that I
_ have ethaction "vanMelle type labels to sheets not so designated
4 by me". Regardless of the intent, these words are an accusation
_ of forgery. WanMelle has cited the thesis pages (280,298) on
- which appear photographs of the allegedly mutilated sheets.
_ There the interested taxonomist will see routine annotation
slips with my own name conspicuously printed in black typeface.
Credits for assistance in the preparation of a voluminous
_ paper are traditionelly rendered in the Preface. Since I shall
_ not again have occasion to publish what I said there, I quote
; the pertinent passage:
. "Mr. P. J. vanMelle of the Fougnkeepsie Nursery has pointed
4 Out many cogent facts about cultivated Junipers, both in the
;.
"
;

field end in correspondence. This assistance was continued long
efter it became evident that our viewpoints were widely ache
ent. Rather strong criticisms are made here of Mr. vanvelle's

_ treatment of the J. chinensis group. Such disagreement with so
good a friend end ~puide is regrettable. It is directed solely
j at his interpretations ond treatment. The writer retains a

i 401

402 Rok YT Ob oO: Gates Vol. 2, no. 10

great respect and admiration for Mr. vanMelle's ability as a
plantsman, for his knowledge of his particular group, and es-
pecially for his patient good nature."

(1) vanMelle, P. J. in Fhytologia 2 :353-363, 1947.
(2) vanvelle, P. J. Review of Juniperus chinensis et. al. New
York, April 1947.

NOTES ON POLYGONUM. IIT

J» F. Brenckle

Polygonum Exsiccatum (Avicularia) Fascicle 1, issued in Jan-
uary Toty, is the first of a series to be issued at irregular
intervals. There will be 36 copies of this exsiccatum, the
specimens of each number to be as much alike as possible and to
be collected on the same date and at the same place. The speci-
mens have been selected to bring together forms that are often
mistaken for each other end also to group species for a phylo-
geographical study end classification which is to be published
later. Severel new species and forms are introduced, specimens
of which in this fascicle may be considered as co-type material.

Tne exsiccatum is being distributsd to active collaborators
and some larger herberia. Zontributions to this series are sol-
icited.

List of species and forms in Fascicle One:

No. 1. Polygonum interior new species, forna vernalis.
No. 2. Folygonum intsrior new species.
No. 3. Polygonum interior new species, forma rostratum.
No. 4. Folygonum interior var. Turneri Brenckle new variety.
No. 5. Folygonum remosissimum Michx., forma vernalis.
No. 6. Polygonum ramosissinum Michx.
No. 7- Polygonum remosissimum Michx., forma rostratum.
No. 8. Polygonum latum Small, forma vernalis.
No. 9. Polygonum latum Small.
No. 10. Folygonum prolificum (Small) Robins. var. autumnale

Brenckls new name.

No. 11. Polygonum prolificum (Small) Robins. var. profusum
Brenckle new variety.

No. 12. Polygonum prolificum (Small) Robins. var. profusum
Brenckle forma rostratum.

No. 13. Folygonun achoreum Blake.

No. 14. Folygonum cumporum Meisn., forma varnalis

1948 Brenckle, Notes on Polyzonum 403

No. 15. Folygonum camporum Meisn., forma rostratum.

No. 16. Polygonum argyrocoleon Steud.
No. 17. Folygonum argyrocoleon Steud.
No. 18. Folygonum Faronychia Cham. & Schl.

No. 19. Polygonum Faronychia Cham. & Schl.
No. 20. Polygonum ma jus (Meisn.) Piper.
No. 21. Polygonum emaciatum A. Nels.

No. 22. Folygonum emaciatum A. Nels.
No. 23. Folygonum Englemannii Greene.
No. 24. Polygonum sawatchnense Small.
No. 25. Polygonum tenue Michx.
Polygonum interior new species.

Herba annua atroviridis rectis gracilis multibrachiata
striata 4--83 dm. alta dein rubescens; ramis patentibus attenua-
tis argute angulatis; foliis caulium oblongo-lanceolatis ad
basin apicemque acutis 3--6 cm. longis 3--8 mm. latis fugaceis;
foliis ramorum reductis linearibus vel nullis; ocreis pallidis
ad basin rubellis inconspicuis.

Annual, dark green, erect, slender, much branched from the
base, striate, 4--8 dm. high, becoming reddish with age;
branches spreading, attenuate, sharply angled; stem-leaves ob-
long-lanceolate, pointed at each end, short-petioled, 3--6 cm.
long, 3--2 mm. wide, fugacious; branch-leaves reduced, linear
or absent; ocreae pale, reddish at the bese, inconspicuous; per
fanth sharply triangular, carinate, divided to near the base
into 5 segments, the 3 outer longer, white-margined or pink-
edged, 3 mm. long, pedicellate; achenes narrow, pointed, with
the apex edges sharper than those of the body, one of the faces
slightly umbonate, chestnut-brown, smooth, vaguely puncticu-
late, 2--3 mm. long, some later achenes becoming attenuated to
6 mm. long and exserted.

Among vegetation at the margins of ponds or in ditches. The
species differs from Polygonum exsertum and Folygonum leptocar-
pum in thet it matures most of its achenes within the pericarp
and produces exserted achenes late in the season or none. The
plants are generally smaller and more slender than F. exsertum.

Specimens examined: South Dakota, Brenckle nos. 3726, 572 ls
3652, 3848, 3865, 4165, distributed as P. exsertum; North Dako-
ta, Stevens nos. 212, 442, 463, 656; Alberta, Turner nos. 2314,
4283; Saskatchewan, W. P. Fraser no. 10, Aug. 1939.

Type locality: margin of a pond fed by artesian wells, half
mile northeast of Mellette,* South Dakota. The type specimen is
deposited in the Britton Herbarium at the New York Botanical
Garden. Co-type material is distributed in Fascivle 1, nos. l,
ea, De

Folygonum interior var. Turneri new variety.
Herba annua flavido-viridis; caule recto 6--9 dn. alto, med-

404 PHYTOLOGITIA Vol. 2, no. 10

fam elongato 6--7.5 cm. longo, basin versus incrasaates foliis
3--7 cm. longis 3--8 mm. latis.

Annual, yellowish green; stem erect, €--9 dm. high, the mid-
dle sections elongated, 6--7.5 cm. long, the lower half thick-
ened; leaves 3--7 cm. long, 3--8 mm. wide.

Collected by Dr. George H. Turner near Fort Seskatchewan,
Alberta, the type locality being one mile north of Fort Saskat-
chewan. The type specimen is deposited in the Britton Herbarium
at the New York Botanical Garden. Co-type material is distribu-
ted in Fascicle 1, no. 4.

Folygonum prolificum (Small) Robins.
This species is widely distributed, its area extending from

the Atlantic seaboard to the Inter-Mountain regions, Utah, and
from Cenada to Texas. It is completely at home on the Inland
Plains in various semi-srid and humid locations where it ~
assumes a variety of forms. It may be erect, slender and spar-
ingly branched, or it may become a robust, bushy, much branched
and spreading plant, or be completely prostrate. It was first
described as a variety of P. ramosissimum, but to this species
it has no close phylogenetic relationship. The general struc-
ture of the plant, leaves and achenes are quite distinct, nor
do the species hybridize. My conclusion is that this species
and its ancestor have long occupied this inland American erea
and were comnon on the shores of our anciert Inland Sea. The
flowing two verieties are distributed as nos. 10, ll, and 12
in Fascicle l.

Folygonum Toe (Small) Robins. var. autumnale Brenckle,
new name.
Published as P. autumnale in the Bulletin of the Torrey Bot-
anical Club, vol. 68, Pe 495.

Folygonum prolificum (Small) Robins. var. profusum new variety.
Herba annua robusta late patens ramosissima, seminibus

plerumque autumnale productis.

A robust wide-apreading annual, bushy, branching heavily
from the base. Its main crop of seed is produced in the fall.
Common in South Dakota. The type locality is a wet meadow half
a mile northeast of Mellette, South Dakota. The type specimen
is deposited in the Britton Herbarium at the New York Botenical
Garden. Co-type material is distributed as nos. 11 and 12 in>
Fascicle 1.

The Phylad Polyzonum Paronychia.
The species represented by nos. 13 to 25 of Fascicle 1 are

evidently genetically related and with some others form a well-
defined phylad of polygonums. The area of origin and the known
distribution are indicated. The characters comnon to the spe-

1948 Brenckle, Notes on Folygonum 405

cies of this section are (1) 6 raven-black achene, which, while
maturing, does not pass through shades of brown or chestnut
coloration. Usually it is smooth and shining, but may become
more or less punctate or striate in some species; (2) a dis-
tinctive and often large colored perianth; (3) the structure of
the leaves is suggestive, mostly narrow with revolute edges, a
prominent mid-rib and plication in some species,

Folygonum Faronychia Cham. & Schl. is 4 perennial with woody
stems. Its area is restricted, extending along the seacoast
from mid-Cealifornia to British Columbia; clearly a relic
species. The habitat given is "prostrate on sand along the
coast." Fascicle 1, specimen no. 18. Material collected from a
clay bluff was erect and bushy, and is represented by specimen
no. 19.

Folyzgonum majus (Meisn. ) Piper is represented by two forms:
(1) a robust, lerger, woody-stemmed form which at times is bi-
ennial and perhaps perennial. The area of this form is along
the shores and benches of the Columbia and Snake Rivers at ele-
vations of 50 to 200 feet. The benches mentioned represent old
sescoast lines before elevation of the country and were then
occupied, no doubt, by the ancestral PF. Faronychia. (2) A more
slender herbaceous form which occurs over the same area and to
higher elevations surrounding it. This form is often difficult
to distinguish from F. spergulariaeforme because the achenes,
leaves, flowers and distribution are similar.

Additional members of this phylad will be distributed in
future fascicles.

Folygonum emaciatum A. Nels.

This is a subspecies of Folygonum Douglassii Greene. Its
area is the more arid mountain regions. The distinguishing
characters are the linear leaves, slender stems, and striated
achenes which are somewhat shorter, 2.5--3 mm. long. Represent-
ed by nos. 21 and 22 in Fascicle l.

Among a number of polygonums sent me by Reverend arnest
Lapage of Quebec, collected in northern Canada and Alaska, is
one which is here described as a new species:

Fersicaria Oneillii new species.

Herba annua parva; caule breve basin versus brachiato pros-
trato; ramis gracilibus obscure 3- vel 4-angulatis rubello-
brunneis 1--6 cm. longis foliosis; foliis oblongo-lenceolatis
vel spathulatis ad epicem rotundatis, ad basin in petiolum
brevem attenuetis, 1.5--%5 cm. longis, 2--5 mm. latis, saepe
subtus tomentosis; in*lorescentiis axillaribus terminalibusque,
racemis interruptis ad apicem sphaericis vel ovalibus, 5--8 mm.
longis, 5 mm. latis.

A small annual; stems short, several- to many-branched near

406 PH Y°T.O LO GFA Vol. 2, no. 10

the tap-root, prostrate; branches slender, obscurely 3- or 4-
angled, reddish-brown, 1--€ cm. long, leafy; leaves oblong-
lanceolate or spatulate, rounded at the end and tavering to a4
snort petiole, often tomentose on the lower side, 1.5--3 cm.
long, 2--5 mm. wide; inflorescence in the axils of leaves and
in terminal, interrupted racemes, the terminal section being
spherical or oval, 5--3 mn. long, and 5 mm. wide; perianth
greenish, flettened, 5-perted to near the base, the sections
white-edged and sometimes reddish-tinted; echenes lenticular,
circular, biconcave, slightly umbonate, dark brown, dull, 2.5
mm. in diameter.

Type locelity: on the Nabesne Road, Mile 89, in Aleskea, June
24, 1947, collected by Dutilly, Lapage, ‘and O'Neill no. 21556.
This dwarf subarctic plantas related to Fersiceria tomentosa
(Schrenk) Bicknell end Fersiceria scabra (Moench). The leafy
inflorescence and the dull, reddish-brown “achenes clearly dis-
tinguish it. I teke pleasure in naming this species in honor
of Rev. Hugh C'Neill, who has long collected and studied the
northern Cenadian and Alaskan floras. The type specimen is de-
posited in the Langloie Herbarium at the Catholic University
of America, Washington, D. C.

A NEW SFECIES OF EUPATCRIUM FROM THE WEST INDIES

Joseph V. Monachino

In 1945 I had the pleasure of identifying specimens of
phanerogams collected by José I. Ctero and C. E. Chardon on
Mona Island, an island having an area of about twenty square
miles and lying midway between Puerto Rico and Hispaniola.
These specimens ere to serve as botanical vouchers for species
to be discussed in Dr. Ctero's forthcoming article on the veg-
etation of Mona Island. A new species of Eupatorium was dis-
covered among the collection. As it mignt be a rather long
time before his manuscript appears in print, Dr. Ctero has re-
guested that-I legitimize the name of this novelty by formal
publication at this time.

EUPATORIUM OTEROI Monechino, sp. nov. Fruticulus gluber;
foliis oppositis non punctatis, petiolis ca. 1--2 cm. longis,
laminis ovatis vel oveto-lanceolatis, ca. 2.5--5 cm. longis,
1.5--3.5 cm. latis, ad apicem acutis vel obtusis, ad basin
late cuneatis, plerumque serratis; inflorescentiis corymbosis
multicapitatis, capitulis ca. 12-floribus, involucris cylin-
drico-ellipticis, 4.5--5 mm. longis, brecteis imbricatis ca.
4-serietis oblongis vel anguste spathulato-oblongis, ca. 2--4

19428 Monachino, A new Eupatorium 407

mn. longis, 0.6 mm. latis, ad epicem rotundatis vel obtusis, 3-
striatis, apicem versus parce cilistis, caeterum glabris; sem-
inibus ca. 2--3 m. longis glabris, pappi setis ca. [l--] 1.6--
2-5 mm. longis.

Shrubby, glabrous; leaves opposite, not punctete nor resin-
ous, pergameneous, the petioles about 1--2 cm. long, sparsely
minutely ciliate at the base, the blades ovate to ovate-lance-
olete, ca. 2.5--5 cm. long, 1.5--3.5 cm. broad, acute to obtuse
at apex, broadly cuneate at base, serrate to coarsely crenate-
dentate or rarely entire, 3-nerved from the base, the reticula-
tion moderately expressed; inflorescence in many-headed cor-
ymbs, glabrous; cepitulum sessile or short-peduncled, ca. 12-
flowered (number of achenes], the flowers light purple [fide
Britton], the involucre cylindric-elliptic, 4.5--5 mm. long;
brects imbricated in ebout 4 series, rether rigid, oblong to
narrowly spetulate-oblong, about [1.6--] 2--4 mm. long, 0.6 mm.
broad, rounded or obtuse at apex, usuelly 3-striate (sonetimes
with 1 or 2 fainter lines), sparsely ciliate towerd the apex,
otherwise glabrous, achenes 3--S-angled, dark brown, ce. 2--3%
mm. long, completely glabrous, smooth, the pappus of ca. 20--25
bristles, white, barbellate, ca. [1--] 1.6--2.3 mm. long.

Type: Ctero & C. E. Chardén 821, limestone plateeu, Mone Is-
land, March 9, 1944, deposited in the Britton Harbarium at the
New York Botanical Garden, New York City, with a fragment in
the United States National Herbarium, Washington.

Additional material examined: Mona Island, Fuerto Rico:
Britton, Cowell & Hess 1672, coastal rocks, Sardinera, Feb. 20-
26, 1914; F. L. Stevens 6376, Dec. -20=-21., -1913:.(N. -Y. <Bot. Ga.)

Hupatorium Oteroi obviously belongs under the genus Osmia in
Britton & Nilson's Botany of Forto Rico and the Virgin Islands
(Selentific Survey of Forto Rico and the Virgin Islands, New
‘York Acad. Sci. 2 (2): 287. 1925). It fe easily distinguished
from all the species of Osemia described therein, however, by
the non-punctate character of its ieaves.

In a very superficial way, E. Oteroi ‘bears 6 resemblance to
E. corymbosum Aubl. It belongs in the Ser. Imbricata, Sect. Cy-
lindrocephala, as defined by De Candolle (Frod. Syst. Nat. 5:
141. 1836) or Sect. Cylindrocephala as defined by B. L. Robin-
son (Froc. Am. Aced. Arts & Sci. 54 (4): 269, 270. 1918).

Two species of Hupatorium are reported from Mona Island by
N. L. Britton (The vegetation of Mona Island, Annals Vissouri
Bot. Gard. 2: 49. 1915). The one named £. atriplicifolium Lam.
is &. Oteroi, judging from the material deposited in the herb-
erium of the New York Botanicel Garden. The second species is
E. odoratum L.

Se F. Blake was kind enough to conpere e fragment of the
type collection, and reports that he was not able to match it
in the West Indian material of Supatorium at the United States
Netional Herbarium.

NOTES ON NEW AND NOTEWORTHY PLANTS. IV

Harold N. Moldenke

ALOYSIA VIRGATA var. PLATYPHYLLA (Briq.) Moldenke, comb. nov.

Lippia virgata var. platyphylla Briq., Ann. Conserv. & Jard.
Bot. Genev. 7--8: 304, 190 °

BOUCHEA FLUMINENSIS var. PILOSA Moldenke, var. nov.

Haec varietas a forma typica speciei ramulis foliisque
rhachideque calyceque insigniter longe pilosis recedit.

This variety differs from the typical form of the species in
having the branchlets, leaves, rachis, and calyx conspicuously
long-pilose. The leaves are alternate or subalternate, deeply
serrate.

The type was collected by Christopher Sandeman (no. 4776) in
shade and semi-shede at Iguazu Falls, alt. 500 feet, Misiones,
Argentina, in “ay, 1944, and is deposited in the herbarium of
the Royal Botenic Gardens at Kew. The collector describes the
plent as a weak-growing shrub with rosy-lilac flowers which
fade and drop very rapidly after being gathered.

CLYRCDZENDRUM WALLII Moldenke, sp. nov.

Fruticulus; ramulis subgracilibus dense fusco-tomentosis;
foliis ternatis; petiolis gracilibus densissime fusco-tomentos-
is; laminis membranaceis ovato-ellipticis vel ellipticis vel
lenceolatis, ad apicem acutis vel sliquatenus attenuatis, ad
basin acutis vel scvminetis, reguleriter serratis, supra dense
puberulis, subtus densissime cinereo-tomentellis.

Bush; branchlets rather slender, densely tomentose with fus-
cous hair, less so in age; nodes annulate; principal internodes
1.5--3.5 cm. long; leaves ternate; petioles slender, 4--6 mm.
long, very densely fuscous-tomentose; blades membranous, dark-
green above, much lighter beneath, ovete-elliptic, elliptic, or
lanceolate, 4--7 cm. long, 1.5--3 cm. wide, ecute or somewhat
ettenuate-acute at apex, acute or acuminate at base, regularly
serrate from the widest part to the apex with acute or obtuse
antrorse teeth, rather densely puberulent above, very densely
tomentellous beneath with cinereous hair; midrib very slender,
plane above, prominulous beneath; secondseries very slender, 4
or S per side, ercuete-ascending, plane above, subprominulous
beneath, arcuetely joined near the margins, not leading direct-
ly into the teeth; vein and veinlet reticulation fine, usually
visible on both surfaces but not at all prominulous; inflores-
cence apparently terninal, about 3.5 cm. long and 2--2.5 cm.
wide, many-flowered, composed of several pairs of small cymes,
densely fuscous- or inc&énous-tomentellous throughout, the low-
est pair of cymes sometimes subtended by 3 foliaceous bracts

408

1948 Moldenke, New and noteworthy plants 409

similar to the leaves in all respects but smaller; peduncles

slender, 1--2.5 cm. long, densely tomentellous; pedicels very

slender, 1--4 mm. long, densely tomentellous; calyx tubular, a-

bout 4 mm. long and 1.5 mm. wide, densely short-pubescent, its

rim S-toothed, the teeth narrow, erect, about 1.5 mm. long,

_ subacute, pubescent; corolla white, exserted, its tube about 5
wm. long, lightly glandular or granular-puberulent on the outer
surface.

The type of this species was collected by Erik Wall -- in
whose honor it is named -- at Fort Ball, Uganda, on August 2,
°1926, and is deposited in the Erik Wall Herbarium at Stockholm.
It was identified by Berthold Thomas as "Clerodendrum aff.

Odontocalyx Thomas".

DURANTA SPRUCEI var. BREVIRACEMOSA Moldenke, var. nov.

Haec varietas a forma typica speciei recedit racemis usque
ad 3 cm. longis paucifloris et laminis foliorum supra glabris
subtus parce disperso-puberulis.

This variety differs from the typical form of the species in
having racemes only to about 3 cm. long and few-flowered, and
leef-blades that are glabrous above and merely sparsely scatter-
ed-puberulent beneath, the puberulence more dense on the midrib
and secondaries.

_ The type was collected by Oscar Haught (no. 6097) along
roadsides on the Zipaquira-Facho highway, at an altitude of
2000 m., Cundinamarca, Colombia, on August 20, 1947, and is de-
posited in the Britton Herbarium at the New York Botanical Ger-
den. The collector describes the plant as a stout shrub, 2 m.
tall, with rather showy purple flowers, and says it is abundant
at the type locality.

GYPSOPHILA PANICULATA f. PLENIFLORA Moldenke, f. nov.

Haec forma a forma typica speciei corollis plenis recedit.
This form differs from the typical form of the species in its
"doubled" corollas.

The type was collected by H- N. Moldenke (no. 8069) from
cultivated material at Watchung, Somerset County, New Jersey,
on July 4, 1934, and is deposited in the Britton Herbarium at
the New York Botanical Garden. Although common in cultivation,
this form does not appear to have been validly named hitherto.

JUNELLIA SERIPHIOIDES var. GLABRA Moldenke, vare nov.

Haec varietas a forma typica speciei ramis ramulisque spin-
isque foliisque calicibusque glabris vel subglabris et calici-
bus 5--8 mm. longis recedit.

This variety differs from the typical form of the species
in its glabrous or subglabrous branches, branchlets, spines,
leaves, and calyx, and in its calyxes being 5--8 mm. long.

The type of the variety was collected by Carlos A. O'Donell

- 410 Pa Y T’OLOG Pa Vol. 2, How 10”

(no. 3240) at Puerto Madryn, Chubut, Argentina, on October 24
or 25, 1945, and is deposited in the herbarium of the Naturhis-
toriska Riksmuseum at Stockholm.

LANTANA ARISTATA var. SUBSESSILIS Moldenke, var. nov.

Haec varietas a forma typica speciei recedit inflorescentiis
numerosis axillaribus congestis brevissime pedunculatis vel
subsessilibus; laminis foliorum subtus dense pubescentibus vel
subvelutinis; caulibus densiuscule treviterque pubescentibus,
pilis contortis patentibus; bracteolis dense hirtellis.

This variety differs from the typical form of the species in’
having ite rather numerous inflorescences crowded on very short
peduncles in the leaf-axils, or subsessile; the under surface
of the leaves is densely pubescent or subvelutinous; the stems
are rather densely snort-pubescent with twisted spreading hairs;
and the bractlets are densely hirtellous.

The type was collected by A. G. Schulz (no. 1456) on hill-
sides at Jujuy (Capital), alt. 1200 m., Jujuy, Argentina, in
February, 1936, and is deposited in the Britton Herbarium at
the New York Botanical Garden.

LANTANA CIFERRIANA Ekm. & Moldenke, sp. nov.

Frutex humilis perbrachiata; ramis gracilibus acutiuscule ©
tetragonis prorsus albido-strigillosis; foliis numerosis parv-
is; petiolis dense albo-strigillosis saepe submarginatis; lam-
inis leviter chartaceis oblongis vel ovalibus vel subrotundis,
ad apicem rotundatis, ad basin acutis vel plerumque acuminatis,
regulariter serrulatis, supra strigilloso-puberulis, subtus
dense puberulis et parce resinoso-granulosis; corolla alba.

Low shrub, abundantly branched; branches slender, rather a-
cutely tetragonal, rather uniformly whitish-strigillose
througnout; nodes rather obscurely annulate; principal inter-
nodes 0.4--4 cm. long; leaves numerous, decussate-opposite,
small; petioles very slender, 2--5 mm. long, densely white-
strigillose, often submargined; blades thin-chartaceous, deep-
green above, somewhat paler beneath, oblong or oval, varying to
subrotund, 5--14 mm. long, 4--10 mm. wide, rounded at the apex,
acute or usually acuminate at base, uniformly serrulate except
at the very base with rounded rather appressed teeth, strigil-
lose-puberulent above, densely puberulent and somewhat resinous
granular beneath; midrib and the 3 or 4 pairs of ascending sec-
ondaries very slender, often slightly subimpressed above, sub-
prominulous beneath; veinlet reticulation often subimpressed a-
bove, obscure beneath; inflorescence axillary near the tips of
the twigs, 1 or 2 per node, capitate; peduncles very slender or
filiform, 1.5--4.8 cm. long, rather densely white-strigillose
like the branches and twigs; heads hemispheric, to 1.3 cm. long
and 1.5 cm. wide in anthesis, ovate in fruit; bractlets broadly
elliptic or oval, 5--7 mn. long, 4--6 mm. wide, blunt at apex,

ee
a

1948 Moldenke, New and noteworthy plants 411

lightly strigillose on the upper surface, more densely whitish-
strigillose on the lower surface; corolla pure white, its tube
about 5 mm. long, densely puberulent outside, the limb about 4
mae wide, lightly puberulent outside and somewhat resinous-
granular.

The type of this species was collected by E. L. Ekman (no.
H.15967) on dry sterile hillsides at Hatillo, Valle del Cibao,
prov. Santiago, Dominican Republic, on September 17, 1930, and
is deposited in the herbarium of the Naturhistoriska Riksmuseum
at Stockholm. The species is dedicated to Dr. R. Ciferri, who
urged Skman to pay more attention to the species of the Lantana
reticulata-Lantana involucrata complex.

LANTANA CUJABENSIS var. PUNCTATA Moldenke, var. nov.

Haec varietas a forma typica speciei foliis minoribus plus-
minusque bullatis subtus dense resinoso-punctatis et in venas
plusminusque breviter pilosis recedit.

This variety differs from the typical form of the species in
having smaller more or less bullate leaves which are densely
resinous-punctate beneath and more or less short-pilose on the
venation beneath.

The type was collected by Fred Alexander Barkley, Jairo
Correa Velasquez, and Gabriel Gutiérrez Villegas (no. 1536) in
an open pasture close to Ceja, altitude about 2180 m., Antioqui-
a, Colombia, on November 1, 1947, and is deposited in the her-
barium of the Facultad Nacional de Agronomia, Medellin,
Colombia. |

LANTANA GLUTINOSA var. ORIENTALIS Moldenke, var. nov.

Heaec varistas a forma typica speciei caulibus ramisque foli-
isque bracteolisque densissime puberulis, bracteolis breviori-
bus angustioribusque ad apicem obtusis, et corollis albis rece-
dit.

This variety differs from the typical form of the species in
neving the pubescence on the stems, branches, leaves, and bract-
lets much shorter (very densely puberulent rather than villous-
hirsute), the bractlets much shorter and narrower, obtuse at
the epex, and the corollas white.

The tyre of this variety was collected by J. Hanbury-Tracy
eae) in fairly dry ground among spaced scrub and low trees
at Le “esa, dist. Campo Ella, altitude 5000 feet, Mérida, Ven-
ezuela, on August 14, 1938, and is deposited in the herbarium
of the Royal Botanic Gardens at Kew.

LANTANA HAUGHTII var. OBTUSIBRACTEATA Moldenke, var. nov.

Haec varietas a forma typica speciei bracteolis ad apicem
regulariter obtusis vel rotundatis recedit.

This variety differs from the typical form of the species in
having its bractlets uniformly obtuse or even rounded at the a-

412 PR ¥°TO°R"O' Gres Vol. 2, noe 10

CXe

: The type was collected by Oscar Haught (no. 5132) in dry
pastures at Mercaderes, altitude 1100 m., Cauca, Colombia, on
October 24, 1946, and is deposited in the Britton Herbarium at
the New York Botanical Garden. The collector describes the
plant as a slender shrub to 2 m. tall, with showy inflorescen-
ces of rose-colored flowers. :

LANTANA HAUGHTII var. PARVIFOLIA Moldenke, var. nov.

Haec varietas a forma typica speciei bracteolis ad apicem
uniforme obtuseque rotundatis et laminis foliorum 1--1.8 cm.
longis et 6--13 mm. latis recedit.

This variety differs from the typical form of the species in
having its bractlets all obtusely rounded at the apex and the
leaf-blades only 1--1.8 cm. long and 6--13 mm. wide.

The type was collected by Oscar Haught (no. 5902) near
Chocanta, Cundinamarca, Colombia, on June 30, 1947, and is de-
posited in the Britton Herbarium at the New York Botanical Gar-
den. The collector describes the plant as a low spreading
shrub, 50 cm. tall, with flowers that open cream-color and turn
red, “unusually beautiful for the genus".

LANTANA LEUCOCARPA Urb. & Ekm., spe nov.

Frutex humilis; ramis elongatis horizontaliter reclinatis
gracillimis dense hispidulis tetragonis; foliis numerosis; pet-
jolis gracillimis hispidulis; laminis mambranaceis atroviridi-
bus triangulari-ovatis, ad apicem obtusis, ad basin truncatis
vel subtruncatis, regulariter serratis, supra parce puberulis,
subtus dense puberulis; corollis roseo-purpureis; fPructibus
albis.

Low shrub; branches elongated, horizontal-reclining, very
slender, densely hiespidulous with divaricate grayish rather
stiff hairs, less so on the more exposed parts, rather acutely
tetragonal or sometimes obtusely so; nodes not plainly annu-
late; principal internodes 1.5--4 cm. long; leaves abundant,
decussate-opposite; petioles very slender, 1--3 mm. long, his-
pidulous like the branches; blades membranous, rather uniform-
ly deep-green on both surfaces, triangular-ovate, 1.5--2.8 cm.
long, 9--16 mm. wide, obtuse at apex, truncate or subtruncate
at base, uniformly serrate from the widest part to the apex
with rounded teeth, lightly puberulent above, densely puberu-
lent beneath; midrib and the 4--6 pairs of ascending second-
aries very slender, obscure above, very faintly prominulous be-
neath; veinlet reticulation obscure or indiscernible; inflores-
cence axillary, capitate; peduncles very slender or filiform,
2.5--5.5 cm. long, spreading-pubescent, 2 per node, abundant;
heads hemispheric, about 1 cm. long and 1.5 cm. wide at anthe-
sis, many-flowered; bractlets ovate, 5--6 mm. long, 5 mm. wide
at the base, triangular-acute at apex, puberulent on the upper

ee

1948 Moldenke, New and noteworthy plants 41%

surface, densely short-pubescent on the lower surface; corolla
rose-purple, its tube about 5 mm. long, very densely puberulent
outside, the limb about 5 mm. long and 4 m. wide, densely pub-
erulent on the back; fruit perfectly white.

The type of this curious species was collected by E. L.
Ekman (no. H.15998) in fields on hillsides at Hato del Yaque,
Valle del Cibao, prov. Santiago, Doninican Republic, on Septem-
ber 27, 1930, and is deposited in the herbarium of the Natur-
higtoriska Riksmuseum at Stockholm.

LANTANA PARANENSIS Moldenke, sp. nov.

Herba perennis vel suffrutescens; caule ut videtur simplex
obtuse tetragono dense glanduloso-pubescente and albo-hirsutu-
lo; petiolis gracillimis gleanduloso-pubescentibus et hirsutulis
vel subobsoletis; laminis firme chartaceis utrinque griseo-
viridibus ellipticis vel suboblanceolatis, ad apicem obtusis
vel rotundatis, ad basin acutis vel attenuatis, crassiuscule
serrato-dentatis utrinque dense glanduloso-pubescentibus, sub-
tus parce hirsutulisj venis supra arzute impressis.

Perennial herb or subshrub; stem apparently simple, to 4 dm.
tall, obtusely tetragonal, densely glandular-pubescent and also
hirsutulous with longer, divaricate, white, non-glandulose
hairs, the hairs densest toward the apex of the stem; nodes not
plainly annulate; principal internodes 1.5--4 cm. long; leaves
decussate-opposite; petioles very slender, 1--2 mn. long and
glandular-pubescent and hirsutulous, or subobsolete; blades
firmly chertaceous, rather grayish-green on both surfeces, el-
liptic or almost oblanceolate, 2.5--4 cm. long, 5--11 mm. wide,
obtuse or rounded at the apex, acute or ettenuate at base,
rather coarsely serrate-dentate to below the middle with rether
bluntish revolute-margined teeth, rether densely glandular-
pubescent on both surfaces, the lower surface also bearing
scattered hirsutulous hairs like the stems; the midrib and ven-
etion deeply impressed above, giving the blade a decidedly bul-
late appearance, very prominent beneath; inflorescence axil-
lary, 2 at each of the upper nodes, about equaling the subtend-
ing leaves; peduncles very slender, to 4.5 cm. long, densely
glandular-pubescent and hirsutulous like the stems; heads hemi-
spheric, about 6 mm. long and 10 mm. wide; brectlets nerrow-
elliptic, 4.5--5 mm. long, ebout 1 mm. wide, densely glandular-
pubescent and hirsutulous, blunt or subacute at apex; corolla
slightly surpassing the bractlets, puberulent on the outer sur-
face,

The type of this species was collected by Fer Karl Hialmar
Dusen in the campo at Lago, Parana, Brazil, on December 2,
1910, and is deposited in the herbarium of the Naturhistoriska
Riksmuseum at Stockholm. Its narrow, blunt, bullate leaves and
glandulose pubescence render this s very distinct and unmistek-
able species.

414 PHY’ TO b-OG@ BS Vol. 2, no. 10

LIPFIA CAMPESTRIS Moldenke, sp. nov.

Planta perennis humilis ad basin lignosa; caulibus numerosis
simplicibus dense glanduloso-pubescentibus et albo-hirsutulis;
foliis sessilibus ovetis, ad apicem subacutis vel obtusis, ad
basin rotundatis vel cordatis, integris ciliatis utrinque hir-
sutulo-pubescentibus, sibtus parce glandulosis.

Dwarf perennial, woody at the base; stems numerous, simple,
15--17 cm. tall, rather densely glandular-pubescent and also
hirsutulous with much longer, divaricate, white, glandless
heirs; nodes not noticeably annulate; leaves decussate-oppo-
site, sessile; blades ovate, 7--16 mm. long, 4--10 m. wide,
the lower pairs increasingly smaller, subacute or obtuse at a-
pex, rounded or cordate at base, entire, ciliate, hirsutulous-
pubescent on both surfaces and somewhat glandulose beneath;
larger venation rather obscure above, very faintly subprominu-
lous beneath; inflorescence axillary, 2 per node, in the median
or upper axils; peduncles very slender, 1.5--2.4 cm. long,
rather densely glandulose-pubescent and hirsutulous; heads hem-
ispheric, less than 1 cm. long; breactlets ovate, about 5 mm.
long, 2 mm. wide at the base, triangular-attenuate at apex,
densely glandular-pubescent and hirsutulous; corolla about 7 or
7-5 mm. long, the tube about equaling the bractlets, uniformly
puberulent outside, the limb puberulent on the back, glabrous
within.

The type of this species was collected by Per Karl Hjalmer
Dusén on a campo at Jeguariahyve, Parana, Brazil, on October 9,
1911, and is deposited in the herbarium of the Naturhistoriska

Rikemuseum at Stockholm.

LIPPIA CHACENSIS Moldenke, sp. nov.

Frutex; caulibus gracilibus obtuse tetragonis canescento-
puberulis dein strigillosis; ramis ut videtur paucis brevibus;
foliis numerosis; petiolis gracillimis dense canescento-strigo-
sis; laminis leviter chertaceis anguste ellipticis, ad apicem
acutis, ad basin attenuatis vel subacuminatis, uniforme serru-
latis, supre dense adpresso-strigillosis, subtus densissime
flevescento-velutinis.

Shrub, 6--7 dm. tall; stems slender, obtusely tetragonal,
canescent-puberulent, strigillose on the older parts; nodes
rather indistinctly annulate; branches apparently few and
short; leaves decussate-opposite, abundant; petioles very
slender, 2--6 mm. long, densely canescent-strigose; blades
thin-chartaceous, lighter beneath, narrowly elliptic, 2--6 cm.
long, S--15 mm. wide, acute et the apex, uniformly serrulate a-
long the margins to below the middle, attenuste or subacuminate
at base, densely appressed-stririllose above, very densely vel-
utinous with very short flavescent hairs beneath; the slender
midrib and 5 or 6 pairs of secondaries, and often the larger
veinlets, slightly subimpressed above, prominulous beneath; in-

1948 Moldenke, New and noteworthy plants 415

florescence exillary, 1 or 2 per node, shorter than the sub-
tending leaves; peduncles very slender, 1--1.5 cm. long, canes-
_ cent-strigillose; heads hemispheric or oblong, to about 1 em.
long; bractlets broadly ovate, 4--4.5 mm. long, about 2 mm.
_wide at the base, acuminate at apex, densely strigose or strig-
illose, canescent, glandulose; corolla-tube about 6 mm. long,
very densely strigose on the outside, the limb about 4 m.
wide, glabrous or slightly pubescent at the base.

The type of this species was collected by Robert =. Fries
(no. 1445) in ean open grassy cempo at Tatarenda, Gran Checo,
Bolivia, on March 22, 1902, and is deposited in the herberium
of the Neturhistoriska Riksmuseum at Stockholm. The collector
describes the species as rare and the flowers as "lividis".

LIFFIA LORENTZII Moldenke, sp. nov.

Frutex; ramis ut videtur multibrechiatis irregularibus
griseis; ramulis gracilibus obtuse tetragonis canescento-
strigillosis; internodiis abbreviatis; foliis confertis; petio-
lis gracillimis vel obsoletis dense canescento-strigillosis;
laminis parvis lanceolato-ellipticis utrinque dense canescento-
strigillosis acutis uniforme serrulatis, ad basin longe attenu-
atis; spicis elongato-capitatis; bracteolis ovatis acuminatis.

Shrub; stems apparently much branched and irregular, gray;
branchlets slender, obtusely tetragonal, canescent-strigillose;
nodes rather indistinctly annulate; internodes abbreviated, 5--
30 mm. long; leaves decussate-opposite, mostly clustered on
very short twigs; petioles very slender, 1 mm. long or obso-
lete, densely canescent-strigillose; blades small, lanceolate-
elliptic, densely canescent-strigillose on hoth surfaces, 5--15
mm. long, 3--5 mm. wide, acute at apex, uniformly serrulate al-

most to the long-attenuate base; the slender midrib and second- ~

aries more or less impressed above and very prominent beneath;
inflorescence axillary, 2 per node; peduncles very slender, 1
cm. long or less, densely canescent-strigillose; spikes elon-
gate-capitate, about 1 cm. long; brectlets ovate, 2.5--3 mm.
long, acuminste at apex, densely canescent-strigose, ciliate;
corolla 4--5 mm. long, puberulent outside.

The type of this species was collected by P. G. Lorentz and
G. Hieronymus at Dragones, Salta, Argentina, in the middle of
August, 1873, and is deposited in the herbarium of the Natur-
historiska Riksmuseum at Stockholm.

LIPPIA ROSMARINIFOLIA var. STEWARTI Moldenke, var. nov.

Haec varietas a forma typica speciei laminis foliorum pin-
nato-lobatis recedit.

This variety differs from the typical form of the species in
having pinnately lobed leaf-blades, the lobes on smaller leeves
being tooth-like, divergent, 1--3 per side.

The type was collected by Alban Stewart (no. 3307) on the

416 Put Tt .O:b 0-4 Tesh Vol. 2, noe 10

sides of the mountain, to 4000 feet altitude, Tagus Cove, Albe-
marle Island, Galapagos Islands, on March 27, 1906, and is de-
posited in the herbarium of the California Academy of Sciences.

PAEPALANTHUS ANDICOLA var. VILLOSUS Moldenke, var. nove

Haec varietas a forma typica speciei per omnes partes den-
siore albo-villosis et praecipue vaginis longe albo-villosis
recedit.

This variety differs from the typical form of the species in
being more densely white-villous throughout and especially in
having the sheaths villous with long white hairs from the base
to within about 1 mm. of the truncated apex.

The type was collected by Oscar Haught (no. 5878) on steep
dry slopes, altitude 1800 m., along the Gacheta-Ubala highway,
Cundinamarca, Colombia, on June 20, 1947, and is deposited in
the Britton Herbarium at the New York Botanical Garden.

STACHYTARFHETA JAMAICENSIS f. MONSTROSA (Moldenke) Moldenke,
combe NOVe
Stachytarpheta indica f. monstrosa Moldenke, Prelim. List
Invalid Names 7, hyponym (1940); Phytologia 1: 433--434. 1940.

STACHYTARFHETA RIVULARIS Moldenke, sp. nov.

Suffrutescens; ramis acutiuscule tetragonis stramineis, ju-
ventute et ad nodos parciuscule lLongeque pilosis, dein et ad
internodos glabris; petiolis late alatis utrinque parcissime
pilosulis vel glabris; laminis leviter chartaceis ellipticis
breviter acuminetis, ad basin longe acuminetis, supra subscab-
ridis, utrinque parcissime strigilloso-pilosulis glabrescenti-
bus; rhachide profundo excavato dense puberulis sub anthesin.

Suffrutescent, 3--4 m. tall; branches rather acutely tet-
ragonal, stramineous, rather sparsely long-pilose at and near
the nodes, especially on the younger parts, glabrous on the
internodes and older parts; younger nodes annulate; principal
internodes 2--4.5 cm. long; leaves decussate-opposite; petioles
indistinct, broadly winged, about 2 cm. long, the wings taper-
ing into the base of the blade, very sparsely and indistinctly
pilosulous or glabrate on both surfaces; blades thin-chartace-
ous, elliptic, uniformly green on both surfaces, 7--9.5 cm.
long, 3--4.5 cm. wide, short-acuminate at apex, long-acuminate ~
into the petiole at base, somewhat scabridous above, very
sparsely strigillose-pilosulous on both surfaces, especially
along the larger venation, when young, glabrescent in age; mid-
rib slender, plane above, prominent beneath; secondaries very
slender, 5--7 per side, ascénding, slightly arcuate, indis-
tinctly: anastomosing in many loops near the margins, plane a-
bove,prominulous beneath; veinlet reticulation rather sparse;
inflorescence spicate, terminal, to about 30 cm. long in fruit;
rachig stout, deeply excavated, densely puberulent in anthesis,

1948 Moldenke, New and noteworthy plants 417

obscurely so in fruit; peduncle short, 2.5--3.5 cm. long; flow-
ers imbricate; bractlets ovate-lanceolate, 5--6 mm. long, ca.
1.5 mm. wide near the base, subacuminate at apex, glabrous or
subglabrous except for the minutely ciliolate margins; calyx 9-
10 mm. long, subdglabrats or glabrous; corolla about 2.3 cm.
long, maroon.

The type of this distinct species was collected by Ynes
Mexia (no. 1789a) along a stream at Arroyo de Los Tapeistes,
altitude 1425 m., Hacienda del Ototal, San Debastian, in the
Sierra Madre Occidental, Jalisco, ee on March 3, 1927, and
is deposited in the herbarium of tne Celifornia Academy of
Sciences at San Francisco. Tne collector records the common
names “chupa-miel" and "chupa-muerto". The species seems to be
related to S. acuminata F. 5C., but mey be distinguished at
once by its foliar and inflorescence characters.

STILBACEAE Lindl.

This family name pig oa as such in Lindley's "The Natural
System of Botany", ed. 2, p. 279 (1836), where there is a fine
description of the family and the neme is validly published in
every respect. In Lindl., Veg. Kingd., 4d. 2, pp. 594 & 607
(1847) Lindley repeats the name Stilbsceae as a valid and ac-
cepted family segregated from the Verbenaceae, although Lindley
always referred to families as "netural orders" -- a practice
continued even in such manuals as Asa Gray's “anual until a
rather recent date. On page 43 of the 1847 work Lindley cites
also a “Suborder” Stilbacei in the fungi -- obviously as a sub-
family and not of family rank. He gives no indication that any-
one had proposed or regarded it as of “order” or family rank up
to that date. This name Stilbacei for a group of fungi starts
in Link, Abhandlungen der Koniglichen Akadenie der Wissenschaf-
ten zu Berlin 1824: 181 (1826). It has been claimed by some
botanists that this name is a family name, is an orthographic
variant of Stilbaceae, antedetes Lindley's name by ten years,
and therefore invalidates Lindley's name.

A study of Link's paper, entitled “Entwurf eines phytolog-
ischen Pflanzensystems nebst einer Anordnung der Kryptophyten",
which covers pages 145 to 194 of the volume cited above, shows
conclusively that Link did not regard this group as e family in
our present sense of the term, and was not proposing the name
as a family name. He slesstiies the group under his "Cl. I.
Cryptophyta"»[p. 154], "O. 1. Fungi" [p. 162], “Subordo 3.
Vycetes" [p. 168], “Reihe XI" [p. 177; see p. 179, lines 31 and
33, for proof of his application of the term "Reihe" to these
categories], and "Familie IV. Gastronycetes" [p. 181]. That he
regards his groups I. Epiphyti, II. Sclerotiaceae, III. Tremel-
loidei, IV. Gastromycetes, V. Sphaeriacei, VI. Sarcomycetes,
VII. Fhalloidei, and VIII. Agaricini as families is plainly in-
dicated by the text. For instance, under Epiphyti he says

418 PH YY Ont-Oet eA Vol. 2, no. 10

"Diese Familie enthélt die Anfinge viele andern Familien". Un-
der Gastromycetes he says “Auch diese Familie ist aus mehreren
kleinern Haufen zusammengesetzt", and he then proceeds to enum-
erate and (in most cases) describe ten such "Haufen" or groups
within the family, namely, (1) Dimidiati, (2) unnamed, (3) Nem-
asporei, (4) Sporigastrei, (5) Stilbacei, (6) Lycoperdei, (7)
Cyathoidei, (8) Carpobolei, (9) Tuberacei, and (10) unnamed.

It thus seems apparent that Link proposed the name Stilbacet
as a subfamily group. The earliest reference that I heve been
able to discover for Stilbaceae as a family name in the fungi,
with e validating description or reference to an earlier valid-
ating description, is by Lindau in Rabenhorst, Krypt. Fl., ed.
2, 8? 5 (1904). The earlier reference to "Stilbaceae Fries" in
Saccardo, Syl. Fungi 16: 1082 (1902) is unaccompanied by a val-
idating description or reference to a validating description.
The possibility that Link's Stilbacei was validly raised from
subfamily to family rank between 1826 and 1836 seems remote,
especially since Lindley in 1836 knew of no such action. I am
therefore.maintaining Lindley's Stilbaceae as a legitimate and
valid name in the phanerogams, and am regarding Lindau's and
Fries' "Stilbacese" forthe funzgous group an illegitimate
homonym which must be replaced.

SYNGONANTHUS STEYERMARKII Moldenke, sp. nov.

Herba minuta caespitosa; caulibus valde abbreviatis; foliis
linearibus vel ecicularibus rectis numerosis arcte imbricatis
(non adpressis) obtusis glabris; pedunculis solitariis ca. 5
mm. lorgis teretibus non striatis non contortis glabris; vag-
inis adpressis glabris non striatis non contortis, ad apicem
fissis 2-lobatis, lobis subspathulatis rotundatis; ceapitulis
solitariis subglobosis sordidis 4--€-floris.

Minute tufted herb, forming moss-like mats; stems greatly
abbreviated, the entire plants usually less than 1 cm. tall;
leaves bright-green, linear or acicular, 2--3 mm. long, erect,
numerous, closely imbricate but not appressed, blunt-pointed,
glabrous but often enclosing a cushion-like mass of whitish
hairs at the tip of the stem; peduncle solitary, about 5 mm.
long, terete, not striate nor twisted, glabrous; sheath appres-
sed, about 4 mm. long, glabrous, not striate nor twisted, split
at the apex, the 2 lobes completely separate, each about 1.5
ma. long, subspeatulate, rounded at the apex; heads solitary,
subzlobose, sordid-white, about 1.5 mm. wide, 4--6-flowered;
involucral bractlets € or less, dark-brown or brownish-black,
shiny, firm-textured, broadly ovate or suborbicular, about 1.3
mme long and wide, acute at the apex, densely ciliate-margined
with whitish cilia; receptacle densely long-villous; receptacu-
lar brectlets brown, elliptic-oblanceolate, about 0.8 mm. long
and 0.3 mm. wide, acute at the apex and there densely bearded;
pistillate florets only seen: sepals 3, brown, separate almost

1948 Moldenke, New and noteworthy plants 419

to the base, broadly elliptic, about 1 mm. long and 0.6 mn.
wide, more or less concave on the inner and convex on the outer
surface, acute at the apex, more or less long-pilose on the
back with easily rubbed-off hairs, densely and persistently
white-bearded at the apex on the back; pstals 3, connete into a
slender, erect, hyaline tube about 0.9 mm. long or less, long-
pilose; overy deeply 3-lobed, about 0.6 mm. long, glabrous, 3-
cglled, 3-ovulate; style about 0.2 mm. long, glabrous; the 3
stigmas and 3 style-appendages all about 0.2 m. long and
arising from the same point.

The type of this very distinct and amazing species was col-
lected by my good friend, Julian A. Steyermark (no. 57372) --
in whose honor it is named -- on limestone outcrops of the
Piramo de Tama, altitude 3045--3475 m., near the Colonbia-
Venezuelan boundary, Téchira, Venezuela, on July 15, 1944, and
is deposited in the Britton Herbarium at the New York Botanical
Garden. The general aspect of the plant is that of 4 very min-

ute Paepalanthus.

VERBENA AUSTRALIS Moldenke, sp. nov.

Herba; caulibus decumbentibus, ad apicem adscendentibus, non
velde brachiatis, gracilibus acute tetragonis sulcatis leviter
strigilloso-pubescentibus; foliis paucis; petiolis gracilibus
marginatis strigillosis; laminis leviter chartaceis ovatis pro-
funde trilobatis, lobis dissectis obtusis, supre parce obscure-
que strigillosis, subtus punctatis et densiore strigillosis;
bracteolis lanceolatis albo-ciliatis caeterum glabris.

Herb with decumbent stems, ascending at their tips, not much
branched; stems and branches slender, acutely tetragonal, sul-
cate, lightly strigillose-pubescent with spreading or subap-
pressed hairs; principal internodes 2--8.5 cm. long; nodes an-
nulate; leaves rather sparse, decussate-opposite, sometimes
with a few small ones in their axils; petioles slender, 5--5
mm. long, margined, strigillose; blades thin-chartaceous,
rather uniformly bright-green on both surfaces, ovate in out-
line, 1--3 cm. long, 0.8--1.8 cm. wide, deeply 3-lobed, the
lobes again dissected with rather broad and blunt secondary
lobes, very sparsely and obscurely strigillose above, somewhat
more densely so and runctate beneath, the very slender midrib
and seconderies often slightly subimpressed above and prominu-
lous beneath, veinlet reticulation indiscernible on both sur-
faces; inflorescence spicate, terminal and in the uppermost
axils, densely congested in anthesis, later elongating to a-
bout 5 cm.; peduncles slender, tetragonal, rather densely strig¢
illose-pilogulous with very short white antrorse hairs; rachis
densely puberulent; bractlets lanceolate, 3--4 mm. long, sbout
1 mm. de, acute et apex, glabrate except for the white-
ciliolate margins; calyx tubular, about 5 mm. long, very minu-
tely strigillose on the 5 ribs or glabrate, the teeth short, a-

420 PA YO) OG ek Vol. 2, no. 10

cute, not appendaged, membranous and purplish between the ribs
toward the apex; corolla-tube about 6 mm. long, very sparsely
and minutely pilosulous on the outside above the calyx; corolla-
limb about 5 mm. wide, very minutely and sparsely pilosulous on
the outside.

The type of this species was collected by Per Karl Hjalmar
Dusén (no. 15190) in wet almost swampy soil at Jagueriahyva,
Farané, Brazil, on October 10, 1911, and is deposited in the
herbarium of the Naturhistoriske Riksmuseum at Stockholm.

VERBENA CHEITMANIANA Moldenke, sp. nov.

Herba porrecta; caulibus saepe procumbentibus gracillimis e-
longatis acute tetraconis strigillosis vel glabrescentibus; ram-
is pauciusculis grecillimis elongetis porrectis acute tetragonis
saepe subsulcatis densiuscule albido-strigillosis, pilis brevis-
simia reflexis; foliis'ut videtur ad apicem ramulorum aggrega-
tis; petiolis grecilibus marginatis dense piloso-pubescentibus,
pilis subadpressis albidis antrorsis; laminig membranaceis non
brunnescentibus, triangulari-ovetis attenuato-acutis, ad besin
cunesto-acuminatis, irregulariter incisis saepe plusminusque
distincte trilobetis utrinque adpresso-strigillosis, lobis in-
tegris vel plerumque incisis, dentibus acutis.

Sprawling herb; stems often procumbent, very slender, elon-
gate, scutely tetragonal, strigillose or glabrescent; branches
rether few, very slender, elongate, sprawling, ecutely tetrago-
nel, often slightly sulcate between the angles, rather densely
strigillose with very short and whitish reflexed hairs; nodes
annulate; principal internodes 1.i-~7.5 cm. long; leaves decus-
sate-opposite, apparently often numerous only toward the tips
of the branches; petioles slender, 1--1.5 cm. long, margined,
densely pilose-pubescent with subappressed, whitish, antrorse
hairs; blades membranous, somewhat lighter green beneath, not
brunnescent in drying, triangular-ovate in outline, 1.5--4 cm.
long, 1--3 cm. wide, attenuate-acute at the apex (or merely a-
cute on smaller leaves), cuneately acuminete into the petiole at
base, irregularly incised, often more or less distinctly 3-
lobed, appressed-strigillose on both surfaces, more densely so
when immature, the lobes entire or more usually incised, the
teeth all acute at epex; midrib and the 1--3 pairs of second-
aries very tenuous, plane or obscure above, very slightly pro-
minulous beneath; veinlet reticulation indiscernible above, ob-
scure beneath; inflorescence terminal and in the uppermost leaf-
axils, the floriferous portion congested-spicate, densely many-
flowered, apparently 2--3.5 cm. long, showy: peduncles very
slender, 6--10 cm. long, rather obtusely tetragonal, sometimes
slightly sulcate, stramineous, fairly densely short-pilose or
strigillose with whitish reflexed hairs; bractlets lanceolate,
4--€ mm. long, about 1 mm. wide at the base, long-attenuate at
the epex, rether densely strigose with antrorse white hairs;

1948 Moldenke, New and noteworthy plants 421

calyx cylindric, 8--S mm. long, densely strigose with closely
appressed, antrorse, white hairs, its rim unequally toothed,
the teeth long-eristate, the tips connivent before and after
anthesis, purplish; corolle-tube 1C--11 mm. long, zlabrous or
very obscurely pilosulous above the calyx outside, villous in
tne throat within; corolla-limb 7--& mm. wide, patent, the
lobes deeply cordate at the apex, glabrous on both surfaces.
The tyre of this most interesting species was collected by
Robert =. Fries (no. 477) in shady pleces under Salix Humboldt-
fans at Piquete, on the sandy banks of Rfo San Frencisco,
Jujuy, Argentina, on August 21, 1901, and is deposited in the
herberium of the Naturhistoriska Riksmuseum at Stockholm. The
species reminds one in the general appearance of its leaves of
the North American V. canadensis (L.) Britton. It is named in
honor of Philip Cheitman, Americen educator and ardent student
of Nature, who has re-dered considerable help to me in my re-
searches on tropical and extra-tropical American plants.

X VERBENA CLEMENSORUM Moldenke, hybr. nov.

Herba rudis ut videtur alta et multibrechiate; caulibus ram-
ulisque acutiuscule tetragonis striato-canaliculatis glebris;
nodis plerumque elongatis; foliis chertaceis rigidis ovatis
profunde irregulariterque incisis saepe trilobatis, supra plus-
minusque scabris vel scabro-marginatis, subtus scabris.

Coarse herb, apparently quite tall and much-branched; stems
and branches rather acutely tetraconal, striate-canaliculate,
glabrous; nodes annulate; principal internodes mostly elongate,
3.5--€.5 cm. long; leaves chartaceous, stiff, uniformly green
on both surfaces, ovate in outline, deeply and irregularly in-
cised, the larger more or less 3-lobed, 2.5--8 cm. long, 1--3
cm. wide, the smaller ones very scabrous above, the larger ones
glabrous or scabrous-margined, all more or less scabrous be-
neath (especially tne smaller ones just beneath the inflores-
cence), the slender midrib and secondaries usually impressed a-
bove and quite prominent beneath, the margins usually more or
less subrevolute; petioles absent or so strongly alate as to
merge completely with the blade; inflorescence spicate, com-
pound, the peduncles and rachis acutely tetragonal, minutely
and rather sparsely puberulent-pulverulent, apparently somewhat
glandular, the floriferous portion of the evikes elongate (of-
ten to 20 cm. long), rather densely flowered;bractlets ovate,
upwardly curvate, about 2 mm. long, subacuminate at apex, keel-
ed on the beck, minutely puberulent-pulverulent on the back,
ciliolate-margined, about equaling or slightly shorter than the
calyx; calyx 2--2.5 mm. long, glandular-puberulent; corolla-
tube about 3 mm. long, subglabrate or very minutely pulverulent
outside, the limb about 2 mm. wide; cocci about 1.6 mm. long.

The type of this natural hybrid was collected by Mary Knapp
Clemens at Jackson, Amador County, California, on Septemher 13,

422 PAY? OOB*O% TMA Vol. 2, no. 10

1920, and is deposited in the herbarium of tne California Aca-
demy of Sciences. It is named in honor of Mre. Clemens and her
husband, Joseph Clemens, who have done such noteworthy collect-
ing of members of this group in Michigan, Oklahoma, Texas, U-
tah, California, the Philippines, Australia, end French Indo-
china. The plant seems to be a hybrid, but its parentage is not
certain. Verbena officinalis L. is probably one parent end the
other may be V. robusta Greene, as both these species are known
from Amador County.

VERBENA DISSECTA f. ALBA Moldenke, f. nov.

Haec forma a forma typica speciei corollis albis recedit.

This form differs from the typical form of the species in
having white corollas.

The type was collected by Santiago Venturi (no. 7068) ina
prado at Balcozna, dept. Del Alto, Catamarca, Argentina, at an
altitude of 1250 m., on January 18, 1928, and is deposited in
the herbarium of the California Academy of Sciences.

VERBENA DUSENII Moldenke, sp. nov.

Herba mediocriter alta recta; caulibus ramisque argute tet-
ragonis minute puberulis vel dein glabrescentibus sarmentosis,
apicem versus purpureis; petiolis grecilibus marginatis minute
parceque puberulis; laminis ovatis leviter chartaceis trifido-
dissectis utringe parcissime minuteque strigilloso-puberulis,
lobis irregulariter incisis acutis.

Apparently a rather tall erect herb; stems and branches
sharply tetragonal, minutely puberulent or eventually glabres-
cent on the larger parts, the upper portions often decidedly
purplish, twiggy; principal internodes 2--9.5 cm. long; nodes
annulate; leaves rather abundant, decussate-opposite, often with
several smaller ones in their ax‘ls; petioles slender, 3--7 mm.
long, margined and on the larger leaves often not distinct from
the blade, minutely and sparsely puberulent, especially on the
margins; blades ovate in outline, thin-chartaceous, somewhat
lighter green beneath, 1.5--%3.5 cm. long, 1--3.3 cm. wide, tri-
fid-dissected, the lobes irregularly incised, acute, the lower
ones widely divergent, very sparsely and minutely strigillose-
puberulent on both surfaces (mostly on the lamina above and on
the venetion beneath), the very slender midrib and secondaries
plane or subimpressed above, slightly prominulous beneath, a.
few short tertieries often also discernible beneath; inflores-
cence spicate, terminal and in the uppermost axils, subcapitate
flattened in anthesis, the floriferous portion later elongating
to 7 cm., dense, showy; peduncles rather stoutish, tetragonal,
saostly purplish, 6--10 cm. long, rather sparsely strigillose
with reflexed whitish hairs; bractlets conspicuous, green,
rather broadly elliptic, 5--8 mm. long, 2--4.2 mm. wide, acum-
inete at apex, glabrous except for the long-ciliete margins;

1948 Moldenke, New and noteworthy plants 423

calyx tubular, 9--10 mm. long, 1--1.5 mm. wide, S-costate, pur-
plish (especially on the ribs), densely white-hispidulous on
the ribs, less so in age, the 5 teeth caudate-apiculate, 1.5--2
mm. long, purple; corolla showy, its tube about 15 mm. long,

_ puberulent above the celyx, its Limb 10--13 mm. wide, very
sparsely puberulent on the outside, the lobes deeply cordate-
notched at the apex; anthers glandular-anpendaged.

Tne type of this very showy species was collected by Per
Karl Hjalmar Dusén (no. 710€) -- in whose nonor it is named --
in rather swaztpy ground at Pinhees, Faranaé, Brazil, on Cctober
29, 1908, ard is deposited in the herbarium of the Naturhistor-
iska Rikemuseum at Stockholm.

VERBENA EUMIFUSA var. RETICULATA Moldenke, var. nov.

Haec varietas a forma typica speciei recedit laminis folio-
rum ellipticis vel obovatis vel suborbicularibus acutis, ad
basin cuneatis vel subacuminatis, in petiolum aletum brevem an-
gustatis vel subsessilibus, crasse dentatis supra pustulato-
bulletis et albo-pilosis (pilis adpressis antrorsis), subtus
densiuscule patento-nirsutulis.

This variety differs from the typical form of the species in
having its leaf-bledes varying from elliptic to obovate or sub-
orbicular, 0./--3 cm. long, 0.€--1.5 cm. wide, usually acute at
apex (rounded on the smallest leaves), cuneate or subecuminate
at base, narrowed into a very snort winged petiole or subses-
sile, coarsely dentate from the widest part to the apex with
acuts or obtuse rether reguler teeth, not lobed, pustulate-
bullete above and rather abundantly hairy with white appressed
entrorse hairs, rether densely spreading-hirsutulous on the ve-
nation beneath; the midrib, secondaries, and veinlet reticula-
tion deeply impressed above and very uniformly prominent be-
neath.

The type of this variety was collected by Fer Karl Hjalmar
Dusen (no. 15714) on the campo, altitude 800 m., between Lago.
and Desiro Ribas, Parana, Brazil, on October 22, 1914, and is.
deposited in the herbarium of the Naturhistoriska Riksmuseum at
Stockholm.

VERBENA LOBATA var. HIRSUTA Moldenke, var. nov.

Haec varietas a forna typica speciei ramis ramulisque netio-
lisque pedunculisque bracteolisque calicibusque et pagina infer
iora leninorum densissime albo- vel flavescento-hirsutis recedit

Tnais variety differs from the typical form of the species in
having the branches, branchlets, and twigs, as well as the pet-
loles, lower leaf-surfaces, peduncles, bractlets, and calyxes
very densely hirsute with widely spreading white or flavescent
hairs. The upper leaf-surface is also mors hirsute than in the
typical form.

The type of this variety was collected by Gustef Cskar An-

4o4 PHY TODOS TA Vol. 2, no. 10

dersson Malme (no. 1260) in the grassy edges of a marsh et Pin-
hal, near Santa Maria, Rio Grande do Sul, 8razil, on January
27, 1902, and is deposited in the herbarium of the Naturhistor-
iska Riksmuseum at Stockholm. The collector describes the cor-
olla as blue.

VERBENA MACDOUGALII f. ALBIFLORA Moldenke, f. nov.

Haec forma a forma typica speciel corollis albis recedit.

This form differs from the typical form of the species in
naving white corollas.

The tyre was collected by Francis Ramaley (no. 16847) at
Alamosa Canyon, Alamosa County, Colorado, at an altitude of
8000 feet, on September 14, 1938, and is deposited as sheet no.
42339 in the herbarium of the University of Colorado at Boul-
der. It was incorrectly labelled as Verbena stricta Vent.

VERBENA MORICOLOR Moldenke, sp. nov.

Herba; caulibus ramisque grecilibus tetrazonis perce irregu-
lariterque albido-pilosis, juvantute densiuscule hirsutulis;
petiolis irregulariter brevissimeque pilosis et parce hirsutu-
lis; laninis leviter chartaceis lanceoletis attenuato-acutis,
ad basin obtusis vel abrupte acutis, irregulariter dentatis,
supra densiuscule strigoso-pubescentibus, subtus dense pubes-
centibus; corolla purpurea.

Herb, to 1 m. tall; stems and branches slender, more or less
tetragonal, sparsely and irregularly pilose with whitish hairs
ef various length, the youngest parts rather densely hirsutu-
lous with sharp-peinted hairs about 1 mm. long borne at right
angles to the branch; nodes annulate; principal internodes e-
longated, 7.5--13 cm. long; leaves decussate-opposite; petioles
slender, 1.5--2 cm. long, irregularly pilose with very short
hairs and sparsely hirsutulous with ‘ong sharp-pointed hairs
like on the younger branches; blades thin-chartaceous, somewhat
lighter beneath, lanceolate, 4--8 cm. long, 1--2.3 cm. wide,
attenuate-acute at the apex, obtuse or abruptly acute et the
base, irresularly dentate from base to apex with rather broad
subacute teeth, the larger irrezularly interspersed among small-
er onss toward the base of the blade and often doubly dentate,
rather densely strigose-pubescent above, densely pubescent be-
neath with hairs of various lengths; midrib slender, plane or
slightly subimpressed above, slightly prominulous beneath; sec-
ondaries about 7 pairs, very slender, ascending, hardly arcu-
ate, indiscernible above, rather obscure beneath; veinlet re-
ticuletion indiscernible; inflorescence terninal, spicate, the
flowering portion apparently elongating to about 7 cm.; pedun-
cles similar to the upper branches or stems in texture, shape,
and pubescence, 1--7.5 em. long; rachis rather densely pilose-
pubescent with hairs of various lengths; bractlets lanceolate,
about 4 mm. long, attenuate to the apex, sparsely puberulent on

Sra

1948 Moldenke, New and noteworthy plants 425,

the back, the margins regularly ciliate; calyx cylindric, 6--7
mm. long, ebout 1 mm. in diameter, rether sparsely puberulent-
pilosulous, the teeth 0.5 and 0.9 mm. long, subulate; corolla
hypocrateriform, purple, its narrow-cylindric tube 13--14 mm.
long, densely pilose-puberulent above the calyx, its limb 6--7
mm. wide, much darker in the throat, glabrous within, subglab-
rate outside.

The type of this handsome species was collected by Santiago
Venturi (no. 5397) among spiny plants on hillslopes at Sterra
de Calilegua, altitude 800 m., dept. Ledesma, Jujuy, Argentina,
on October 11, 1927, and is deposited in the herbarium of the
California Academy of Sciences at San Francisco. The species is
obviously related to V. phloziflora Cham. and V. incisa Hook.,
but differs conspicuously in its much smaller calyx and corolla.

VSRBENA REGNELLIANA Moldenke, sp. nov.

Herbea; caulibus saepe decumbentibus radicantibus; ramis ad-
scendentibus vel rectis obtuse tetragonis brunnescentibus saepe
submarzinatis parce pilosis vel glabrescentibus; foliis numero-
sis brunnescentibus; petiolis gracilibus valde alatis parce
pilosis; laminis leviter chartaceis ovatis irregulariter incis-
is vel trifidis supra percissime pustulato-pilosis (pilis al-
bidis brevissimis adpressis), subtus parce adpresso-pilosulis,
lobis acutis vel apiculatis subrevolutis.

Herb; stems often decumbent and rooting eat the nodes;
branches ascending or erect, obtusely tetragonal, brunnescent
in drying, often slightly margined, sparsely pilose or glabres-
cent; principal internodes 1--7.5 cm. long; nodes annulate;
leaves abundant, decussate-opposite, often with a cluster of
small ones in their axils, brunnescent in drying; petioles
slender, 2--5 mm. long, plainly winged, sparsely pilose; blades
thin-chartaceous, darker above than beneath, ovate in outline,
1.5--4.5 em. long, 0.8--2.6 om. wide, irregularly incised, the
larger trifid, the lobes often with ea single exterior divergent
tooth, acute or apiculate, very sparsely pustulate-pilose above
with extremely short appressed whitish hairs, sparsely aprress-
ed-pilosulous (but not pustulate) on the larger venation and on
the margins beneath, the margins usually very slightly revo-
lute; venation mostly indiscernible or obscure above, the mid-
rib and very slender secondaries conspicuously prominulous be-
neath, but the veinlet reticulation indiscernible; inflores-
cence spicate, long-pedunculate, terminal and also exillary in
the uppermost axils, the floriferous portion apparently short
and dense or elongating to about 4 om. after anthesis; pedun-
cles very slender, brunnescent, 3--7.5 cm. long, very sparsely
pilosulous; brectlets ovate-lanceolate, about 3 mm. long and 1
mm. wide, sharply attenuate or subacuminate at apex, subglab-
rate except for the ciliate margins and sometimes e very few
microscopic hairs at the apex; calyx tubular, about 4 mm. long

426 Pry Pe oe tes Vol. 2, no. 10

and 1.3 mm. wide, very sparsely and minutely pilosulous, the
teeth about 0.5 mm. long; corolla-tube 5--6 mm. long, sparsely
and very minutely pilosulous above the calyx; corolla-limb
about 4 mm. wide, minutely pilosulous on the outer surface.

The type of this distinctive species was collected on the
swampy banks of the rapidly-flowing Rio Verde at Caldas, Mines
,Geraes, Brazil, on November 20, 1867, by Anders Frederik
Regnell (no. III.939) -- in whose honor it is named -- and is
deposited in the herbarium of the Naturhistoriska Riksmuseum at
Stockholm.

VERBENA SPECTABILIS Moldenke, sp. nov.

Herba; caulibus brevibus obtuse tetragonis brachistis dens-
fuscule breviterque brunneo-pubescentibus, pilis reflexis; ram-
is gracilibus rectis vel adscendentibus obtusiuscule tetragonis
plerumque’ sulcatis dense breviterque sordido-pubescentibus,
pilis reflexis; petiolis plerumque obsoletis; laminis chartace-
is ovatis acutis, ad basin angustatis, irregulariter dentatis
vel subincisis, supra densiuscule adpresso-strigosis, subtus
adpresso-pubescentibus; bracteolis lanceolatis.

Herb, about 45 cm. tall; stems rather short, obtusely tet-
ragonal, mostly branched at or near the base, rather densely
short-pubescent with reflexed brownish hairs; branches slender,
erect or ascending, rather obtusely tetragonal, usually sulcate
between the angles, densely short-pubescent with reflexed sord-
id-gray heirs; nodes annulate; principal internodes 2--4 cm.
long; leaves decussate-opposite; petioles very snort or usually
obsolete; blades chartaceous, rather uniformly green on both
surfaces, ovate, 2.5--4 cm. long, 0.8--2 cm. wide, acute at a-
pex, acuminetely narrowed into the broadly winged petiole at
base, irregularly dentate or occasionally subincised from the
apex to the widest point, the lowest teeth sometimes lobe-like
and divergent on larger leaves, rather densely appressed-
strigose above, appressed short-pubescent beneath; midrib and
the 5--8 pairs of secondaries very slender, usually impressed
above, prominulous beneath, the secondaries rather straight,
ascending, branching at the apex and a branch extending to the
tip of each tooth; inflorescence terminal and in the uppermost
leaf-axils, congested-spicate, the floriferous portion 2.5--3.5
cme long, densely many-flowered, apparently not elongating
after anthesis; peduncles slender, 4--€.5 em. long, densely
short-pubescent with reflexed whitisn hairs, often jointed at
about the midpoint and there bearing a pair of lanceolate
bracts 7--5 mn. long; floral bractlets lenceolate, about 4 mn.
long, about 1 mm. wide at the base (or narrower),ettenuate et
the apex, closely sppressed to the calyx, densely short-pubes-
cent with spreading whitish hairs; celyx cylindric, about 9 mm.
long, S-costate, densely short-pubescent with spreading whitish
hairs, its rim unequally earistate-toothed, purplish, the longer

1948 Moldenke, New and noteworthy plants 427

appendages about 1 mm. long, often twisted-connivent before and
efter anthesis; corolle purple, showy, its tube 10--15 mn.
long, rether densely puberulent above the calyx outside, its
limb about 9 mm. wide.

The type of this handsome species wes collected by rik
Leonard Ekman (no. 1980) along the small stream called ‘agdele-
na et Loreto, Fosedas, Misiones, Argentina, on February 6,
1908, end is deposited in the herbarium of the Naturhistoriske
Riksmuseum at Stockholm.

VERBENA SWIFTIANA Moldenke, sp. nov.

Herba pumile gracilis; ceulibus gracilibus saepe semi-
procumbentibus et radicantibus, acute tetregonis glabris brun-
nescentibus; remis numerosis saepe simplicibus rectis grecilli-
mis acute tetragonis glabris brunnescentibus; foliis numerosis
membranaceis utringue uniforme brunnescentibus; petiolis grac-
{llimis glabris, vel alatis obscuris; laminis linearibus vel
angustissime ellipticis et integris vel ovatis et trifidis
_utrinque glabris, lobis terminelibus integris vel paucidenta-
tis vel incisis.

Low slender herb to about 3 dm. tall; stems slender, often
partly procumbent and rooting at the nodes, acutely tetragonal,
glabrous, brunnescent; brenches numerous, often simple or near-
ly so, erect, very slender, acutely tetragonal, glabrous, brun-
nescent; nodes annulete; principal internodes 1--2 cm. long;
leaves numerous, membranous, uniformly brunnescent on both sur-
faces in drying, variable in shape; petioles very slender and
1--45 mm. long, glabrous, or obscure and winged, merging into
the blade; blades 1--2 cm. long, varying from linear or very
narrowly elliptic end entire to ovate and trifid with the large
terminal lobe entire or several-toothed or -incised, glabrous
on both surfaces; midrib very tenuous, mostly indiscernible a-
bove, very slightly proninulous beneath; secondaries and vein-
lets indiscernible on both surfaces; inflorescence terminal,
spicate, rather few-flowered, to about 5 cm. long, the flowers
rather distant during and after anthesis; peduncles slender,
exactly similar to the branches in color and texture, 1--1.5
cm. long; rechis exactly similar to the peduncle; bractlets
lanceolate, about 1 mm. long and O.5 mm. wide, acuminate at the
apex, ligntly strigillose; calyx tubular, about 1.5 mm. long
and O.75 mm. wide, lightly strigillose outside, the rim very
shortly toothed; corolla-tube very slender, sbout 3 mm. long,
very sparsely and minutely strigillose toward the apex; corolla
rose, its limb about 5 mm. wide, very lightly and minutely
strigillose toward the base outside.

The type of this species was collected by G. J. Schwarz (no.
3402) at Corpus, dept. San Ignacio, Misiones, Argentina, on
September 19, 1946, and is deposited in the herbarium of the
Naturhistoriska Riksmuseum et Stockholm. The species is named

428 PHY TCECe DA Vol. 2, no. 10

{in honor of Josiah Otis Swift (1870-- ), well-known naturalist,
founder of the Yosian Brotherhood which in the past auarter of
a century has brought the beauties of the great out-of-doors to
over 150,000 Americans, chiefly in the vicinity of New York
City, and author of the daily column "News Outside the Door"
which has popularized Nature-study among millions of newspaper
readers.

NOTES ON SOUTH AMERICAN MZLASTOMES

H. A. Gleason

The genus Centradenia, with seven known species, has always
been considered endemic to Central America, ranging from
Fanama to. southern Mexico. Haught has recently collected
plants along the western coast of Colombia which, although
bearing only immature flowers, undoubtedly represent C.
Maxoniana Gl. So far as known to me, this is the first collec-
tion from the South American continent, and leaves Heterocen-
tron the only endemic genus of the family in Central America.

Haught has also collected in Colombia plants of Miconia
barbicaulis Gl. which exactly match the type specimen. The spe-
cies was originally described from the eastern slope of the
Andes in Scuador. Heugnt's collection was made on the western
slope of the mountains and approximately four hundred miles
north of the type locelity.

CALYFTRELLA STELLATA Gl., sp. nov. A speciebus sex differt
petalis subrotundis nec acutis nec acuminatis; a C. littorali
Gl. differt floribus C-meris magnis foliis S-nerviis; a C.
denticulata Gl. differt foliis et hypanthiis stellato-
tomentosis atque longe villosis, floribus majoribus, dentibus
calycis exterioribus multo majoribus 2.5 mm. longis.

A large shrub or smell tree with clear red flowers. Younger
stem densely villous with short crowded hairs. Leaves obovete-
oblong, 5S-nerved, entire, abruptly short-acuminate, obtuse or
rounded at base, glabrous above, densely white-stellate-tomen-
tose beneath and also villous with pale brown hairs. Hypanth-
ium densely stellate and also villous, 8 mm. long to the torus.
Calyx 4.5--5 mm. long, irregularly ruptured at anthesis, usual-
ly into 3 lobes, pubescent like the hypanthium but with shorter
hair; exterior teeth triangular, 2.5 mm. long. Petals rotund-
obovate, 14 mm. long and wide. Filaments 8.5 mm. long; anthers
11.6 mm. long, tangentially flattened, the stout connective
prolonged 2 mm. to the filament and terminating in an obscure

1948 Gleason, South American Melastomes 429

obtuse basel sour.

Type, Espinosa 1544, in the herberium of the New York Boten-
ical Garden, collected at Huiaco, nesr Loja, Ecuador, alt.
2250 meters.

A key to the eight species of Calyptrella then known wes
published in Phytologia 2: 301 in 1947, Cur plant traces
through the key directly to C. denticulata, aleo from icusdor,
but differs from it conspicuously in certain characters not
mentioned in the key. In C. denticulata, the pubescence of the
leaves and the hypanthium is restricted to stellate hairs,
lacking the long simple ones, the leaves are nerrowed to the
base, the hypanthium and calyx are only half as large, the ex-
terior teeth are merely minute points, the petals and stamens
sre considerably smaller.

CONOSTEGIA HAUGHTII Gl., sp. nov. Frutex 1.5 m. altus, ramis
foliisque juvenilibus arctissime furfuraceis mox glabrescenti-
bus. Folia membranacea, elliptico-oblanceolata vel oblanceolata,
usque ad 22 cm. longa 7.5 cm. lata, acuminata ad epicem obtusun,
integra, basi longe cuneata fere ad caulem, petiolo libero
tantum 2--3 mm. longo, valde S-pli-nervia, subtus pallida. Pan-
icula laxe pauciflora, 4--6 cm. longa. Flores S-meri in cymulis
5-floris terminalibus. Alabastrum obovoideum, obtusum, 4.5 mm.
longum, tenuissime furfuraceum; hypanthium 2.2 mm. longum.
Petala alba, ovata, obtusa, equilatera, 3.2 mm. longa, 1.9 mm.
lata. Stamina 10, isomorpha; filamenta leviter complanata, 1.5
mm. longa; antherae oblongae, 1.5 mm. longee; connectivum sim-
plex. Ovarium inferum; stylus gracilis, 4.7 mm. longus; stigma-
te fere punctiformi.

Type, Haught 4939, collected at Quebrado Isaias, east of
Turbo, Dep't. Antioquia, Colombia, altitude about 50 meters, in
the herbarium of the New York Botanical Garden. The species
stends out sharply from the other Colombian plents of the genus
by its apparently glabrous foliage, strongly 5-pli-veined
leaves, and small few-flowered cymes.

MICONIA MEDUSA Gl., sp. nov. Sect. Cremanium. Frutex; rami
graciles, petioli, et foliorum pegina inferiore tomentosi, pil-
is elongatis contortis parce ramosis. Folia longe petiolata,
membranecea, oblongo-oblanceolata, breviter acuminsta,
spinuloso-ciliata (dentibus adscendentibus, 0.5 mm. longis), 3-
nervia, supre glabra, Fenicula terminalis, valde reducta, non
vel vix ramosa, 2--5 cm. lonz@e- Flores S-meri ad nodos sessiles
fasciculati. Hypenthium poculiforme, glabrum. Calyx ad anthesin
in lobos 5 triangulares hyalinos 0.4--0.5 mm. longos ruptus;
dentes exteriores trianguleares, erecti, C.2 mn. longi. Fetala
alba, fere orbicularia, 1.25 mm. longa. Filamenta 1.6 mm.
longa, ultra medium geniculata. Antherae oblongae, 0.8 mm.
longae, poris 2 latis ventro-terminalibus dehiscentes. Cvarium

430 PHYTOLOGIA Vol. 2, no. 10

inferum, (?) 3-loculare. Stylus rectus, 3 mm. longus, apicem
versus clavatus ad stigma rotundatun.

Type, Espinosa 1559, collected 5 km. southeast of Loja, Zcu-
ador, alt. 2300--2400 meters, in the herbarium of the New York
Botanical Garden.

There is no doubt that this plant finds its nearest relative
in the Feruvian M. aprica Gl., in which the minute flowers are
similarly congested and the pubescence is irregulerly branched.
It differs from our plant in its much wider end thicker leaves,
scabrous above, with shorter end sparser tomentum and larger
spinulose teeth, in the well developed branched panicle, the
much larger anthers, and the prolonged connective.

PACHYDESMIA Gl., gen. nov.

Among the Haught collection is another melastome which dis-
plays such remarkable differences from all other members of the
family known to me thet I am constrained to describe it as a
new genus.

Caulis circum basin petioli valde expansus. Panicula trich-
otoma, axi centrali non evoluto et panicula corymbiformi.
Ovarii summum ad parietam hypanthii connexum per membranas 10
redietim divergentas. Connectivum antherae valde dilatatum et
thecas ubique excedens, sursum in lobum unum rotundatum vel
truncatum, deorsum in lobos 2 rotundatos. Thecae antherae
parvae, in medio connectivi affixeae et vix elevatae, sursum in
partem sterilem divergentem breviter productae, late poculi-
forme, poro magno terminali dehiscentem.

FACHYDESMIA HAUGHTII Gl., sp. nov. Frutex 1 m. altus, caule
ramoso, sparse furfuraceo. Fetioli 6--12 mm. longi. Laminae
elliptico-ovatae, usque ad 120 mm. longae 57 mm. latae, subac-
uminatae, ciliatee (ciliis spinulosis 1--2 mm. longis), basi
late rotundatae, 3-nerviae, supra glabrae, subtus sparsissime
furfuraceae secus venas primerias. Penicula fere 1 dm. longa
lataque. Hypanthium late campanulatum, 1.5 mm. longum ad torun,
3 mm. in diametro, minutissime furfuraceum. Calycis tubus 0.5
mme productus, sinubus late rotundatis; lobi obtuse triangular-
es, 1 mm. longi, minute eroso-ciliati; dentes exteriores ad-
nati, trienguleres. Fetala alba, late elliptica, 4.3 mm. longa,
inequilatera. Stamina 10. Filamenta ce. 4 mm. longa, supra med-
ium geniculate, ad epicem valde dilatata. Connectivum 2 mm.
longum, in ser. staminum exteriore apice truncatum, in ser. in-
teriore ovatum obtusum. Thecae vix elevatum, poro divergente
C.8 mm. in diametro dehiscentes.

Type, Haught 5201, collected west of Tambo, on the west
slope of the Cordillera Occidental, Dep't. Cauca, Colombia, al-
titude 2200 meters, in wet forest.

The peculier features of the plant are summarized in the

1948 Gleason, South American Melastomes 431

generic diagnosis. The spathe-like expansion of the stem about
the base of the petioles is known also in four species of Mi-
conia, M. scutata, annularis, stipularis, and manicata. A cor-
ymbiform panicle
very rarely ap-
pears in the gen-
use The connection
of the ovary-
summit with the
wall of the hypen-
thium has not been
observed by me in
the Miconieae: it
consists of ten
tnin radiating
membranes extend-
ing from the very
summit of the o-
vary to the wall
of the hypanthium
as far as the tor-
us, and dividing
the cavity into as .
meny chambers each
of which is in the
bud occupied by an
anther. The re-
merkebly large
connective extends
well below the
minute thecee,
which is common in
meny species of

“elastomes, and Front and side view of anther; hypanthiunm,

also well beyond calyx, and style. x 8.3
the thecae, a con-

dition which I
have not noted elsewhere in the family. The two thecae are
small and tangentially flattened. Before dehiscence, as seen in
an unopened flower-bud, they are slightly distended; after de-
hiscence they are scarcely eleveted above the surface of the
connective. At the upper end of the thecae the broadly cup-
shaped sterile portion diverges at right angles from the con-
nective and terminates in s large pore. The total length of the
thecae, including the pore, is about half that of the connect-
ive.

In general aspect and in most technical characters the plant
suggests Miconia, to which it must be related, but it differs
so greatly in its anthers from every species known to ma that

432 PHYTOLOGIA Vol. 2, no. 10

it can not be associated with any of them, nor assigned to any
ot the eleven sections of the genus.

A NEW SPECIES OF PTYCHCCARPUS FROM PSRU

Joseph V. Monachino

The genus Ptychocarpus Kuhlmann belongs in the tribe
Casearieae (Benth. & Hook. Gen. Pl. 1: 795; ingl. & Prantl
PPlanzenf. 3©2:46).It is easily distinguished at sight from
other American genera in the Flacourtiaceae by its inflores-
cence habit. P. apodanthus, the only hitherto known species,
was reported as seemingly frequent in the state of Para, Brazil;
it has also been collected in the Matto Grosso (Krukoff 1342;
Tabe jaza,, upper Machado River, terra firma; November 13, 1941;
distributed as "Ferebea ?"). The simple, clearly defined posi-
tion of this genus end species made it an easy matter to recog-
nize the following novelty, which was chanced upon among a set
of unidentified specimens filed as Sapotaceae at the New York
Botanical Garden.

PTYCHOCARPUS KILILIPII Monachino, sp. nov. Arbor glaber 6--8
m. altus; petiolis 4--6 mm. longis; leminis ca. 15--20 em. lon-
gis et 4--7 cm. latis punctatis oblongo-oblanceolatis, ad api-
cem cuspidato-acuminatis, ad basin acutis, nerviis primeriis
lateralibus utroque ca. 12 distantibus adscendentibus arcuatis;
inflorescentiis axillaribus sessilibus glomeratis ca. 8 mm.
longis paucifloris dense bracteatis; floribus masculis: perian-
thio urceolato ca. 6 mn. longo intus prope basin dense piloso,
caeterum glabro, lobis 4 imbricatis ovetis ca. 1.5 mm. longis;
staminibue 8, filamentis ligulatis ca. 1.5 mm. longis et 0.5
mm. latis.

Glabrous tree 6--8 m. tall; stipules caducous; leaves alter-
nate, the petioles 4--6 mm. long, thick, the blades about 15--
20 cm. long and 4--7 cm. broad, closely translucent-punctate,
oblong-oblanceolate, abruntly cuspidate-acuminate, nerrowed and
acute at base, faintly serrulate on the upper margins, the
principal lateral nerves about 12 pairs, widely spaced end
clearly ascending-arcuate, connected near the margins of the
blade, the reticulation loose, a little raised on the underside
of the blade; inflorescences closely resembling those of Pp.
apodanthus, axillary, sessile, glomerate, about 8 mm. long,
few-flowered, densely bracteate, the bracts overlapping so ag
to impart a strobile-like appearance to the inflorescences, or-
bicular to oblong, up to 6 mn. long and 3 m. broad, flet to

1948 Monachino, New Species of Ptychocarpus 433

cucullate, rounded at the apex, punctate; only male flowers
seen, the perianth urceolate, about 6 mn. long, densely pilose
near the base within, otherwise glabrous, punctate, the lobes
4, imbricate, ovate, about 1.5 mm. long and as broad; stamens
8, the filaments strap-shaped, flat, inserted at about the mid-
dle of the perianth-tube, alternately longer and shorter, those
opposite the perianth-lobes twice as long, about 1.5 mm. long
and 0.5 mm. broad, the anthers oblong, about 1 mm. long,
reaching the throat of the perianth; rudimentary ovary as in P.
apodanthus.

Type: &. P. Killip & A. ©. Smith 29936, collected in dense
forest, alt. about 100 m., at “ishuyacu, near Iquitos, dept.
loreto, Peru, September 24--28, 1929; deposited in the Britton
Herbarium at the New York Botanical Gerden.

F. Killipii is easily distinguished from the only other
known species in the genus, P. apodanthus Kuhlmann. The princi-
pal lateral nerves of the leaves are about half as many and
ascending-arcuate, not spreading and straight. Tne indumentum
within the perianth-tube is denser than that of P. eapodanthus.
The filaments are strap-shaped, and those alternate with the
perianth-lobes ere manifest, although about half the length of
the longer ones; they are inserted at about the middle of the
perianth-tube. The filaments opposite the perianth-lobes in FP,
avodanthus are broadly deltoid, and tne alternate ones hardly
apparent; they are inserted at the throst of the perianth-tube.

ADDITIONAL NOTSS ON THE GENUS AEGIFHILA. IX

Herold N. Moldenke

AEGIPHILA BRACHIATA Vell.

The species has been collected in low woods among fields at
an altitude of 5O meters in a region where tne average annual
rainfall is 1.5 m. and the average temperature varies from <i
to 35° C. during the year. It has been collected in anthesis in
February and October, and has been confused with Citherexylum
by some herbarium workers.

Additional citations: BRAZII: Rio Grande do Sul: Friedrichs
32928 (N); Henz 32936 (S); Rambo 990 (N). State undetermined :
Sellow 1269 [Macbride photos 17590] (N--photo).

AEGIFHITA CANDELABRUM Briq.
Additional citations: PARAGUAY: Hassler 8120 [Macbride
photos 24621] (N--photo of type).

434 PHYTO L OoGtok Vol. 2, no. 10

AZGIPHILA CHRYSANTHA Hayek

Additional citations: ECUADOR: Guayas: Eggers 14348 [Mec-
bride photo 20349] (N--photo). PERU: Loreto: Poeppig 2314
[Macbride photo 34313] (N--photo of logotype).

AEGIFHILA CONTURBATA Moldenke
Additional citations: BRAZIL: Maranhaéo: Newman sen. [Mac-
bride photo 28377] (N--photo of type).

AEGIPHILA CORDATA Poepp.
Additional citations: PERU: Loreto: Poeppig 2158 [Macbride
photo 34312] (N--photo of type).

AEGIFHILA CORDIFOLIA (Rufz & Fav.) Moldenke
Additional citations: PERU: Hudnuco: Rufz & Pavon sen.
[Mifia, Panatahua; Macbride photo 28378] (N--photo of isotype).

AEGIPHILA COSTARICENSIS Moldenke
Additional citations: MEXICO: Chiapas: Matuda 572 (Mh), 2101
(Mh).

AEGIPHILA CRENATA Moldenke
Additional citations: BRAZIL: Parana: Dusen 10541 [Macbride
photo 30182] (N--photo of isotype).

AEGIPHILA DEPPZANA Steud.

Additional citations: MEXICO: Chiapas: Seler & Seler 2005
(Gg--245897). JUANA RAMIRZZ ISLAND: &. Palmer 464 (Gg--34499).
ens Magdalena: Balbis sen. (Macbride photo 33932] (N--
photo). :

AEGIPHILA ELATA Sw.

Additional citations: MEXICO: Tabasco: Matuda 3031 (Mh),
a (Mh), ae (Mh). BRITISH HONDURAS: Gentle 2633 (Mh), 2843
Mh), 3350 (Mh); Schipp 216 (Gg--172812). JAMAICA: W. Harris

11746 (Gg--31928); Swartz s.n. (S--isotype).

A=SGIPHILA FALCATA Donn. Sm.

Two vernacular names not. before recorded for this species
are reported by Matuda from Chiapas. They are "taco" and
"taquito".

Additional citations: MEXICO: Chiapas: Matuda 666 (Mh).

oe fi Tiguesata: E. Wall sen. [21/4/28] (Ew), sen. (24/4/
2 Ew).

ASGIPHILA FERRUGINEA Hayek & Spruce

Additional citations: ECUADOR: Imbabura: Lehmann 4700 [Mac-
bride photo 17584] (N--photo). Pichincha: Spruce 5473 [Macbride
photo 34311] (N--photo of type).

bracteolosa Moldenke

436 PHYTOLOGIA Vol. 2, no. 10

AZSGITFHiLA FITIPESS Mart. & Scheu.

Haught describes this plant as a slender shrub with cream-
colored flowers.

Additionel citations: COLOMBIA: Cundinamarca: Haught 6124
(N). BRAZIL: Fara: “Martius s.n. [Macbride photo 20350] (N--
photo of cotype).

AEGIPHILA FLORIBUNDA Moritz & Moldenke
Additional citations: VENEZUZLA: Aragua: Moritz 1765 [Mac-
bride photo 34310] (N--photo).

AEGIFHILA FOSTIDA Sw.
Additional citations: JAMAICA: Swartz sen. (S--isotype).

AZGIFHILA GLANDULIFERA Moldenke
Romero C. describes the plant as a shrub 6 meters tall.
Additional citetions: COLOMBIA: Choco: Romero C. 503 (N).

ASGTFHILA GRANDIS Moldenke
Additional citations: COLOMBIA: Cundinamarca: Triana 2080
{Macbride photo 28379] (N--photo).

AEGIFHILA GUIANENSIS Moldenke
Additional citations: COLOfBIA: Cundinamarca: Triana 2084
[Macbride photo 28380] (N--photo).

AEGIFHILA HASSLERI Bria.

Additional citations: BRAZIL: Rio Grande do Sul: Buck
31389 (N). PARAGUAY: Hessler 3193 [Macbride photo 24613] (N--
photo of cotype). URUGUAY: Fuques e.n. [Lombardo 4153] (N).

AEGIPHILA HERZOGII Molderke
Additional citetions: BOLIVIA: Santa Cruz: Herzog 1369 [Mac-
bride photo 2€381] (N--photo of isotype).

ASGIFHILA INTSGRIFOLIA (Jacq.) Jacks.
Sandeman describes the flowers of this species as cream-
colored.

Additional citations: PERU: Cuzco: Sandeman 3676 (K). BOLIV-
TA: Sante Cruz: Peredo sen. [21-III-1946] (N).

AEGIPHILA INTSRMEDIA Moldenke
Additional citations: BRAZIL: Maranhaéo: Herb. Gen. Mus.
Para. 2270 (Macbride photo 28382] (N--photo of isotype).

AZGIPHILA LANATA Moldenke
Additional citations: BRAZIL: Goyaz: Glaziou 21917 [Macbride
photo 28383] (N--photo of isotype).

qy

948 Moldenke, Notes on Aegiphila- 427

~

y ‘s
oN
IS Se SN
Fi

8: .
«
2G, 1 Meee
ZEON) Psy;
a A Y

(= Sal? CM
ZZ de
Wid) LAWET, if
GL jf

CALA

AMMA LEN ol
PLE RAL GIL MOH
VEAL] aU
( ny Ga f CM I)
of eGy

At Ayr
A

38 | 7 MMA ui

de Ale: 23 gg oe)

4 s SED GEG

¢ y; OES MCLEE
CTE age

Safe

ROG ACA See
Le CUT HEE ee

Fig. 2. Aegiphila gloriosa Moldenke

438 PO Yet O Deo Gt Vol. 2, no. 10

AEGIPHILA LAXIFLORA Benth.
Additional citations: BRITISH GUIANA: M. R. Schomburgk 772
[Macbride photo 28384] (N--photo of isotype).

AEGIPHILA LEHMANNII Moldenke e
Additional citations: COLOMBIA: Choco: Triana 2083, in part
{[Macbride photo 28385] (N--photo).

AEGIPHILA LHOTZKIANA Cham.

Additional citations: BRAZIL: Bahia: Casaretto 2022 [Mac-
bride photo 24614] (F--686352, F--772034--photo, Kr--photo, N--
photo). Minas Geraes: Heringer sen. [Herb. Esc. Sup. Agr. Lav-
ras 274; Herb. Dept. Bot. Est. S. Paulo 42456] (N); Markgraf
3281 [Brade & Mello Barreto 12140; Herb. Jard. Bot. Belo Hori-
zonte 28444] (F--1009600); Mello Barreto 3270 (Herb. Jard. Bot.
Belo Horizonte 11249; Herb. Rio de Jan. 32273] (F--933076, Ja),
9347 [Herb. Jard. Bot. Belo Horizonte 25608] (F--948145), 9736

Herb. Jard. Bot. Belo Horizonte 25641] (F--948135), 9796
(Herb. Jard. Bot. Belo Horizonte 25515] (F--948143); Sempaio
Pa (Herb. Rio de Jan. 32270] (Ja). Parané: Jonsson 10288 (F--

4735). S&o Paulo: Zagatto s.n. [Herb. Inst. Agron. Est. S.
Paulo 2469;. Herb. Dept. Bot. Est. S. Paulo 40171] (Sp). State
undetermined: Herb. Rio de Jan. 32272 (Ja); Sellow an. [Bras-
ilia; Macbride photo 17585) (F--663064--photo, Kr--photo, N--
photo, Vt).

AEGIPHILA LONGIFOLIA Turcz.

Cuatrecasas describes tnis plant as a scandent shrub,
blooming in November, inhabiting savannas at an altitude of 240
meters.

Additional citations: COLOMBIA: Meta: Cuetrecasas 7730 (Jc).
Santander Sur: Schlim 688 [Macbride photo 24616] (F--772036--
photo of isotype, Kr--photo of isotype, N--photo of isotype).

AEGIPHILA LUSCHNATHI Schau.

Synonymy: Aegiphila Luschnatii Schau. apud Hook. f. & Jacks.,
Ind. Kew. }: 46, sphalm. 1895; Aegiphila Luschnathii Schau.
apud Briq., Bull. Herb. Boiss., ser. 2, 4: 1167, sphalm. 1904.

This species has occasionally been confused in the herbar-
ium with A. laxiflora Benth.

Additional citations: BRAZIL: Rio de Janeiro: Guillemin s.n.
(St. Thérése 1839] (Du--166414); Luschneath s.n. (Herb. Martius
1040; Macbride photo 7879] (F--645715--photo of cotype, Kr--
photo of cotype, N--photo of cotype).

AEGIPHILA MACRANTHA Ducke

Additional items for the description: fruiting-calyx accres-
cent, woody, light-brown, 2.5--3.5 em. in diameter, 1.2--2 cm.
long, with a rough warty surface, otherwise glabrous, borne on .

Exe

~SSZayss
SAS

a

fia WV
ROR,

Wy Ss

Pines o Aegiphila obducta Vell.

440 PHYTOLOGIA Vol. 2, noe 10

a short pedicel; fruit broadly ovoid-ellipsoid, about z--4 om.
long and 2.4--3.4 cm. wide. "green, speckled with brown" (when
immature?), brown when dried, minutely asperous-granular, some-
times only 1 or 2 maturing in an inflorescence. It is a woody
climber found in mixed forests.

Additional citations: BRITISH GUIANA: De la Cruz 2836 (Cm);
Herb. Forest. Dept. Br. Guian. 3011 (S), 5295 [F.2550] (N);
Sandwith 1202 (s) BRAZIL: Pard: Ducke 843 (N).

AEGIPHILA MAGNIFICA Moldenke

This plant is said by Standley to be "rare" or "scerce". He
found it at an altitude of 700 m. in Guatemala, fruiting in -
March in wet thickets. The celyx and fruit are very similar to
those of A. paniculata. Baker describes it as a woody vine
climbing over trees in shaded places.

Additional citations: GUATEMALA: Escuintla: P. C. Standle
89280 (N). Suchitepéquez: P. C. Standley 62196 (F--982650).
NICARAGUA: Chinandega: C. F. Baker 204 (psa 76166. ae

Gg--31927=-isotype).

AEGIFHILA MARTINICENSIS Jacq.

References: Seymour, Host Ind. Fungi N. Am. 588--589. 1929;
Fittier, Supl. Plant. Usual. Venez. 54. 1939; Roig y Mesa,
Plant. Medic. Cuba 410--411 & 770. 1945.

An additional synonym is Aegiphyla martinicensis Jacq. ex
Moldenke, Suppl. List Invalid Names 1, in syn. (1941). Roig y
Mesa, on page 410 of the reference cited above, records the
common name "bois cobr{" from Martinique and Guadeloupe. He
quotes Gomez de la Maza to the effect that it is a diuretic
medicinal tea; a syrup made from it is used in the treatment of
asthma. Shafer reports the corollas 4 white or yellow The spec-
ies has been collected in anthesis in January, February, and
October, and in fruit in February and March. Holdridge found it
in brush pastures at an altitude of 300 feet in Fuerto Rico.
The Dudley Herb. 166413, labeled as this species, is ectually
Psychotria sulzneri Small. Cooper describes our plant as a tall
shrub or small tree, 10--15 feet tall, with creamy-white flow-
ers, and found it at altitudes of 1800 to 2000 feet on Dominica.
He records the common name “sylvania" from there.

Additional citations: PUERTO RICO: Holdridge 423 (N); Otero
308 (Bt--39906), 696 (Bt--52435). MONTSERRAT: ee 162 (Cm),
197 (Cm), 539 (Cm), 652 (Cm), 668 (Cm). DOMINICA: G. P. Cooper
60 (F--766225), 167 (Ca--549811, F--771385); Eggers 501 (Herb.
Prager 18669] (Gg, Gg--31926). MARTINIQUE: Bailey & Bailey 206
(Ba); Fonthieu s.n. (S). WEST INDIES (island iedvel eee
Swartz 9 (S). COLOMBIA: Norte de Santander: Quatrecasas 12845
(W--1851043). LOCALITY UNDESIGNATED: Collector undesig. 158 (Q).

AEGIFHILA MEDITERRANEA Vell.

1948 Moldenke, Notes on Aegiphila 441

SD)

v7 At
KG Ds CS
WGA

Fah

44a PHY To0. LO Gta Vol. 2, no. 10

References: Moldenke, Phytologia 1: 238 (1937) & 393. 1940;
Sampaio & Peckolt, Arquiv. Mus. Nac. Rio de Jan. 37: 334. 1943.
In the first of the references cited above I ventured the
opinion that the Rodriguez specimen there recorded was collect-
ed in the state of Rio de Janeiro, Brazil. It now appears that

the specimen actually was collected in Misiones, Argentina.
The plant was confused by the collector with Cordia Sellowiana
Cham.

The herbarium labels which accompany Macbride's photograph
no. 34358 are inscribed "Brazil Moldenke", implying that I col-
lected the plant there depicted. Actually it was collected by
George Gardner in 1836.

Additional citations: BRAZIL: Rio de Janeiro: G. Gardner 100
(Macbride photo 34358] (F--977197--photo, Kr--photo, N--photo).
ARGENTINA: Misiones: D. Rodriguez “87 (Herb. Inst. Miguel Lillo
57251] (N), s.n. [Lillo TObde} (G

AEGIFHILA MEDULLOSA Moldenke
Additional citations: BRAZIL: Rio de Janeiro: Saint-Hilaire
C.50 (F--977114--fragment of type).

AEGIPHILA MEMBRANACEA Turcz.

Williams describes the plant as a shrub 2 m. tall, with or-
ange fruit. His collection is a splendid fruiting specimen.
Triana's collection was made at an altitude of 450 m. and was
in anthesis in January. Steyermark'’s notes indicate a shrub 10
feet tall, with greenish-white corollas and filaments, grass-
green calyx and rachis, and membranaceous leaves which are
deep-green above and dull-green beneath. He found the plant in
rich damp forests, at an altitude of 1925--2075 m., blooming in
May. The label on the Chicago Natural History Museum's specimen
of Hostmann 89 is inscribed “French Guiana?" The collection was
made in Surinam.

Additional citations: COLOMBIA: Méta: Triana 3713, in part
(5] (Jc). VENZ ae Bolfvar: Ll. Williams 11200 (Ve). Mérida:
Steyermark 56379 (F--1205613, N). SURINAM: Hostmann 89 [Mac-
bride photo 2 ae (F--6333529--photo of isotype, F--686351--
fragment of isotype, F--77203l--photo of isotype, Kr--photo of
isotype, N--photo of isotype).

AEGIPHILA MOLLIS H.B.K.

An additional synonym is Aegiphila pubescens (H.B.K. ) Willd.
ex Moldenke, Suppl. List Invalid Names 1, in syn. (1941). All
the Bolivian material cited by me as this species in Brittonia
1: 406 (1934) and Phytologia 1: 239--240 (1937) & 394 (1940) is
probably A. Steinbachii Moldenke. The Steinbach 3168 cited in
these references is the type collection of A. Steinbachii. The
Woodson, Allen, & Seibert 1174 from Fanama and Dugand G. 639
from Colombia are anomalous in their extremely short pubescence

g 7
q\ V A WG,
Cares:
Y W4 \Y \4
“a Ss a.
> Siem y/ * aN
AEGAN /AS
Ay YS \\ \
\ \"
\
j,
es NX
\}\
\

\
Fig. 5. Aegiphila Valerii Standl.

dy ddy PHYTOLOGIA Vol. 2, no. 10

and may actually represent A. puberulenta Moldenke. H. H. Smith
870 in the University of California herbarium may be regarded
as typical of the long-pubescent A. mollis. The lias 581 and
Heught 2288, cited as A. mollis in Brittonia 1: Lo5 (1934) and
Fhytologia 1: 239 (1937) & 394 (1940) prove actually upon re-
examination to be A. puberulenta. The Goudot 2 cited in Fhyto-
logia 1: 239 as from an undetermined department of Colombia is
actually from either Bolfvar or Magdalena. It was collected at
"El Volador", and there is a "Volador" in each of these depart-
ments.

The original description of A. salutaris H.B.K. states that
the type was collected on the shorss of the Orinoco close to
Santo Thomas del Angostura. The Humboldt 983 cited by me in
Fhytologia 1: 283 (1938) and in the present contribution is al-
so labelled “Herb. Willdenow 2834" -- which is the herbarium
number givenfor the type collection -- but was collected at San
Carlos on the R8{o Negro, Amazonas, Venezuela.

Dugand G. collected the species at an altitude of only 60--
150 m. and describes it as a “shrubby small tree 4--5 m. tall"
with the stem 15 cm. in diameter at the base. OQuatrecasas des-
cribes it as a very leafy shrub with green branches and green-
ish-yellow or ochraceous corollas or a small tree to 6 m. tall,
growing at altitudes of 1400 to 1750 m., fruiting in March. He
hes found it along roedsides and in cultivated places. Haught
describes it as 4 scrambler to 4 m. tall. Steyernark notes the
corolla as pale-yellow, the rachis pale-green, and the leaves
membranaceous, deep-zgreen and reticulated above, gray-green
beneath, growing at altitudes of 1065 to 2430 m.

Additional citations: PANAMA: Chiriquf{: Woodson, Allen, &

Seibert 1174 (F--969465, N). COLOMBIA: Atlantico: Dugand G. 639
(F--744565). Cauca: Lehmann 6692 [Mecbride photo 17568] (Kr--
photo, N--photo). El Valle: Cuatrecasas 13869 (N), 14462 (N, 7
1652246). Cundinamarca: Bonplend sen. (F--976412); Guatrecesas

8287 (W--1774636); Triana 3713 (2; 300; 678] (Je). Magdalena:
Haught 3686 (W--1708528); H. H. Smith 868 (Cm, Vt)/ 870 (Ca--
58 908 , Cm, Te) ee aie ae Meta: Jaramillo, Mesa, Idrobo,
& Fernandez 332 (W--1900437). Tolima: Guatrecasas 10516 (W--
1796541). eat undetermined: Apolinar-Marfa 122 (F-~
1007353). VENEZUELA: Amazonas: Humboldt 983 (Herb. Willdenow
2834; Macbride photo 17588] (F--663067-- photo, Kr--photo, N--
photo). Federal District: Pittier 7855 (Du--297798, Gg--311279).
Guanico: Chardon 117 (W--1801748). Mérida: Steyermark 56314
(N). Miranda: T. Gonzdlez a.n. [Petere arriba] (Ve). State un-
determined: Humboldt XII [Herb. Willdenow 2838; Vecbride photo
17586] (F--663065--photo, Kr--photo, N--photo).

ASGIPHIIA MOLIIS var. INTSRMEDIA Moldenke
Zlias describes this plant as a tree 3--5 m. tall, with a
smooth trunk, dark berk marked with gray spots, brenching from

1948 Moldenke, Notes on Aegiphila 445

near the ground, the primary branches ascending, the secondary
branches spreading or nutant, and the flowers gray-green.
Dugand G. says it is a shrub with rather elongate branches, op-
posite leaves, and small tubular white flowers, growing on
shaded banks of small streams in forests. It has been collected
at altitudes up to 400 m., blooming in August and fruiting in
January. It has been confused with A. cordifolia (Rufz & Pav.)
Moldenke. The Slias 673, cited as Aas puberulenta Moldenke in
Brittonia 1: 415 (1934) and Phytologia 1: 259 (1937), is actu-
ally A. mollis day intermedia.

Additional citations: COLOMBIA: Atlantico: Dugand G. 1137
(F--930328); Elias 1458 (F--859231). Bolfvar: Slias 673 ee
photo, W--1442993, Z--photo). Magdalena: H. H. Smith 329 (Ca--
584590--isotype, Om--isotype, Vt--isotype).

AEGIFHILA MONSTROSA Moldenke
Williams describes this plant as a slender tree or tall
shrub, up to 25 feet in height, the bark light-gray, with a
pale-brown tinge, moderately smooth, less than 1/4 inch thick,
the inner bark light-brown, the trunk branching from the base,
4 inches in diameter, the fruit rounded, attached to the
branches. He found it at the edge of roadways in fairly dense
forests, and records the common names "café cimarron" and
"talalachi". Edwards found it in a dense tropical forest;
Standley and Steyermark in low or wet thickets. The fruit is
described by the two latter collectors as green or light-green,
appearing in March. Standley collected it at sea-level. B. P.
Reko in Mitobotanica Zapoteca, pp. 97 & 127 (1945) records the
common name "tala lachi", which, he says, is probably a corrup-
tion of "be-la lachi" or "be-laga lachi".
Additional citations: MEXICO: Caxaca: Ll. Williams 9120 (F--
haga BRITISH HONDURAS: — = Bartlett 11941 (F--659095),
11 (F--659092 ); Forestry (F--1001403); Gentle 263
Cam 63977), 404 get Pi bombo Gg--237829). GUA-
TEMALA: El Feten: C. L. Lundell 1 1492 (F--662818). Izabal: P. OC.
Standley 73129 ats Lappe Steyermark 38038 (F--1034591). H HON-
DURAS: Atlantida: P. Standley 55268 (F--583996). Cortés: J.
B. Edwards P.717 (eee. N). Yoro: Von Hagen & Von Hagen

1020 (F--943155).

tater MULTIFLORA Rufz & Pav.

e Dombey sen. from "Pasoén Huaru-hueri", cited in Phytolo-
gia te 251 (1937) as fron an undetermined department, is ectu-
ally from Madre de Dfos. Metcalf describes the species as a
bush 2--5 m. tall, with green calyx and levender corollas, in-
hebiting dry open places in rocky soil in a recion of much fog,
at an altitude of 2700 m., blooming in May.

Additionel citations: PRU: Huanuco: Rufz & Favon s.n. [Hua-

ssachuess & Felleo] (F--633375--photo of isotype). Funo: Red.

446 PHY TOLOGTIA Vol. 2, no. 10

Metcalf 30550 (W--1834986); Vargas ©. 1311 (F--989516). Depart-
ment undetermined: Dombey s.n. Chili et Perou] (F--998416);_
Rufz 187 [Macbride photo 17587] (F--663066--photo, Kr--photo,
N--photo); Rufz & Pavon 12/70 (F--712584, F--845337). _

AEGIPHILA NERVOSA Urb.
Additional citations: JAMAICA: Swartz s.n. (F--633326--photo
of type). HISPANIOLA: Haiti: Ekman H.472 (F--642166--photo).

AEGIFHILA NOVOFRIBURGENSIS Moldenke

The photograph of the label of the Delessert Herbarium iso-
type of this species, cited below, seems to prove definitely
that the number of the type collection of this species is "194"
{not "134" as hitherto cited by me in Brittonia 1: 350 (1934)
and Phytologia 1: 251 (1937)].

Additional citations: BRAZIL: Rio de Janeiro: P. Clausen 194
(Macbride photo 28386] (F--830244--photo of isotype, Kr--photo
of Lsotype, N--photo of isotype).

ABGIFHILA OBDUCTA Vell.

References: Lewkowitsch, Chem. Tech. & Analys. Oils, ed. 6,
678. 1922; Hoehne, O Jard. Bot. S. Paul. 576--577. 1941; Sampa-
io & Peckolt, Arquiv. Mus. Nac. Rio de Jan. 37: 334. 1943,

An additional synonym is Aegiphila obducata Vell. apud Samp-
aio & Peckolt, Arquiv. Mus. Nac. Rio de Jan. 37: 334, in syn.
1943. The accepted specific name is also spelled with a capital
initial letter in this reference.

Dusén reports the plant as growing at the edge of virgin for-
ests; Mello Barreto describes it as a tree 3--5 m. tall. The
Mello Barreto specimens are typical of the large-flowered bull-
ate-leaved form of the species; the Handro represents the thin
non-bullate-leaved form with smaller flowers. The flowers of at
least the former form are described as odoriferous. Williams &
Assis describe the plant as a “vine", with purple flowers,
growing in forests at an altitude of 1400 m. Lewkowitsch, in
the reference cited above, reports that a little-known Brazil-
fan oil is extracted from this species. Niederstadt has deter-
mined that the yield is 21.6 percent. The oil has a specific
gravity of 0.9579 at 26° C., its saponification value is 199.5,
its iodine value 64.15, and its acid value 72.2. The species
has been collected in anthesis in June, and in fruit in Decem-
ber.

eu tional 1), til ( BRAZIL: reste Geraes: Mello Barreto
2109 (F--955081), Q1ll (F--933073); Williams & Assis 714 (6,

N). Parana: Dusen 8143 (F--668475); Jonsson 3798 (Ca--533218,
F--668474). Rio de Janeiro: Brade 10542 [Herb. Rio de Jan.
22947] (Ja); Herb. Rio de Jan. 31720 (Ja). Santa Catherine:
Schwacke IV.184 [Herb. Rio de Jan. 32271] (Ja). Sao Paulo:
Herb. Rio de Jan. 32268 (Ja). State undetermined: Herb. Rio de

1948 Moldenke, Notes on Aegiphila 447

Jan. 32267 (Ja), 32274 (Ja). CULTIVATED: Brazil: S&o Paulo:
Handro, pl. viv. 442 [Herb. Inst. Biol. S. Faulo 33523] (F--
695762 ).

AEGIPHILA ODONTOPHYLLA Donn. Sm.

Additional citations: COSTA RICA: Guanacaste: Orsted 11174
1([Macbride photo 22775] (Kr--photo, N--photo). Heredia: Pittier
288 (F--635320--photo of type).

AEGIFHILA FPANAMENSIS Moldenke

Woodson has described the corolla of this plant as lemon-
yellow or "greenish-cream", Matuda as yellowish-white. It has
been collected in anthesis in August, and at altitudes of 1200
to 1500 m.

Additional citations: MEXICO: Chiapas: Matuda 2115 (Mh),
16791 (N). COSTA RICA: Alajuela: Brenes 4320 [105; 9516] (N),
9557 [180; 4395] (N). PANAMA: Chiriquf: Woodson & Schery 758
(N). Coclé: Woodson, Allen, & Seibert 1247 (F--969482, N), 1249
(F--969481, N), 1756 (N). Darien: P. H. Allen 856 (F--1005206).

AEGIFHILA PANICULATA Moldenke

Steyermark describes this plant as a vine, with pale-green
leaves and orange fruit; Allen says it is a tree to 3m. tall.
It has been collected in fruit in October, November, and Janu-
ary. The Brenes 16823 and Steyermark 50757, cited below, have
very typical fruit for this species and very typical fruiting-
calyxes. They need only be compared with fruiting sheets of A.
panemensis for proof that the two species are quite distinct.
The Allen 1124 cited below was distributed as A. falcata Donn.
Sm., but has the closely investing cupuliform calyx and con-
spicuously punctate leaves of A. paniculata. It was collected
at an altitude of only 35 m.

Additional citations: GUATEMALA: Chicuimula: Steyermark
30757 (F--1057191). COSTA RICA: Alajuela: Brenes 1€823 [2] (F--
858958, N). PANAMA: Penama: P. H. Allen 1123 (F--1005205).

AEGIFHILA PARAGUARIENSIS Briq.

Rojas collected this especies "hanging over cliff", and Dusén
found it at an altitude of 730 m.

Additional citations: BRAZIL: Mattogrosso: H. H. Smith s.n.
(Herb. Rio de Jan. 32269] (Ja). Farand: Dusén 15963 (F--668476).
Séo Faulo: Heiner 270 (S). PARAGUAY: Hassler 4498 Rinsiencs
photo 24618] (P--772032--photo of cotype, Kr--photo of cotype,
N--photo of cotype); Rojas 12768 (N).

AEGIPHILA PARVIFLORA Moldenke
The label on the Macbride photographs, cited below, reads
"2296" in error. The plant depicted ie Spruce 589.
Additional citations: BRAZIL: Fara: Spruce 589 [Mecbride

448 Pill: ¥.20: bcos Tek Vol. 2, noe 10

photo 28387] (F--830276--photo of isotype, Kr--photo of iso-
type, N--photo of isotype).

AEGIPHILA PAVONIANA Moldenke

Haught describes this as a small tree about 5m. tall, with
fairly conspicuous inflorescences of cream-colored flowers,
blooming in December at an altitude of 1CO m.

Additional citations: ECUADOR: Guayas: Heught 3004 (N).

ASGIFHILA PERNAMBUCENSIS Moldenke

Noraes Vasconcellos has found this plant in woods and re-
cords the common names "caféeiro de cabra" and "cafeliro de
cabra". The species is closely related to A. australis Moldenke
and not to A. crenata with which it has been confused, but
which belongs in a different subgroup of the genus.

Additional citations: BRAZIL: Parahyba: Moraes Vasconcellos
841 (N), 853 (N), sen. (Herb. Serv. Florest. Est. S. Paulo 841]
W--1564377), sen. (Herb. Serv. Florest. Est. S. Faulo 455] (W
--1564367.). Pernambuco: Fickel 3042 (Du--255725--isotype, Mi--

isotype).

AEGIPHILA FERPLEXA Moldenke

Steyermark describes this plant as a tree 20--25 feet tall,
with firmly membranous leeves, rich-green above, pale dull-
green beneath, calyx rich-green, corolla greenish-yellow, and
style whitish, blooming in April at an altitude of 1200--1480 m
“aah eapaaaan citations: VENEZUELA: Monsgas: Steyermark 62260
N s

AEGIPHILA FERUVIANA Turcez.

Klug has collected this species at altitudes of 1200 to 1600
meters.

Additional citations: FERU: San Mart{n: Klug 3511 (FP--736324)
Spruce 4275 [Macbride photo 24619] (F--772033--photo of iso-
type, N--photo of isotype).

AEGIPHILA FLATYFHYLIA Briq.
Additional citations: PARAGUAY: Hassler 8056 [Macbride photo

24620] (F--772028--photo of isotype, Kr--photo of isotype, N--
photo of isotype).

AEGIFHILA PUBERULENTA Moldenke

This plant is described as a shrub 2--4 m. tall or a tree,
with white or creamy-yellow corollas which soon fall off, Prag-
rant, bloomirgin June and October, at altitudes of 50 to 510 m.
It inhabits thickets and is abundant on limestone soil. It has
been widely confused with A. mollis, A. mollis var. intermedia,

and A. plandulifera.The Elias 675 cited as A. puberulenta in
Brittonia 1: 413 (1934) and Phytologia 1: 259 987) proves to

1948 Moldenke, Notes on Aegiphila 449

be A. mollis var. intermedia. Common names recorded by Dugand
are “bollo limpio" and "San Juan de la verdad". This distingu-
sshed botanist notes that his no. 720, cited below, is "probab-
ly the same as no. 639", but I regard the latter as A. mollis.

Additional citations: COLOMBIA: Atléntico: Dugand G. 256
(Mus. Yale School of Forestry 22545] (F--664070), 720 (F--
744907); Elias 581 (N), 1102 (F--699290, N, N, N). Bolfvar:
Dugand & Jaramillo 3431 (W--1852289). Magdalena: Haught 2288
Or VENEZUELA: Aragua: Ll. Williams 10222 (F--946533).

AEGIFHILA QUINDUENSIS (H.B.K.) Moldenke

Williams reports this to be a shrub or small tree, 1.5 me
tall, with light-gray fairly smooth bark, creamy-white corollas,
and globular yellowish or vermillion fruit. The fruit is very
large, 1.3--1.5 cm. in diameter, the fruiting-calyx very shal-
lowly cupuliform, wide-spreading, about 1 cm. wide. heavy,
glabrous, its rim distinctly lobed. The species has been found
in bloom in May, and et altitudes of 450--960 m. It has often
been confused with A. martinicensis. Steyermark describes it as
a tree 20 feet tall, with membranous leaves that are dark-green
above and pale-green beneath, the calyx pale yellow-green, the
corolla pale-yellow, and the filaments whitish. He found it at
altitudes of 1200 to 1450 m.

Additional citations: VENEZUELA: Aragua: Pittier 14993 (W--
1833196), 15474 (W--1909582), 15481 (W--1909584); Ll. Williems
10251 (F--946419, F--989699, Gg--295555), 10391 (F--948392).
Carabobo: Karsten s.n. (F--642170--photo). Monagas: Steyermark
62046 (N).

AEGIPHILA RACEMOSA Vell.

Sampaio & Peckolt in Arquiv. Mus. Nac. Rio de Jan. 4/: 334
(1943) reduce this species to A. cuspidata Mart., but this is
an error. Martius' species is conspecific with A. vitellini-
flora. Steyermark describes the plant as a “vining shrub", 15
to 25 feet tall. Monteiro da Costa calls it a “vine", blooming
in January, called “cawuirea", inhabiting lowlands, and used in
aromatic baths for nervous diseases.

Additional citations: VENEZUELA: Merida: Steyermark 56728
(F--1221912, N). BRITISH GUIANA: De la Cruz 4292 (Cm), 4552
(om); Herb. Forest Dept. Br. Guian. 3081 [F.345] (K). BRAZIL:

id

Para: Monteiro da Costa 240 (F--693925).
AEGIPHILA RIEDELIANA Schau.

Sampaio & Peckolt in Arquiv. Mus. Nac. Rio de Jan. 4/7: 334
(1943) reduces A. serrata Vell. to A. graveolens Mart. & Schau.,
but as has been pointed out by me in Brittonia 1: 311 (1934) it
seems more properly to belong with A. Riedeliana.

Additional citations: BRAZIL: Bahia: J. &. Fohl 4392 [Mac-
bride photo 34309] (Kr--photo of cotype, N--photo of cotype).

“450 PHYTOLOGIA Vol. 2, no. 10
Rio Grande do Sul: Rambo 29169 (N).

AEGIFHILA RORAIMENSIS Moldenke

Steyermark describes this species as a shrub 5--8 feet tall,
with subcoriaceous leaves thet are dark-green above and dull-
green beneath with gray-buff pubescence, stems buff-pubescent,
and calyx gray-buff. He found it in woods bordering a savanna
on ea ridge above La Laja, at the base of Sororopan-tepui, alt.
1375--1460 m., blooming in November. It has been confused with
the genus Citharexylum.

giao citations: VENEZUELA: Bolivar: Steyermark 60812
(N).

AEGIFHILA SALTICOLA Moldenke

The supplementary characters given by me for this species in
Fhytologia 1: 397--398 (1940) applies only to the Ducke speci-
men there cited. It seems, on re-examination, that this speci-
men may actually be A. intermedia Molénke. It is possible that
A. salticola is actually conspecific with A. intermedia. Mexia
records the common name "genipapo do matta",

Additional citations: BRAZIL: Maranh&o: Froes 11856 (N).
Pard: Mexia 5922 (Gg--286582--isotype).

AEGIPHILA SCANDENS Moldenke

Ducke describes this as a woody climber with greenish-white
flowers, blooming in March, growing in old secondary forests on
terra firma.

Additional citations: BRAZIL: Amazonas: Ducke 1190 (N, W--
1832444),

AEGIPHILA SCHIMPFII Moldenke

The type collection of this species, cited by me in Fnytolo-
gia 1: 266 (1937) as from "Biscay, Ecuador", was actually col-
lected at Bucay, Guayas, Ecuador. Svenson in Am. Journ. Bot.
33: 480 (1946) describes the species as a shrub 6--9 feet tall,
with yellow flowers and exserted stamens, found along streams
near sea-level, blooming in April.

Additional citations: ECUADOR: Guayas: Svenson 11448 (N).

AEGIPHILA SELLOWIANA Cham.

References: Hoehne, Kuhlmann, & Handro, O Jard. Bot. S. Paul.
577. 1941; Instit. de‘Botan. Observ. Ger. Contrib. 5:19 & I.
1942; Hoehne, Relat. Anual Inst. Bot. 1944: 118. 1944.

It has been collected in fruit in March and April. Additional
common names are “pau de tamanco", "tamanqueira", "tamanqueiro",
"cinzeiro", and "papagaio". Mexia describes it as a slender
tree 10 m. tall, with a pithy stem and long straggling branches
and slightly fragrant greenish-white or white flowers and fruit
in heavy clusters, growing in second-growth woods comnonly.

A PROPOSAL TO STABILIZE PLANT NAMES

Elbert L. Little, Jr.

The essential points in botanical nomenclature are fixity
of names and rejection of names which may cause error or
"throw science into confusion" (Art. 4, International Rules of
Botanical Nomenclature. Ed. 3. 1935). All systematic bota-
nists should strive towards a more stable nomenclature, espe-
cially for the benefit of workers in other brancnes of plant
science throughout the world, so that botany can make satis-
factory progress (Art. 1).

THE PROBLEM

A serious obstacle to the gosl of stability of names is the
revival in recent years of many old, abandoned names. Some
were so obscurely published that they were unknown to contem-
porary botanists and escaped notice of indexers. Other names
were poorly described in the first place and are of doubtful
application in the absence of type specimens. Except for the
fact that under the Rules they retain priority from their
Original publication, these long-lost names are new names.
However, as old names under the Rules, these names must be ac-
cepted, even if other names meanwhile have become established
in usage. Also, these old names must be credited to their
original authors, who scarcely deserve to be so honored now at
this late date.

Four recent changes in names of trees of the United States
will serve as examples. These old names upsetting existing
nomenclature were not in Index Kewensis.

Abies nobilis A. Dietr. (Fl. Berlin 793. 1824), an obscure
synonym and earlier homonym, was the basis for the rejection
in 1940 of A. nobilis (Dougl.) Lindl. (Penny Cycl. 1:30. 1833),
a name universally established in usage without synonyms. As
the latter technically was invalid as a later homonym, it was
renamed A. procera Rehd. (Rhodora 42: 522. 1940).

Juglans microcarpa Berland. in Berland. & Chovel (Diario
Viage Comisién Limites Mier Terén 276. 1850). This briefly
described name concealed in a Mexican diary of travels appar-
ently was unknown to botanists until adopted by Johnston (Arn-
old Arboretum Jour. 25: 436. 1944) to replace the familiar
name, J. rupestris Engelm. ex Torr. (in Sitgreaves, Rpt. Exped.
Zuni Colo. Rivers 171, pl. 15. 1653), which was without known
synonyms.

Ulmus rubra Muhl. (Amer. Phil. Soc. Trans. 3: 165. 1793).
This name in a local flora ee proposed merely as a new name

Sl

450 PHYTOLOGIA Vol. 2, no. 10

without description for U. americana Marsh. (Arbustr. Amer.
156. 1785), not L. (Sp. Pl. 226. 1753), was revived in 1945,
after 152 years of dormancy, by Fernald (Rhodora 47: 203-204,
1945). The name in universal use which now must be rejected
as a synonym is U. fulva Michx. (Fl. Bor.-Amer. 1: 172. 1803).

Cotinus obovatus Raf. (Autikon Botanikon 82. 1840), brief-
ly described in a rare work of Rafinesque, apparently was not
again accepted until a facsimile reprint of this rare book was
published in 1941. The established name, C. americanus Nutt.
(No. Amer: Sylva 3: 1, pl. 81. 1849), thus was technically
invalid asa synonym. Accordingly, C. obovatus Raf. was
adopted by Little (Okla. Acad. Sci. Proc. 23: 21-23. 1943).

Other illustrations will be familiar to readers. Changes
such as these, not uncommon in current taxonomic publications,
not only do no good but create confusion in violation of
Art. 4. Asa result, taxonomy is injured in its relations
with other branches of botany, whose workers do not understand
how continual changes in names can constitute progress towards
stability.

The problem, therefore, is to find a way to prohibit or
lessen the revival of old, abandoned names.

POSSIBLE SOLUTIONS

Several solutions of the problem may be considered. FPer-
haps the simplest would be the establishment of a code of eth-
ics among taxonomists, a gentlemen's agreement not to take up
these old names. Fossibly Art. 5, to follow established custom
in the absence of a rule, might be stretched to cover these
cases. However, the prevailing custom seems to be the op-
posite, to bring to light all these old names as soon as possi
ble. A few botanists say that when they run across an old name
that might upset the accepted nomenclature, they put the book
back on the shelf. This admirable practice, though, merely
‘postpones the upheaval and permits it to become greater, for
sooner or later another worker with different ideas probably
will discover the same name in the same book. Then, the appar
ent oversight of the old name by the first monographer may be
interpreted by the second as evidence of lack of thoroughness
in bibliographic work. Seldom do new combinations follow
revival of old names. The reward for the discovery is the
example of careful bibliographic work and perhaps a sense of
importance in causing the change. As the temptation to revive
an old name is great, voluntary agreement seems unlikely as a
solution.

The problem may become progressively less important in the
future, as more and more old names are adopted, because, after
all, the number of different rare books printed in the past
from 1753 to date does have a limit which eventually will be

1948 | Little, To Stabilize Flant Names 453

approached. On the other hand, publication of obscure scien-
tific books and journals continues.

Good modern library facilities, including bibliographic,
abstract, and indexing services and wide circulation of publi-
cations tend to prevent recently published names from being
overlooked. However, the enormously increased quantity of
botanical publications in recent years operates to offset the
library aids.

Art. 38, requiring Latin diagnoses for names of new groups
of plants published after Jan. 1, 1935, probably will be of
great value in the future in making illegitimate various ob-
scurely, inadequately, and incidentally published names other-
wise valid.

Other solutions involve exception to the fundamental prin-
ciple of priority (Art. 16). During certain times in the past,
retention of names lacking priority was accomplished through
the influence of leading workers. In some ways priority seems
to conflict with stability. That priority is not sacred is shown
by the long list of nomina generica conservanda adopted under
Art. 21 and without which, nomenclature would be chaotic and ex
ceedingly unstable. Under this rule any rediscovered old ge-
neric names which would cause disadvantageous changes can be
formally rejected.

Conservation of specific names in exception to priority has
been rejected decisively at previous Botanical Congresses and
is not a likely solution. It does not seem feasible to makea
special exception in the Rules for a single specific name,
when it is simpler to retain the older name. Rules affecting
names in general published under similar conditions are less
complex in operation than rules permitting special exceptions
and requiring action by an International Botanical Congress
upon each name.

One attempt toward stability was the adoption at the last
Congress in 1935 of a motion for a committee to draw up a list
of economic plants under the Rules and that this list remain
in use for a period of ten years. Though the list was not
prepared, a list of standard generic names was issued. In one
country an official tree list including a few invalid but well-
known names was adopted by foresters.

Proposals have been made to amend the Rules to reject names
in certain old or rare works. For example, at the last Con-
gress a proposal to reject names ina list of old works not
using binomials was referred to a committee for study. How-
ever, a rule containing a list of books would be of question-
able value and would not eliminate confusion, because there
would still be other and rarer books not covered.

A radical suggestion has been. made to establish new starting
points of priority, such as modern monographs. Even the Rules
(Art. 20) permitted later starting points than 1753 fora few

j
454 Po XoMeO Le Oeiten Vol. 2, no. 10

groups. Perhaps in the distant future, when nomenclature be-
comes extremely complicated, this suggestion may be adopted by
necessity.

Another but rather discouraging possible solution is that,
if world peace is not established, the atomic bomb and global —
warfare might lead to the destruction of civilization, includ-
ing the botanists with their books, herbaria, and Rules. Then,
at some later date there might arise an altogether different
system of botanical nomenclature with a new set of rules, new
starting date, and entirely new names.

THE PROPOSAL

I believe an addition to the Rules is desirable to help
maintain stability by prohibiting the revival of old, aban-
doned names. An informal note that I favor “amending the rules
to disallow priority changes due to later discoveries in obscure
books 100 years or more old" has been published (W. A. Dayton,
Jour. Forestry 41: 373. 1943). My proposed addition to the
International Rules of Botanical Nomenclature follows:

Article 63 bis. A name (of a taxonomic group) more than one
hundred years old but which has not been accepted as valid, so
far as known, by any subsequent author (exclusive of indexes of
nomenclature) within the first one hundred years after publi-
cation (or by Jan. 1, 1950, in the case of a name published
before 1850) must be rejected as a nomen extinctum if it is an
earlier synonymor earlier homonym of any name otherwise valid
and accepted in use.

In other words, an extinct name, or nomen extinctum, is a
name which was accepted by no other authors within the first
hundred years after publication but which during this time has
been replaced by another name or has been used for another
group. As both the old, unused name and its synonym or homonym
cannot be retained in use, the old name, upon its discovery is
retained in accord with the principle of fixity of names.

Though this proposal would apply to all taxonomic groups,
its chief value would be for names of species and their subdi-
visions. Retention of generic names in exception to strict
priority as nomina conservanda has been provided under Art. 21.

Under this proposal, acceptance by a second author within a
century automatically guarantees a name its priority. However,
mere listing of the name as a Synonym by later authors would
not constitute acceptance, Neither publication of the name in
&@ second work by the original author nor reprint of the origi-
nal work, such as a facsimile edition of a rare book, would
count. It has seemed best to exclude indexes of nomenclature
as not constituting acceptance of the name by a second author.

1948 Little, To Stabilize Plant Names 455

i)
Some indexes do not attempt to pass upon the validity or syn-
onymy of their names. Many overlooked specific names were
omitted from Index Kewensis, though upon discovery afterwards
were included in the Supplements. Also, some names of doubt-
ful identity are listed by indexes as a bibliographic record.

The year 1950, when this proposal would become effective,
if adopted asa rule, has been set as the starting date to
apply to all names more than one hundred years old; that is,
names published between 1753 and 1850. Without this starting
date the proposal would be retroactive (Art. 2) to names be-
coming one hundred years old in 1853 and successive years, and
some names restored after an interval of more than one hundred
years but now already accepted in usage would be invalidated.
In the future, names published after 1850 would automatically
be rejected upon remaining unknown and unaccepted by a second
author for a century. For example, an obscure name published
in the year 1868 would retain its validity and priority if
discovered and used by a second author before 1968. If not
discovered until after 1968, this name would be rejected pro-
vided it had a synonym or homonyn.

The final clause, “if it is an earlier synonym or earlier
homonym of any name otherwise valid and accepted in use,“ is
essential. When I first discussed my proposal, one botanist
protested that a few namesof texonomic groups of small size or
of restricted geographic distribution might pass a century mown
but dormant because later botanists had had no occasion to re-
fer to them. To invalidate these dormant names without syno-
nyms would leave their taxonomic groups nameless. So, if it
has acquired neither a synonym nor a homonym, the old name re-
tains its priority and is not rejected as an extinct name.

The proposed rule would work like this. A systematic bot-
anist in the course cf his work discovers an obscure name in a
rare book more than a hundred years old and from the descrip-
tion identifies it with a later name in use. Or, he recalls
that e later homonyr is in use. A search through pertinent
literature fails to disclose acceptance of this old name by
another author. Thus, the old name clearly must be rejected
as a romen extinctum. The discoverer then publishes a taxo-
nomic note formally rejecting the name and giving himself due
credit. Thus, one more name in use is retained, and one or
two confusing changes in names ere avoided.

There would also be broader effects. This proposal would
automatically invalidate many known names of doubtful identity,
especially those inadequately described and without type speci-
mens, if afterwards they are ever found synonymous with later
names in use. For example, future workers need not spend time
on tke names of Rafinesque which have not been taken up by an-
other author, probably several thousend names. This proposal
would simplify the nomenclature of varieties by preventing ac-

456 PHYTOLOGIA Vol. 2, no. 10

ceptance of many old, briefly described varieties. Otherwise, —
these old varietal names, which generally are not indexed, may ~
cause confusion as the taxonomists of the future turn more to
the recognition of subdivisions of species.
To a minor. extent, this proposal would contradict Art. 61,1
which rejects later homonyms but which was not adopted until
1930. A later homonym would be legitimate in those infrequent —
cases not already corrected where the earlier hcmonym is more
than one hundred years old and has not been adopted by a second —
author. Thus, some later homonyms invalidated in 1930 — by
Art. 61 but not yet renamed could be retuined in usage. 4
The application of Art. 21, which provides for conserved
names, would be simplified by this proposel. Some very old —
generic names, particularly earlier homonyms not yet formally —
made nomina rejicienda, would automatically be rejected as
nomina extincta. There would be no need to act upon these
names individually and add the later names to the already
lengthy list of nomina conservanda. The following examples of —
generic names of trees proposed by me for conservation
(Madrofio 7: 24C-251. 1944) could be retained without special —
action under this proposal: Cedrus Trew, Condalia Cav., Rha-
coma L., Bucida I., and Halesie Ellis. Ag
THe suggested time limit of one hundred years could be low-
ered, if desired. For example, Art. 21 suggests that in the ©
selection of nomina conservanda preference be given to names —
which have come into general use in the fifty years following —
their publication. 7
In some irstarces it may be difficult to determine whether #
the old name has been taken up by a second author. There is
the possibility that e name once rejected as a nomen extinctum
would afterwards be found in a later work and would have to be
adopted. Also, it may not always be clear whether en author
mentioning a name accepts it as valid. However, all names not —
conservec are subject to some risk of change. i
This proposed addition to the International Rules has bea
submitted to Dr. W. H. Camp, Chairman, Committee on Nomencla-
ture, American Society of Flant Taxonomists, New York Botani- ~
cal Garden, New York 58, N. Y. The Committee is considering —
proposals for amendment to the Rules to be officially spon-
sored by the Society at the next International Botanical Con-
gress in 1950. a
A discussion of this proposal is presented here, in order
that interested botanists may consider it. Perhaps improve- —
ments in the proposal and its phraseology will be suggested and ~
appropriate examples will be offered. Whether a majority of —
systematic botanists would favor adding to the complicated Rules —
@ proposal of this kind to lessen the revival of old, abandoned ~
names is not known. *

=

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BeeHYTOLOGIA ~

Designed to expedite botanical publication

July, 1948 No. 11
sg eee CONTENTS
Bee) : ¢ ’ :
4 “ .
Lamm, E. L. , jr. ‘ Notes on nomenclature Of treeS.eccss.ssssssrseseseee 457
- Motpenke, H. N., Notes on. new and noteworthy plants. V........464

r. _ MowvEnxe, H.N., The known geographic distribution of the

a members the Verbenaceae, Avicenniaceae, i ss—is
7 _ Stilbaceae, and Sym phoremaceae. Supplement 9.......c.00000+. 477 -
7 ~ Monacnuno, J. V., Capparis brevis Sprengel is a Glyphaea..... sie 484
Larne, E. L.,-Jr., David Douglas’ new species of Conifers... 485
Moxpenxz, H.N., The known geographic distribution of the
q _ members of the Eriocaulaceae. Supplement 3......s..s.-0eoe0e 490
| Mowenxe, H. N., Additional notes on the genus |
a OEE a ogc rs ccaeWN aae AL techs ctnages tapmuntlis onc. <seusoas Uaneereatonaoune 499
_ Motpenxg, H. N., Additional notes on the genus oe
a - Amasonia. faa Pisa aiaial sol teh Hale ssl ch Scdnad andtan dvs ose At aioe ata ee
' Gueason, H. A., Specific names in Gratiola.ssessssssesssn is sa Natale 503
a Author index to Volume Tw0..cccccrseossssssesssssresssscssssesssesessseee Sa 504
_ Subject index to Volume MT MOGE, iss abicketesisnstoccessadsiavsslion pf Sakdene aoe
a Publication dates of Volume Tw0......ssissesssrsrseersneen aGeeseng pte 1
a
a Published by H. A. Bouck and Harold N. Moldenke

The New York Botanical Garden,
Bronx Park, New York 58, N. Y.

Price of this number, 75 cents; per volume, $5.00 in advance .

Vol. 2, No. 10, was issued April 22, 1948

nis ined
ae

e

NOTES ON NOMENCLATURE OF TREES

Elbert L. Little, Jr.

Here included are notes on the nomenclature of longleaf
pine (Pinus palustris Mill.), Siebold walnut (Juglans ailan-
tifolie Carr.), and sugar maple (Acer saccharum Marsh.) and
@ new combination in Glycosmis.

LONGLEAF PINE, PINUS PALUSTRIS

The name Pinus palustrie Mill. (Gard. Dict. Ed. 8, Pinus
No. 14. 176&) has been applied, with some confusion, to two
species of pines of southesstern United States. Most recent
authers have adopted Pinus pelustris Mill. for the longleaf
_ pine end Finus ceribaea Mor Norelet (Rev. Hort. Cote d'Or l: 105.

1851 (mot seen); Soc. Hist. Nat. Moselle Bul. 7: 100. 1855)
_ for the slash pine.
. However, Small (Man. Southeast. Fl. 4, 5. 1933) used Finus
palustris for e variation of slesh pine known also as swamp
pine and applied to other veriations of siash pine the names P.
caribaea and FP. heterophylla (Ell1.) Sudw. (Torrey Bot. Club
But, 202.45. 1693; not P. heterophylla K. Koch, 1849, nor
Presl, 1849). rt the longleaf pine, Small sccapbed F. _aus-
tralis Kichx. f. (Hist. Arbr. Amér. Sept. 1: 64, pl. 6, 1810).
Previously, cine (Fi. Southeast. U. S. 27. 1963) had used FP,
palustris for the longlesf pine, with P. australis as a syno-
nym. Sargent (Silva No. Amer. 11: 151. 1897) summarized the
older references adopting F. palustris and F, australis for the
longleaf pine. Recent authors accepting P. palustris for the
swamp pine and P. austreaiis for the longleaf pine inciude Van
Dersal (Native Woody Flants U. S. U. S. Dept. Agr. Misc. Pub.
303: 187, 191. 1938)and De Vall (Fla.Acad. Sci. Froc.5 (1940):
121-132. 1941).

As it,is the oldest name, Finus palustris Mill., "the three~
leaved Marsh, American Fine with the longest leaves," should
be adopted for one particular species, if the description is
considered ia Oph for recognition of a species. Unfortunate-
ly, the original description, based upon Pinus Americana palus-
tris trifolia, foliis longissimis Duhamel (Traité Arbr. Arbust.
France 2: 126. 1755), is rether brief. The specific epithet,
translated by Miller es "marsh," is misleading for the large,
upland longleef pines, as is Miller's statement that they grow
“naturally on swamps in many parts of North America, where I
have been informed they grow to the height of twenty-five or
thirty feet." However, he added: “Their leaves are a foot or

457

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4
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j
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4

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458 Pay? OL te ee Vol. 2, mos 11%

more in length, growing in tufts at the end of the branches,
60 have a singular appeerance ..."

Pinus australis Michx. f., "the long leaved pine,” accom-
panied by a latin diagnosis, a colored plate, and 22 pages of
French descrijytion and discussion, including turpertining, is
identified beyond doubt as the femiliar longleef pine. How- |
ever, F. A. Michaux cited as synonym "P. palustris, Linn." and
indicated that he was renaming P. palustris because that name
was not appropriate for a species not of swamps. His exact
words (p. 65) were: "J'ai pensé égalerent que la dénomination
spécifique d‘australis étoit préférable A celle de palustris,
sous laquelle cette espece est décrite per les botanistes; car
cette derniére donne une idée absolurent fausse de la nature
du sol ob croit cet arbre." The substitute name, F. australis,
is not especially eppropriate either, as there are several spe
cies of southern pines. Miller's name was not cited as author,
but the mention of Linnaeus probably is sufficient to connect
the name and synonyry irregularly through Willdenow's edition
(Ed. 4) of ‘Linnaeus’ Species Planterum (4 (1): 499. 1805) and
older references, such as Michaux (Fl. Bor.-Amer. 2: 204. 1€03),
back te Miller's original publication.

Thus, under Art. 60 (1) of the International Rules, P. aus-
tralis Michx. f. is invalid, as it was nomenclaturally super-~
fluous when published. It must be rejected and cannot be used
for the longleaf pine. Furthermore, under Art. 59, P. palus-
tris must not be rejected merely because it is badly chosen or
disagreeable in stating the habitat incorrectly as marshes.

Pinus palustris Mill., the name generally used, should be
retuined for the longleaf pine. Its identity seems clear in
spite of the mincr inaccuracies in the Original deecription
noted above. No other species of this regicn hes needles more
than & foot long. “Even F. A. Michaux in renaming the species
recognized Miller‘s short description as applying to the long
leaf pine. Continued use of Miller's name for @ seccnd species
would result in further confusion.

The rame Finus caribaea Morelet apparently is the oldest
aveilable name for the slash pine. Whether the more northern
variation merits specific segregation or is more proverly a
geographic race not requiring a separate scientific name is
uncertain. Additional field study of these veriations would
be desirable. The aveilable specific name for this swamp pine
is P. elliottii Engelm. (Acad. Sci. St. Louis Trans. 4: 186,
pl. 1-3. 1880). However, the differences seem no greater than
those cf geograyhic races of certsin other species of pines
with extensive ranges.

1948 Little, Nomenclature of Trees 459

SIEBOLD WALNUT, JUGLANS AILANTIFOLIA

While checking the norienclature of the trees of the United
States, I observed that the scientific name of the cultiveted
Siebold wainut from Japan, Juglans sieboldiana Maxim., was
technically invalid as a later homonym of the fossil species
J. sieboldisna GBppert. Accordingly, I adopted J. ailantifolia
od for the Siebold walnut (Wash. Acad. Sci. Jour. 33: 132.
1943).
| Rehder (Arnold Arboretum Jour. 26: 68. 1945) accepted this
nomenclatural change and made a new combination for the vari-
ety, Juglans ailantifolia var. cordiformis (Maxim.) Rehd. Aft-
_erwards Rehder (Arnold Arboretum Jour. 26: 472. 1945) adopted
for the specific name J. cordiformis Maxim., published simul-
taneously with J. sisboldiana Maxim. and previously united with
the latter as the variety. The new combination J. cordiformis
var. ailantifolia (Carr.) Rehd. was proposed also.

However, by odd coincidence Maximowicz's two new species of
Juglans published on adjacent pages both are invelid as later
homonyms. J. cordiformis Maxim. is not available either, be-
cause of the much earlier J. cordiformis Wangenh., a name not
in Index Kewensis but familiar asthe basonym of Carya cordiformis
(Wangenh.) K. Koch, bitternut hickory. Thus, J. ailantifolia
Carr. remains the valid name for the Siebold walnut.

The essential synonymy of the species and variety are summa-
rized below. Additional later synonyms were cited by Rehder.

JUGLANS AILANTIFOLIA Carr. SIEBOLD WALNUT
Juglens sieboldiana Maxim., Acad. Impér. Sci. St.-Pétersb.
Bul., sér. 3, 16: 6C, fig. 1872. Not Juglans siebold-
jana GBppert, Tert. Fl. Insel Java 154. 1854; nomen nudum.
Not Juglens sieboldiana GUppert, Tert. Fl.Schosenitz Schles.
Bey ple 25 fie. 2. 155 (fossil, Miocene, Silesia).
Juglans cordiformis Meaxim., Acad. Impér. Sci. St.-Pétersb.
Bul., s6r. 3, 10: 62, fig. 1872. Not Juglans cordiformis
Wangenh., Beytr. Forstwiss. Nordamer. Holz. 25, pl. 10,
fig. 25. 1787; as "“Iuglans."
Juglens ailantifolia Carr.,Rev. Kort. [Paris] 50: 414, fig.
5-86. 1678.

JUGLANS AILANTIFOLIA Carr. var. CORDIFORMIS (Makino) Rehd.
FLAT SIEBOLD WALNUT (heartnut )
Juglans cordiformis Maxim., Acad. Impér. Sci. St.-Pétersb.
Bul., sér. 3, lO: 62, fig. 18723; later homonym.
Juglens sieboldiana var. cordiformis [Maxim. | Makino, Bot.
Mag. Tokyo 9: 313. 1895; 15: 94. 1901.
Juglans ailantifolia var. cordiforris Rehd., Arnold Arbore-
tum Jour. 26: 66. 1945.

460 PAY? © L-@6: 5:4 Vol. 2, no. ll

Juglens cordiformis var. ailantifolia (Carr.) Rehd., Arnold
Arboretum Jour. 26: 472. 1945.

SUGAR MAPLE, ACER SACCHARUM

The scientific name of the sugar maple, Acer saccharum Marsh.
(Arbustr. Amer. 4. 1785), has been the subject of much con-
troversy in recent years. Some botenists have rejected this
name as a misspelling or orthographical error of A. saccharinum
L. (Sp. Pl. 1055. 1753) and have taken up A. saccharophorum K.
Koch (Hort. Dendrol. 80. 1853) or A. nigrum Michx. f. (Hist.
Arbr. For. Amér. Sept. 2: 238, pl. 16. 1612), if the two spe-
cies are united. Majority opinion seems to favor retentionof
the widely accepted name, A. saccharum. This name probably can
be retained under Art. 6, which provides for following estab-
lished custom where the consequences of rules are doubtful.
However, it is hoped that the permanent International Executive
Committee to interpret the Rules in doubtful cases (Art. 73)
will issue an Opinion on Acer saccharum. Otherwise the nomen-
clature will remein unsettled and subject to future proposals
for change fron time to time.

So much has been written about the nomenclature ofthe sugar
maple that it is difficult to contribute new information. The
most detailed histcry is that by Rousseau (Nat. Canad. 67: 161-
2CO, 201-224, illus. 1940. Reprinted as: Univ. Montréal Inst.
Bot. Contrib. No. 35, 66 p., illus. 1940. Also, No. 36: 36-
37. 1940). In rejecting A. saccharum Marsh., Rousseau has led
others to accept A. saccharcphorum K. Koch. Attempts to inter-
pret Mershall's intention, of which the latest is by Gleason
(PHYTOLOGIA 2: 201-212. 1947), have not been entirely satis-
factory, because the interpretations have differed.

Though now esteblished in usage, Acer saccharum was not
adopted by other authors until more than a century after its
publication in 1785. Britton (N. Y. Acad. Sci. Trans. 9: 10.
1889; Cat. Fl. N. J. Geol. Surv. N. J. Rpt. 2 (1): 78. 1890)
revived the name in 1889 and made the combination A. saccharum
var. nigrum (Michx. f.) Britton. Widespread acceptance prob-
ably dates back only about forty years to the publication in
1906 of the seventh edition of Gray's Manual by Robinson and
Fernald. Older botanists still active learred the names in the
sixth edition of Gray's Manual by Watson and Coulter (1889), in
which the sugar maple was A. sacchérinum Wangenh. and the sil-
ver maple was A. dasycarpum Ehrh. The double change of A. sac-
charinum from the sugar maple to silver maple and the substitu-
tion of the unfamiliar, almost identical néme, A. saccharunm, for
the sugar maple doubtless caused temporary confusion and was
unpopular. Surely it was a greater disturbance than the present
proposed change from A. saccharum to A. saccharophorum.

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1948 Little, Nomenclature of Trees 461

Sargent (Gard. and Forest 2: 364. 1889; 4: 148. 1891) at
first refused to take up A. saccharum, interpreting it asa
misprint. Noting also that Marshall's plant could not be sat-
isfactorily determined from the description and that Marshall
left no herbarium, Sargent concluded (p. 148) that "the only
safe way is to pass over his name entirely." In his Silva
(Silva No.Amer. 2: 97. 1892) Sargent adopted A. barbatum Michx.
(Fl. Bor.-Amer. 2: 252. 1803). However, ina supplementary
volume (Silva No. Amer. 13: 7. 1902), he rejected that nameas
based on a mixture and reluctantly accepted A. saccharum “for
the sake of uniformity of nomenclature,” while repeating his
objections.

Adoption of Marshall's name has not been universal. In 1913
Nieuwland (Amer. Midland Nat. 3: 182. 1913) rejected A, saccha-
rum as “absurd and besides homonymous" and “ungrammatical." Mac-
kenzie (Rhodora 28:111-112, 233-234. 1926) contended that this
“fictitious name" should be abandoned. Introducing new evidence,
he noted that in the French edition of Marshall's book, published
in 1788, the spelling was corrected to A. saccharinum, and he
cited an earlier spelling, A. sacchatum Mill. (Gard. Dict.
Abridged. Ed. 6, Acer No. 6. 1771). Small accepted Marshall's
name in his Flora (Fl. Southeast. U. S. 741. 1903) but re-
jected it in his Manual (Man. Southeast. Fl. 824. 1933) as
“merely a misspelling."

Marshall's Arbustrum Americanum (169 p. Philadelphia, 1785)
was a popular catalog in English, without Latin descriptions,
authors’ names, and citations, and thus differed from the tech-
nical botanical books of that age. As explained in the intro-
duction (p. viii), the catalog contained Linnaean names and
English names, generic descriptions, and "a plain and familiar
description of the appearance, manner of growth, &c." of the
species and varieties,with notes on the.soil, habitat, and uses.
The book closed with a page devoted to an advertisement stating
that seeds and growing plants were offered at a reasonable rate
by the author.

The arguments for and against Acer saccharum Marsh. as the
name for the sugar maple may be summed up as follows:

AFFIRMATIVE. 1. Marshall in 1785 published the name Acer
saccharum with the common name "sugar maple“ and with a popular,
English description which can be interpreted and accepted as
fitting the sugar maple, at least in part.

2. Technically the sugar maple was then without a scientific
name, as Acer saccharinum L. referred to the silver maple.

3. Positive proof that “saccharum" is a changed spelling
of “saccharinum," whether intentional or accidental, is lack-

ing and probably cannot be obtained.
4. The name Acer saccharum Marsh. is now established in

462 PL A<¥ 20.1, 0,6 -Tea Vol. 2. noe ll
usage, and change of names would create confusion.

NEGATIVE. 1. The popular, English description of Acer
saccharum Marsh.is indefinite. As Marshall left no herbariun,
positive identification of the name cannot be made.

2. In order to account for Linnaeus' four species of maples
native in the United States, Acer saccharum must correspond to
A. saccharinum of Linnaeus. Marshall did not list both names.

3. The name Acer saccharinum was confused at that time and
applied both to the silver maple and the sugar maple.

4. It is highly improbable that a botanist in the year 1785
would have assigned a new specific name almost identical with
the Linnaean name of another species in the same genus and
known from the same region.

5. Positive proof that “saccharum" is a changed spelling
of “saccharinum," whether intentional or accicental, cannot be
offered because Marshall's popular book omitted the technical
details. Authors end citations of previously published names
were not stated, and new species were not indicated.

6. Contemporary authors did not accept Acer saccherum as a
valid name for a new species. Also, in both the French and
German editions of Marshall's book, the translators changed the
spelling to A. saccharinunm.

7. Not until more than a century later, in 1889, was Acer
saccharum finally adopted by another author, one who was making
a revoluticnary attempt to restore old names having priority.
In the meantime other authors, such as Torrey and Gray (1840),
had knowingly passed over the name.

The simplest conclusion from all these lires of evidence is
that Marshall described the sugar maple but that the spelling
“saccharum" was an error for “saccharinum." If Acer saccharum
Marsh. had remained in disuse, would present-day botanists now
revive and accept the name, in view of the above evidence? I
think not. Perhaps Acer saccharum owes its acceptance largely
to the reform movement in which so many names were changed at
the same time.

Acer sacchatum Mill. (Gard. Dict. Abridged. Ed. 6, Acer No.
6. 1771), apparently an error for "saccharinum," can be re-
jected as superfluous when published (Art. 60), because Miller
quoted Linnaeus’ Latin description of A. saccharinum and cited
"Lin. Sp. Pl. 1055." In other editions from 1768 on, Miller
(Gard. Dict. Ed. 8. 1768) used the spelling A. saccharinum
and associated Linnaeus’ name with the sugar maple instead of
the silver maple, as did Wangenheim (Beytr. Forstwiss. Nord-
amer. Holz. 26, pl. 11, fig. 26. 1787) and many later authors.
A. saccharum Marsh. cannot be discarded so readily, because
Marshall did not cite Linnaeus nor even mention authors of

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1948 Little, Nomenclature of Trees 463

previously described names.

Another of Marshall's names revived by Britton as basonym
for the pecan, Juglans pecan Marsh. (Arbustr. Amer. 69. 1785),
has been rejected by Rehder (Arnold Arboretum Jour. 22: 571-
572. 1941), by Little (Amer. Midland Nat. 29: 501-502. 1943),
and by Fernald (Rhodora 49: 194-196. 1947). Anyone verifying
Marshall's Uescription" will see that the name is almost a
nomen nudum. Nevertheless, the name was widely accepted for a
time and now must be discarded.

It seems that a majority of the botanists concerned wish to
retain the widely accepted mame, Acer saccharum Marsh. It cer-
tainly is simpler and less confusing to retein a doubtful name
already in use than to attempt a change. As Gleason (PHYTOLOGIA
2: 203. 1947) has remarked, in all such cases the rules should
be interpreted to fevor the maintenance of a name rather than
ite change. Though my personal choice (Rhodora 46: 445. 1944)
would be A. saccharophorum, I agree that perhaps it is best,
“for the sake of uniformity of nomenclature," to retain Marsh-
ell’s name.

GLYCOSMIS PARVIFLORA (Sims) Little, comb. nov.
CHINESE GLYCOSMIS

Limonia citrifolie Willd., Enum. Fl. Hort. Berol. 448. 1809.
Not Limonie citrifolia Salisb., Prodr. 320. 1796.

Limonie parviflora Sims, Curtis's Bot. Mag. 50: pl. 2416.
1823.

Glycosmis ee (Willd.) Lindl., Roy. Hort. Soc. London

Trans. 6: 72. 1826.

This species, commonly knownas Glycosmis citrifolia (Willd.)
Lindl., is an unarmed, evergreen shrub or srall tree native of
southern China, French Indo-China, and Thailand. It is culti-
vated and naturalized at Key West, Florida, according to Small
(Man. Southeast. Fl. 759. 1933) and Everett (Addisoniea 21: 29.
1940). Everett stated also that it is suitable for cultivation
in the warmer parts of southern United States.

Some authors have included this species in G. pentaphylle
(Retz.) DC., Malay glycosmis. However, in the latest summary
of the genus, Swingle (in Webber and Batchelor, Citrus Industry
1: 157. 1943) maintained the two as distinct.

. Forest Service,
United States Department of Agriculture,
Washington, D. C.

NOTES ON NEW AND NOTEWORTHY PLANTS. V
Harold Ne Moldenke

ALOYSIA LYCIOIDES vare PARAGUARIENSIS (Briqe) Moldenke, comb.
novVe

Lippia ligustrina vare paragueriensis Briqe, Anne Conserve
& iene Bote Geneve 7--8: Ee TOCL-

ALOYSIA IYCIOIDES vare SCHULZII (Steandl.) Moldenke, comb. nov.

Lippie ligustrina vare Schulzii Standle, Field Muse Puble
Bote 4: 2560 19296

DURANTA PARVIFOLIA Moldenke, Spe NOVe

Frutex ramosus; ramis ramulisque graciliusculis rigidis
griseis glabratis, juventute acutiuscule tetragonis, senectute
obsolete tetragonis; hornotinis gracilibus brunnescentibug ad-
presse strigillosis; nodis saepe spinosis;:foliis oppositis
numerosis;'petiolis strigilloso-puberulis; foliis crassis ob-=
lanceolato-ellipticis, ad apicem rotundatis vel obtusis, ad
basin longe acuminatis, conspicue revoluto-marginatis, integ-
errimis vel subintegris, supra glabris, subtus minute puberu-
lis vel glabrescentibus.

Shrub, about 2e5 me tall, abundantly pranched; branches and
brenchlets rather slender, stiff, light-gray, glebrate, rather
acutely tetragonal when young, obsoletely tetragonal in age;
youngest twigs slender, brownish in drying, strigillose with
closely appressed hairs; nodes not amulate but often bearing
a pair of stiff ascending spines less than 1 eme longs, very’
sharp; principal internodes 0.4--2.5 cme long, usually decided-
ly abbreviated; leaves decussate-opposite, abundant; petioles
very slender, 2--5 mne long, strigillose-puberulent; blades
thick-textured, bright-green above, brunnescent in drying,
lighter beneath, oblanceolate-elliptic, 0«8--206 ome long, 5--
11 mme wide, usually rounded or obtuse at the apex, rarely sub-
acute, long-acuminate at base, definitely and conspicuously
revolute-margined, entire or sometimes with a few tiny erect

teeth at or near the apex, glabrous (except for the midrib) and

very shiny above or with a few widely scattered hairs, very
minutely and inconspicuously puberulent along the venation or
glabrous beneath; midrib slender, deeply impressed and short-
strigillose above, very prominent beneath; secondaries 2 or 3
per side, ercuate-ascending, deeply impressed above and very
prominent beneath, anastomosing near the margins; veinlet re-
ticulation subimpressed above when viewed under a handlens, ob-
scure beneath; inflorescence axillary, abundant, opposite, 2--6
cme long, rather few-flowered, nutant; peduncles (1 cme or less
long) end rachis very slender, brunnescent like the youngest
twigs, densely appressed-strigillose; pedicels filiform, 1--6
mme long, usually quite elongated, demsely appressed-strigil-
lose; a few foliaceous bracts sometimes present tovard the base

ae ee

——a eS eo! a ee

1948 Moldenke, New and Noteworthy Plants 465

_ of the racemes; bractlots and prophylla linear, 1--2 mm- long,
densely apvressed-strigillose, brunnescent; calyx tubular, 4--
6e5 mme long, about 2 mme in diameter, densely appressed-pubes-
cent with more or less antrorse whitish hairs, 5-ribbed, 5-
plaited, the rim shortly 5-toothed and S-apiculate; corolla
blue, its tube 8--9 mm. long, densely sordid-puberulent, its
limb 8=--9 mme widee

_ The oo) of this species was collected by Mello sarreto
(noe 11057) in capo, Campo do Faco, Minas Gernes, Brazil, on
November 6, 1940, and is deposited in the Britton Herbarium at
the New York Botanical Garden.

. JUNELLIA CHUBUTENSIS Moldenke, spe nove

Frutex; remis gracilibus griseis vel albidis suberoso-
striatis glabris; ramulis brevissimis stramineis gracillimis
dense pubescentibus; foliis decussatis caducis; petiolis pubes-
centibus; laminis firmis oblongis integris, ad apicem acutis
vel obtusis, plerumque subrevoluto-marginatis, utrinque densi-
uscule breviter pubescentibus; pilis ad basin bulbosis.

Shrub; branches slender, light-gray or almost white, corky-
ridged, glabrous; branchlets apparently very short, stramine-
ous, very slender, rather abundantly pubescent with erect
whitish hairs; nodes not amulate; principal internodes moh
abbreviated, 1=-5 mm long on the branchlets, 2--17 mm. long on
the branches; leaves decussate-opposite, apparently caduccus;
petioles Oe5 mme long, pubescent; blades firm-textured, oblong,
unifomly dark-green on both surfaces, 4--9 mme long, 1:5--4e
mme wide, acute or obtuse at the apex and base, entire, mostly
subrevolute at the margins, rather densely short-pubescent on
both surfaces with erect, stiff, bulbous-based hairs; midrib
prominent beneath, impressed above; secondaries and veinlets
not visible on either surface; inflorescence subcapitate, 1--2
cme long in fruit, about 1-3 cme wide in fruit, dense; bract-
lets and prophylla linear, about 2 mme long, very densely pub-
escent; calyx tubular, about 5 mm. long and 2 mm. wide, very
densely pubescent with gray hairs, S=-toothed, the teeth apicu-
late; corolla-tube about 8 mme long, glabrous, its lobes about
2 ume long, glabrous; fruiting-calyx slightly enlarged, plainly
5-ribbed, densely pubescent; cocci 4, oblong-triquetrous, about
5 mme long, broadly 2-alate \the wings each almost 1 mm wide),
glabrous and nitid above, densely cinereous-strigillose be-
neath, slightly subcucullate at both ends.

the type of this remarkably distinct species wes collected
by Carlos Ae O'Donell at Puerto Madryn, Chubut, Argontina, on
October 24, 1945, and is deposited in the Britton Herbarium at
the New rork Botanical Garden.

JUNELLIA ECHEGARAYI vare CORDIFOLIA Moldenke, vare nove

Haec varistas a forma typica speciei recedit foliis valde
Variabilibus oblanceolato-spathulatis vel ellipticis vel subro-
tundis vel ovatis, ad basin longe attenuatis vel rotundatis vel

cordatis, et sarmentis pedunculisque rhachideque calicibusque
bracteolisque tentummodo puberulise

466 PHYTOLOGIA Vol. 2, now 22

This variety differs from the typical fom of the species in
having its leaves very variable in size and shape, usually 4--8
mne long and 1--4 mm- wide, sometimes oblanceolate-spatulate,
sometimes ellivtic or subrotund or ovate, the base long-attemu-
ate on the narrow leaves but rounded or cordate on the broader
ones, and the pubescence on the twigs, peduncles, rachis,
calyxes, and bractlets merely puberulent.

‘the ‘type was collected by Ce and G. Grandjot (no. 4714) near
Uspallata, at an altitude of 2300 me, Las Heras, Mendoza, Ar-=
gentina, on December 21, 1937, and is deposited in the Herbario
Ruiz Leal, Godoy Cruz, Mendoza.

JUNELLIA ECHEGARAYI vare PUBERULENTA Moldenke, vare nove -

Haec varistas a forma typica speciei samentis pedunculisqe
rhachideque calicibusque bracteolisque tantummodo puberulis
recedite ~

This variety differs from the typical form of the species in
having the pubescence of its twigs, peduncles, rachis, calyxes,
and bractlets merely puberulente

the type was collected by © and Ge Grandjot (no. 4713) in
the close proximity of Uspallata, at en altitude of 2300 me,
Las Heras, Mendoza, Argentina, on December 21, 1937, and is de-
posited in the Herbario Ruiz Leal, Godoy Cruz, Mendozae The
leaf-shape here is as in the typical form and does not show the
striking variation seen in var. cordifolia.

JUNELLIA LIGUSTRINA (Lage) Moldenke, combe nove
Verbena ligustrina Lage, Gene & Spe Nove 18+ 1816-6

JUNSLLIA O'DONELLI Moldenke, spe nove

Frutex humilis caespitosus; caulibus irregularibus griseis
pulverulentis vel glabris; ramis ramulisque numerosis brevibus
sarmentosis densiuscule puberulis; internodis valde abbreviat-
is; foliis numerosis decussatis sessilibus trifidis viridibus
utrinque puberulis valde revoluto-marginatis, ad apicem subu-
lato-acutis; costa supra valde impressa-

Low matted shrub; stems to 3 dme long, irregular, with
flaky bark, pulverulent or glabrescent, gray; branches and
branchlets numerous, short, tufted, rather densely puberulent,
twiggy; internodes much abbreviated, 1--4 mme long throughout;
leaves numerous, decussate-opposite, sessile, often with minia-
ture ones in their axils, trifid almost to the base, bright-
green on both surfaces, the segments narrowly linear, 2--4 mme
long, O+5 mme wide or less, the 2 lateral ones quite divergent
on mature leaves, puberulent on both surfaces, decidedly revo-
lute-margined, subulate-acute at the apex, the midrib inpressed
above and decidedly prominent beneath on each segment; veinlets
indiscermible; inflorescence terminal, few-flowered, usually
2—--5=flowered; calyx tubular, 6=--7 mme long, about 2 mme wide,
puberulent, distinctly 5-ribbed, 5-toothed, the teeth apicu-
late, ciliate; corolla-tube about 9 mme long, glabrous, the
lobes about 2 mme long, bifid at the apex, often reflexed,
Slabrous; stamens equaling the mouth of the corolla-tube.

a a

1948 Moldenke, New and Noteworthy Plants 467

The e of this species was collected by Carlos Ae O'Donnell
(mo. 4000) -- in whose honor it is named -=- at Glier Aike, Santa
Cruz, Argentina, on December 15, 1945, and is deposited in the
Britton Herbarium at the New York Botanical Gardene

LANTANA CAMARA f. PARVIFOLIA Moldenke, f. nov-

vaec forma a forma typica speciei recedit caulibus ramisque
inermibus, internodis 1--2 cme longis, et foliis uniforme
parvioribus.

This fom differs from the typical form of the species in
being of smaller stature, having completely unarmed stems and
branches, with the principal internodes only 1--2 om. long, and
with uniformly smaller leaves, the ovate or rarely suborbicular
often rather obtuse blades only 1--205 ome long and 1--2 cme
wide, the peduncles 1--le5 cme long.

The type was cultivated in the greenhouses of the New York
Botanical Garden from seed collected by Ee Je Alexander and T.
MacDougall (noe 1580) along a roadside near Tehuantepée,
Oaxaca, Mexico, in late May, 1945, and is deposited in the
Britton Herbarium at the New York Botanical Garden. The char-
acters of the plant seem to preed true and so I am constrained
to award it a scientific designation.

LANTANA DINTERI Moldenke, spe nove

Frutex; caulibus ramisque pallide griseis vel albidis glab-
ris; samentis gracilibus obtuse tetragonis leviter puberulis
resinoso-granulosis; internodis plerumque abbreviatis; foliis
decussatis vel approximatis mumerosis; petiolis gracillimis
dense puberulis et resinoso-gramulosis; foliis leviter charta-
ceéis utrinque viridibus lanceolatis, ad apicem rotundatis vel
subacutis, regulariter serrato-dentatis, ad basin acuminatis,
supra minute asperulis, non scabris, subtus densissime resino-
so-punctatis, juventute advresso-puberulo-pulverulentis.

Shrub; stems and brenches decidedly woody, the bark very
light-gray or white, glabrous, fissured; twigs slender, obtuse-
ly tetragonal, lightly puberulent and resinous-granular, green-
ish-stramineous; nodes not plainly ammlate; principal inter
nodes abbreviated, mostly 3--22 mme long or on the larger
branches to 4e5 cme long; leaves abundant, decussate-opposite
or rarely approximate; petioles very slender, 2=--6 mm. long,
densely puberulent and resinous-granular; blades thin-charta-
ceous, bright-green on beth surfaces or slightly lighter be-
neath, lanceolate, 2=-3e8 cme long, 6--14 mme wide, rounded or
subacute at apex, regularly serrate-dentate almost to the base,
acuminately narrowed into the petiole at base, minutely asperu-
lous above but not rough to touch, very densely resinous-
punctate beneath, mimtely appressed=-puberulent-pulveruient on
the venation beneath when immature; midrib very slender, plans
above, prominulous beneath; secondaries very slender, 6 or 7
per side, ascending, hardly arcuate, plane above, prominulous
beneath; vein and veinlet reticulation abundant, usually indis-
cernible above, conspicuous beneath on immature leaves, often
much less so on mature leaves; inflorescence capitate, axillary

468 PHYTOLOGIA Vol. 2, no. 1l

end usually only 1 per axil, shorter than the suvtending leaf;
peduncles filiform, 6--15 mme long, tetragonal, finely puberu-
lent and resinous-granular; heads dense, rater few-flowered,
7--13 mme long, 10=-15 mme wide; bractlets ovate, large, con-
spicuous, closely imbricate, the lowermost often mech larger
and divergermt, 7--10 or more mme long, 3 or mOre mme wide, ac-
uminate at apex, sparsely and minutely strigillose and resin-
ous-gramular, the lowermost often foliaceous, the margins often
subrevolute; corolla white, its tube 5--7 mme long, puberulent
on the outside, the limb to 5 mme widee

the type of this very distinct species was collected by Kurt
Dinter (noe 6823) at Kalksberg, Karibib, Southwest Africa, on
January 12, 1934, and is deposited in the herbarium of the Nat-
urhistoriska Riksmuseum at Stockholme The species is named in
honor of the collector who has done such very noteworthy work
on the flora of Southwest Africa.

LANTANA FUCATA fe ALBIFLORA Moldenke, fe nov-
Haec forma a foma typica speciei corollis albis recedite
This form differs from the typical form of the species in
having pure white corollas. -
The type was collected by Ae Re Cuezzo (noe 918) at Serrazu-
ela, Punta de Sierra, depte Cruz del Eje, Cérdoba, Argentina,
on November 8, 1945, and is deposited in the Britton Herbarium
at the New York Botanical Garden.

LANTANA MICRANTHA vare ARMATA Moldenke, vare nove

Haec varietas a forma speciei typica recedit caulibus dense
armmatis et corollis atropurpureis.-

This variety differs from the typical form of the species in
having the stems abundantly armed with stout hooked prickles
and the corollas dark=purple in color. )

The type was collected by Martin CArdenas (noe 2380) in dry
argillaceous soil on the road to Vacas, above Arami, at an al-=
titude of 2600 me, Cochabamba, Bolivia, in February, 1944, and
is deposited in the Britton Herbarium at the New York Botanical
Gardene The collector describes the plant as a thorny shrub a-
bout 6 dme tall, with dark-purple flowerse

LANTANA MICRANTHA f. VIOLACEA Moldenke, fe nove

Haec forma a forma typica speciei corollis roseis vel pur-
pureis vel rubellis recedit.

This fom differs from the typical form of the species in
having its corollas lilac, pink, purple, rose, or carmine in
colore

The type was collected by Ae Ge Schulz (noe 1459) at the
edge of mountains, Colonia Benitez, Chaco, Argentina, in Novem-
ber, 1935, and is deposited in the Britton Herbarium at the New
York Botanical Gardene The collector describes the plant as a
shrub 2--3 me tall, the foliage with a disagreeable odor, and
the flowers uniformly lilac in color. Other collections with
pink, purple, rose, or carmine flowers, however, had better be
placed with the type in this color-forme

1948 Moldernke, New and Noteworthy Plants 469

LIPPIA AFRICANA Moldenke, spe nove

Frutex; ramis subgracilibus obtusiuscule tetragonis albido-
strigosis densiuscule resinoso-granulosis; foliis decussatis;
petiolis gracillimis strigosis resinoso-granulosis; laminis
chartaceis utrinque viridibus ellipticis, ad apicem et basin
acutis, regulariter adpresso-serratis, supra bulboso-strigosis
et subbullatis, subtus dense breviterque pubescentibus et res-
inoso-gramlosis-

Shrub; branches rather slender, rather obtusely tetragonal,
strigose with whitish antrorse hairs and rather densely resin-
Ous-granular; nodes annulate; principal internodes %--5e8 am.
long; leaves decussate-opposite, usually with a cluster of
smaller ones on much abbreviated twigs in their axils; petioles
very slender, 1=--3 mm long, antrorsely strigose and resinous-
grenular like the branches; blades chartaceous, bright=-green on
both surfaces, elliptic, 2=-3 cme long, 8=--13 mm- wide, acute
at apex and base, regularly appressed-serrate, strigose above
With bulbous-based whitish antrorse hairs and subbullate, dense
ly short-pubescent and densely resinous-granular beneath; mid-
rib very slender, impressed above, prominent beneath; second-
aries very slender, 2--4 per side, arcuate-ascending, impressed
above, promimlous beneath; vein and veinlet reticulation abun-
dant, impressed above, the larger parts promimlous beneath;
inflorescence spicate, abundant, 2 or 3 per node, usually borne
at each of the upper 8 or more nodes, surpassing the subtending
leaves; peduncles slender, 2e5--4 cme long, densely strigillose
and resinous-granular, rarely terminated by two equal divergent
spikes; floriferous spikes subcapitate or elongating to about
12 mm, to 9 mme wide, densely many-flowered; bractlets ovate,
5-5-5 mme long, about 2-5 mm wide, attenuate-acuminate at the
apex, densely strigose and resinous-granular; corolla 5--6 mme
long, usually subequaling the subtending bractlet, the tube
puberulent-gramilar at the apex on the back, the limb 1-5--2
MMe widéee

The type of this species was collected by Erik Wall in a
forest 9 miles west of Nylstroom, at an altitude of 4300 feet,
Transvaal, Union of South Africa, on October 3, 1938, and is
deposited in the herbarium of the Naturhistoriska Riksmuseum
at Stockholme The species has been collected quite often in re-
cent years, but has hitherto been confused with "Le asperifolia
Riche" with which it has usually been identified with a quest-
ion or of which it has been regarded as a form or varistye the
size of the heads and bractlets distinguishes it at once.

LIPPIA AFRICANA vare VILLOSA Moldenke, vare nove
Haec varietas a forma typical speciei recedit remis sarmen-
tisque petiolisque laminisque foliorum bracteolisque villosis-
This variety differs from the typical form of the species in
the villous pubescence on branches, twigs, petioles, leaf-
blades, peduncles, and bractlets, and in its larger flowering-
heads, the heads being to 13 mme wide and the bractlets to 7.

e longe °
5; The type of this variety was collected by Ake Holm (no- 32)

470 PHYTOLOGIA Vol. 2, noe ll

on a steppe on the western side of Mount Elgon, at al altitude
of 2200 me, Uganda, on March 20, 1938, and is deposited in the
herbarium of the Naturhistoriska Riksmseum at Stockholme

LIPPIA LUPULIFORMIS Moldenke, spe nov. .

Frutex; ramulis gracilibus acute tetragonis griseis pustula-
to-asperis; hornotinis dense substrigosis (pilis albidis bul-
bosis) dense resinoso-gramulosis brunneis; sterignis foliorum
elevatis perspicuis divergentibus; alabastris dense villosis;
foliis decussatis vel ternatis vel approximatis; petiolis den-
siuscule substrigoso-villosulis resinoso-granulosis marginatis;
foliis chartaceis ovatis, ad apicem rotundatis vel subacutis,
ad basin acutis vel acuminatis vel rotundatis, supra valde
scabris bullatis, utrinque dense breviterque pubescentibus,
subtus dense resi noso-granulosis, uniforme serrato-dentatise

Shrub to 1e5 me tall; branchlets slender, acutely tetragonal
and grayish, pustulate=-asperous, the younger parts and twigs
rather densely substrigose with whitish bulbous-based hairs and
densely resinous-granular, brownish; leaf-scars elevated on
conspicuous divergent sterigmata; buds densely white-villous;
nodes annulate on young twigs, not anmlate on older branches;
principal internodes 1]e2--3e2 cme long on older branchlets, ab-
breviated to 2--9 mme on twigs; leaves decussate-opposite or
rarely ternate or approximate; petioles slender, l--5 mm. long,
rather densely substrigose-villosulous and resinous-granular,
margined, deeply canaliculate above; blades chartaceous, dark-
green above, lighter beneath, ovate, 1e7--3e5 cme long, 0-9--2
cme wide, rounded or subacute at apex, acute or acuminate at
base or sometimes rounded, very scabrous and bullate above,
densely short-pubescent on both surfaces, densely resinous-
grenular beneath, uniformly serrate-dentate from base to apex,
the teeth often decidedly revolute-margined; midrib impressed
above, prominulous beneath; secondaries very slender, about 5
per side, ascending, hardly arcuate, impressed above, promim-
lous beneath; vein and veinlet reticulation abundant, usually
deeply impressed above and promimulous beneath; inflorescence
spicate, axillary, surpassing the subtending leaf, 1 or 2 per
node; peduncles slender, 4—-665 cme long, asperous-hirsutulous
and resinous=-gramlar, tetragonal; spikes at first capitate,
later elongating to 2 cme, le5<=-2 cme wide, strobilifom,
densely many-flowered; bractlets large and conspicuous, densely
imbricate, ovate, 8=--10 mme long, 4--7 mme wide, acute or sub-
acuminate at apex, rather sparsely strigillose and granular;
corolla=-tube 6=—-7 mme long, densely gray-pubescent on the out-
side, the limb 4--5 mme wide.

the type of this species was collected by He Rudatis (noe
1145)-at G'Mermum, at an altitude of 650 me, Dumisa, district
Alexandra, Natal, Union of South Africa, on september 11, 1910,
and is deposited in the herbarium of the Naturhistoriska Riks-
museum at Stockholme

LIPPIA VIOLACEA Moldenke, sp- nove
Frutex; ramis stramineis vel brunnescentibus obtusiusocule

Siaiiald

1948 Moldenke, New and Noteworthy Plants 471
tetragonis breviter pubescentibus, pilis uncinatis ad basin
bulbosis; remulis donsissime pubescentibus virgatis; foliis de-
cussatis numerosis; petiolis gracillimis dense cinereo-pubes-
centibus; foliis ellipticis vel ovatis chartaceis, ad apicem a-
cutis vel obtusis, ad basin acutis, serrulatis, supra dense
puberulis st resinoso-punctulatis, subtus dense breviterque
pubescentibus et plusmimsve resinoso-punctulatise

ohrub, about 2 me tall; branches stramineous or brunnescent,
rather obtusely tetragonal, short-pubescent with uncinate
bulbous-based hairs; branchlets much more densely pubescent
with sordid-gray or cinereous hairs, virgate; nodes annulate;
principal internodes 1--4e8 cme long; leaves decussate-oppo-
site, numerous; petioles very slender, 1=-4 mme long, densely
cinereous-pubescent; blades more or less diamond-shaped or el-
liptie, varying to ovate, chartaceous, le3--205 cme long, 9--15
mme Wide, acute or obtuse at apex, regularly serrulate from be-
low the middle to the apex, acute at base, densely puberulent
and resinous-punctulate above, very densely short-pubescent and
more or less resinous-punctulate beneath; midrib very slender,
subimpressed above, prominulous beneath; secondaries very slen-
der, 4 or 5 per side, ascending, not arcuate, often subimpress-
ed above, subpromimulous beneath, not at all anastomosing, each

_ secondary or one of its branches ending in a sinus between two

teeth; veinlet reticulation obscure or subimpressed above,
plane beneath; inflorescence axillary, capitate, 2 per node,
1-5--2 cme long, mostly about equaling the subtending leaves;
peduncles very slender, about 1 come long, densely short-pubes-
cent with rather appressed sordid-cinereous hairs; heads dense-
ly flowered, hemispheric, 1--1e5 cme wide; bractlets lanceo-
late-lingulate, about 5 mme long and 1e5 mme wide, strigillose;
calyx about 5 mme long, densely white-hirsute especially on the
margins and resinous-gramlar; corolla vioiet, its tube 5--6
mme long, rather scattered=-strigillose or puberulent above the
calyx, its limb about 5 mme wide.

The e of this species was collected by Mendes Magalhaes
(no. 1768) on the campo between Capivari and Pico do Itambé,
Serra Quebrada, munic{pio Serro, Minas Geraes, Brazil, on May
3, 1942, and is deposited in the Britton Herbarium at the New
York Botanical Gardene

PAEPALANTHUS STANDLEYI Moldenke, sp- nove

Herbas caule valde abbreviato; foliis rosulatis mumerosis
Viridibus nitentibus ligulatis obtusis firmis satis revoluto-
marginatis, sudtus parcissime obscureque puberulis, supra pul-
verulentis, mlto-nervatis; pedunculis numerosis crassiusculis
sexangulatis subcompressis densissime villosis-

Herbs stem mich abbreviated; leaves tufted, numerous,
bright-green, shiny, strap-shaped, broadest at the pase, 6--8
cme long, 8=--9 mme wide at the middle, obtuse at the apex,

fim-textured, somewhat revolute-margined especially toward

the apex, very finely and usually obscurely puberulent beneath,
pulverulent above, many-nerved; peduncles 15 or more per plant,
rathor stout, 27=--30 ome long, 6<angled, somewhat flattened,

472 P iH Y-T OB OG <A Vol. 2, no. li

very densely and persistently villous from vase to apex with
uniform fulvous-brunneus hairs; sheaths cylindric, usually a-
bout equaling tne leaves, 6--te5 cme long, rather closely ap-
pressed to the peduncle, many-costate, rather densely pubescent
with spreading brovmish hairs, obliquely split at the apex, the
blade lanceolate, about 1 cm- long, erect, appressed to tne pe-
duncle; heads obovate=hemispheric, l--le2 cm» high, 2—-2.3 cm.
wide, very suowy; involucral bracts broadly ovate, leathery,
deep chestmt-browm and very shiny on voth surfaces, 7--9 mme
long, 4=-5 mme wide at the base, long-attenuate or supacuminate
at the apex, in 4 or 5 series, the outermost ones densely tom-
entellous or subvillous, the inner ones white-villosuious on
the lower half and pulverulent above, long-ciliate with white
hsirs on the margins; receptacle very densely and conspicuously
white-villous with mlticellular white hairs about 5 mme -long;
recepteacular bractlets numerous, linear, 6—-6-5 mm long, about
Oo5 mne wide, dark-brown on the upper half, light-brown tovard
the base, densely villous=barbellate on the inner surface at
and near the apex; staminate flowers apparently of two types:
in the one type tne sepals are 3, connave only at the very base,
stramineous, oblanceolate, about 4 mme long and 1] mme wide,
triangular-acute at the apex, glabrate on both surfaces except
for a few, crect, irregular, translucent hairs on the margins,
densely villous=-barbellate at the apex with straight, erect, —
white hairs extending about 4 mne beyond the apex of the sepals;
petals 3, about 2 mme long, connate for about half their length
into a tube about 0-7 mme wide, brownish, the free apex triang-
ular-acute, about 0.5 mme long, densely long-ciliate with regu-
lar, orect, white hairs; stamens 3; filaments Oce6 mme long; an-
thers oblong, about O«8 mme long and 0«4 mme wide; style-vest-
iges 3, club=-shaped, about O«c6 mme longs in the second type the
corolla-tube is 4--5 mm long, subhyaline, densely villous
within, often invaginated at the apex, tne filaments elongated
to 6 mm; pistillate florets: sepals 3, separate to the base or
practically so, oblong, stramineous, about 4 mme long and 009
mme wide, l-ribbed, glabrate on both surfaces except for the a-
pex which is long-barbellate with a dense tuft of erecty
straight, white hairs extending le5 mme beyond the apex of the
sepals; petals 3, separate to the base, elliptic, about 4e]1 mme
long and le5 mme wide, acute or rarely retuse at the apex,
stramineous, glabrate on both surfaces and shiny except for the
rather unifomly white-ciliate margins end epex, not barbell-
ate; staminodes 3, about 1e5 mme longs pistils 3; styles gleb-
rous, 5--5e5 mme long, forked at the apex, the branches about 1
mm- long; ovary 3=-lobed, 3-sulcate, 3-celled, glabrous.

The type of this extremely handsome and distinct species was
collected by Mello Barreto (no. 9688) in a sandy campo at Bia
Vusta - ExtraccHo, municfpio Diamantina, Minas Geraes, Brazil;
on November 9, 1937, and is deposited in the Britton Herbariun
at the New York Botanical Garden. It was determined by the col-
lector of Pe plumosus (Bonge) Kérne, put differs conspicuously
from that species in its mech larger staturee The species was
regarded, on the basis of a Mexia collection, by Dre Pe Ce

ee

eT
7 ¥ as : » x

1948 Moldenke, New and Noteworthy Plante 473

Stendley as Pe Warmingiams (Kore) Kom-, which, however, dif-

fers in its smooth peduncles, less attenuated involucral bract-
lets, and floral characterse

SABINA VIRGINIANA var. CREBRA (Fernald & Griscom) Moldenxe,
combDe nove
Juniperus Virginiana vare crebra Fernald & Griscom, Rhodora
37: 133, ple 3330 19356

STACHYTARPHETA PARAGUARIENSIS Moldenke, sp. nov.

Frutex l--2 me altus; ramis gracilibus acute tetragonis
saecpe marginatis dense breviterque pubescentibus, hornotinis
velutinis; ramulis sarmentisque densissime velutino-puoescen-
tibus cinereis; petiolis obscuris latissime alatis; laminis
submembranaceis utrinque atroviridibus in siccitate brunnes-
centibus, ovatis, at apicem acutis vel obtusis, ad basin longe
acuminatis, crasse serratis utrinque dense breviterque pubes-
centibus; corollis coeruleise

Shrub, l==-2 me tall; branches slender, acutely tetragonal,
often margined, densely short-pubescent with cinereous hairs,
velutinous on the younger parts; branchlets and twigs similar
to the pranches but even more densely velutinous~pubescent,
cinereous; nodes annulate; principal internodes le3--8e4 cme
long; leaves decussate-opposite, often with several smaller
ones in their axils; petioles obscure, 3--15 mme long, very
broadly winged, indistinguishable from the leaf=base; blades
submembranous, dark-green on both surfaces, brunnescent in dry-
ing, ovate, 3--7e5 cme long, le3--302 cme wide, obtuse or acute
at apex (in outline), long-acuminate into the petiole at base,
coarsely serrate from below the middle to tne apex with broadly
triangular acute or subapiculate teeth, densely short-pupescent
on both surfaces, subvelutinous on tile midrib and secondaries
beneath when immature, the hairs canescent or cinereous; midrib
slender, flat above, promimulous beneath; secondaries very
slender, 4--6 per side, ascending, hardly at all arcuate, flat
above, subpromimulous veneath; veinlet reticulation obscure,
except for the larger tertiaries beneath; inflorescence spic-
ate, terminal, to about 20 cme long, closely many-flowered;
flowers imbricate; rachis rather slender, densely canescent=
puberulent, rather deeply sculptured efter anthesis; peduncles
obsolete or to 2 ame long and densely canescent~velutinous;
bracts lanceolate, about 5 mme long, 1--le3 mme wide, long=
attenuate to the subacuminate apex, cinereous-puberulent, usu-

ally densely white-ciliate on the margins (especially when

young); calyx tubular, about 8 mme long, 1—-1le3 mme wide,
densely short-pubescent with brownish hairs, 5-ribbed, the rim
5-toothed, the teeth micronate, almost 1 mme long; corolla
sea-blue, its tube about 1 ome long, obscurely pulverulent-
puberulent or glabrescent above the calyx, its limb about 1 om.
wide, glabratee

The type of this distinct species was collected by Teodoro

ojas (noe 13615) on hillslopes among small trees at Fuerte
Bepas, (ne the ee region oF Paraguay on October 18, 1946, and

474 Pel Vit Ove GS Vol. 2, noe ll

is deposited in the Britton Herbarium at the New york Botanical
Garden.

STILBE VERTICILLATA (Ecklon & Zeyher) Moldenke, combe nove
Trichocephalus verticillatus Ecklon & Zeyher, Emme Ple Afre
Austre 131le 183 e

STILBE VERTICILLATA vare CUSPIDATA (He He We Pearson) Moldenke,
combe nNoVe
Stilbe mucronata vare cuspidata He He We Pearson in Thisel-
ton-Dyer, Fl. Cape 5: 184¢ 1901.

TITHYMALOPSIS IPECACUANHAE fe ORBICULATA Moldenke, fe nove —

Haec forma a forma typica speciei foliis orbiculatis virid-
ibus recedit.

This fom differs from the typical fom of the species in
its green orbicular leaf=bladese

The type was collected by He Ne Moldenke (no. 10476) in
sandy sai] along a roadside at Smithtown, Suffolk Coe, New
York, on May 29, 1938, and is deposited in the Britton Herbar~
ium at the New York Botanical Gardene The form propably cor=
responds, in part, at least, to Boissier's "Euphorbia Ipecacu-
anha vare portulacoides", but anyone who has observed these
plents growing Will agree tnat it deserves only form ranke

VERBENA KUNIZEANA Moldenke, spe nOve

Herba; caulibus basin versus saepe decumbentibus; ramis
graciliusculis obtuse tetragonis saepe suleatis dense hirsut-
lo-puvescentibus griseo-sordidis, pilis plerummque glandulifer-
is; petiolis indistinctis late alatis; laminis chartaceis ova-
tis, ad apicem acutis, ad basin acuminatis, crassiuscule regu-
lariterqw serratis utrinque dense hi rsutulo-pubescentibus,
pilis supra plerumque bulbosis-

Herb, several-branched from the base, the lower part of tne
stems often decumbent; stems and branches ratner slender, ob-
tusely tetragonal, the sides often suicate, densely hirsutu-
lous=-pubdescent with widely divergent, grayish-sordid, mostly
gland=tipped neirs; nodes annulate; principal internodes 2--5
Cme long; leaves decussate-opposite; petioles rather obscure,
broadly winged and merging into tne leaf-base; biades chartace~-
ous, bright-green above, paler beneath, ovate, 2e5=-5 ome long,
le4--2 ome wide, acute at apex, acuminate into the winged pet-
iole at base, rather coarsely put regularly serrate from the
widest part to the apex, the teeth rather broadiy triangular,
obtuse or subacute, densely hirsutulous=-pubescent on both sur-
faces, the hairs often slightly pulbous-based above; midrib
slender, impressed above, promimulous beneath; secondaries
slender, 3--5 per side, ascending, impressed above, prominu-
lous beneatns veinlet reticulation plane or subimpressed above,
tne larger parts subprominulous beneath; inflorescence termin-
al, subcapitates; peduncles slender, 6=-8 cme long, tetragonal,
sulcate, densely glendular-hirsutulous; floriferous portion of

1948 Moldenke, New and Noteworthy Plants 475

tue inflorescence about 365 cme long, to 205 ome wide in an-=
thesis; practlets elongated, linear-lanceolate, 11--13 mme
iong, densely glandular-nirsutulous with wnitish nairs, often
somewhat ancurved after the flowers have fallen; rachis densely
glandular-hirsutulous; calyx tuvular, the tube about 15 m.
long and 2 mme wide, densely glandular-hirsutulous with whitish
hairs, the rim 5-apiculate, the apiculations linear-caudate,
unequal, densely glandular-nirsutulous with wnitish hairs, tne
longest ones about 4 mme long; corolla-tube 15=-20 mme long,
densely short-pubescent on the outside, the limb about 10 mme
wide, minutely puverulent on the outer surface, glabrous within.
the type of this distinct species was collected by Carl Axel
Magnus Lindman (noe Ae3649) at Paraguarf, Paraguay, in October,
1893, and 1s deposited in the nerparium of the Naturnistonska
Rixksmuseum at Stockholme The collector notes that the native
neme is "flor de vovia"e It was first identified by Briquet as
Ve paraguariensis vare latiuscula Briqe and then as Ve platen-
sis Sprenge It is named in honor of Carl Ernst Otto Kuntze,
Who has done such splendid collecting of Verbenaceas and relat=
ed groups in Asia, Africa, and America, whose keen insight has
devected so many varieties and forms wortny of nomenclatural
rank, and who fought so eloquently and neroically for fair play
and honesty in botanical nomenclature, albeit a losing battlee

VERBENA MAIMII Moldenke, spe nove

Frutex usque ad le7 me altus; ramis ramulisque viridibus
acute tetragonis ubique glaberrimis nitidisque saepe sulcatis;
foliis minutis indistinctis sessilibus oblongis vel linearibus
firmis utrinque viridibus decussatis adscendentibus utrinque
adpresso-strigillosis, ad apicem acutis l-nervatis.

Shrub to about le7 me tall, much-branched; branches and
branchlets green, acutely tetragonal, completely glavras
throughout, shiny, often sulcate between the angles; twigs
numerous, ascending-erect, slender, acutely tetragonal, green
end shiny, glabrous; principal internodes 2=-5 cme Long; nodes
noy annulate; leaves very tiny and indistinct, giving the
plent en aphyllous appearance, sessile, oblong or linear, rath-
er firm-textured, uniformly bright-green on both surfaces, de-
cussate-opposite, ascending, 3--8 mme long, about 1 mme wide,
appressed-strigillose on both surfaces, acute at apex, l=
nerved, tne midrib slightly elevated beneath and subimpressed
above; inflorescence spicate, very abundant, usually in grows
of 3 at the tip of each twig, often aggregted in paniculate
fashion, the terminal spike usually short-pedunculate, the Lat-
eral ones longer~pedunculate, the floriferous portion elongat-
ing to almost 3 cme after anthesis; peduncies very slender,
glavrous or minutely strigillose, 3--15 mme long, tetragonal,
green; rachis retuer densely strigillose=puverulent with wnit-
ish neirs especially visible after tne calyxes have fallen off;
bractlets lanceolate, very small and obscure, about 1 mme long,
strigose with appressed antrorse whitish hairs, acute at apex;
calyx tubular, about 3 mme long, densely white-strigose with

476 PHYTOLOGIA Vol. 2, no. ll

appressed antrorse nairs, the rim 5-apiculate; corolla blue,
its tuve about 4 mme long, glabrous except at the very apex
where it is densely white-strigose like the calyx, its limb 3--
4 mm wide, puberulent in the throat within end strigose at the
base outside, the lobes glabrous on both surfacese

the type of this distinct species was collected by Gustef
Uskar Andersson sialme (noe 1141) -- in whose nonor it is named
-- in a swamp at Villa Rica, nio Grande do Sul, Brazil, on Jan-
uary 22, 1902, and is deposited in the herparium of the Natur-
historiska Riksmuseum at Stockholm.

Horbas caulibus procumbentibus vel adscendentibus simplic-
ibus argute vetragonis sulcatis scabris; foliis decussatis;
petiolis gracilibus sparsiuscule strigosis marginatis; laminis
chartaceis ellipticis vel anguste ianceolatis vel oblanceolatis
irregulariter dentatis, ad apicem acutis, ad basin attenuatis
vel acuminatis, supra scabrido-strigillosis, subtus molliter
breviterque pubescentibus, pilis ad basin bulbosis; corolla
pulchrae —

Herb; stems procumbent or ascending, several, not branched,
sharply tetragonal, sulcate between the angles, scabrous with
meny short reflexed hairs; principal internodes 3-=8 cme long;
nodes more or less anmilate; leaves numerais, decussate=-oppo-
site, usually with clusters of smaller ones in their axils;
petioles slender, 5--8 mme long, rather sparsely strigose with
short antrorsely curved hairs, margined; blades chartaceous, -
somewhat lighter beneath, elliptic or narrow-lanceolate, vary-
ang to oblanceolate, le5<-5e5 ome long, 6--15 mme wide, acute
at apex, attenuate or acuminate at base, rather irregularly
dentave from the apex almost to the base with ratner broadly
triangular acute teeth, slightly scabridous-strigillose above,
rather softly short-pubescent beneath with bulbous=based hairs;
midrib very slender, plans above, promimlent beneath; secon-
daries very slender, 4 or 5 per side, plane above, prominulent
beneath; veinlet reticulation mostly indiscernible above, con~
spicuous (but not elevated) beneath; inflorescence terminal and
in the uppermost axils, the floriferous portion very dense, at
first flattened~supcapitate, later elongating to 3 cme but re=
maining very dense; peduncles rather stout, 6—-8e5 ome long,
tetragorial and sulcate like the stems, reflexed=-pilose and more
or less: rough to the touch; bractlets lanceolate, 4==-9 mme
long, l--le4 mme wide, attenuate-acute or subacuminate at the
apex, glabrous (or very sparsely pilosulous) except for the
iong-ciliate margins; calyx tubular, 10--12 mme long, about 1
mme wide, 5-costate, purplish especially toward the apex, oili-
ate=-pubescent on the ribs, the teeth unequal, l--2 mme long,
filiform-eppendaged; corolla showy, its tube 15=-17 mme long,
very sparsely and minutely puberulent or merely pulverulent
outside above the calyx, its limb to 15 mme wide, pilose in the
throate

The type was collected by Per Karl Hjalmar Dusén (noe 9334)
in wet thickets at Calmon, Paran&, Brazil, on March 13, 1910,

*

1948 Moldenke, New and Noteworthy Plants 477

end is deposited in the herbarium of the Naturhistoriska Riks-
museum at Stockholme The calyxes on the type specimen are
mostly infested with a fungus which forms tiny brovm dots over
the surface, which dots under the microscope reveal themselves
to be masses of tint brown filaments.

VERBENA PULCHRA vare PALUDICOLA Moldenke, vare nov.

liaec varietas a fonna tynica speciei recedit laminis folior-
um tantummodo 2e4--4e3 om- longis, l--2 cme latis et corollis
albis oculiroseis.

This variety differs from the typical form of the species in
its leaves being shorter, the blades only 2-4=-4e3 on. long,
1l--2 cme wide, and the corollas white oxcept for a red “eye's

The type was collected by Wilhelm Gustav Herter (noe 99937)
in a rather wet sandy marsh, at an altitude of 150 me, exposed
to the sunlight in an arroyo, Reanqueras, Rivera, Uruguay, be=
trreen March 24 and 27, 1907, and is deposited in the Britton
Hervarium at the New York Botanical Garden.

VITEX RUFESCENS vare ABLUDENS (Moldenke) Moldenke, combe nove
Vitex Perriana vare abludens Moldenke, Alphe List Common
Names Verbenace 21, hyponym (1939); Trope Woods 64: 39-6 1940.

VITEX SPRUCEI vare LONGIDENTATA (Moldenke) Moldenxe, combe nove
Vitex spongiocarpa vare longideiitata Moldenke, Phytologia 2:
31. 1941. —

VITEX TRIFOLIA vare SIMPLICIFOLIA fe ALBIFLORA (Ye Matsumura)
Moldenke, combe nove
Vitex rovwundifolia vare albiflora Ye Matsumura, Amatores

Herbarii 10: O4e 1943.

THE KNOWN GEOGRAPHIC DISTRIBUTION OF THE MEMBERS OF THE
VERBENACEAE, AVICZNNIACEAE, STILBACEAE, AND SYMPHOREMACEAE.
SUPPLEMENT 9

Harold Ne Moldenke

Since the preparation of the eighth supplement to this list
several thousand additional specimens of these groups have teen
exemined from the herbarium of the Chicago Natural History Mus-
eum, the United Staves National Herbarium at Washington, the
University of Massachusetts at Amherst, the United States Field
Station at Sacaton, Arizona, the Britton Herbarium of the New
York Botanical Garden, the Jardin Botanique de l'Etat at Brus-
sels, the Botanisk Museum of the University of Lund at Lund,
Sweden, the Naturhistoriska Riksmuseum at Stockholm, and the
Herbario Ruiz Leal at Godoz Cruz, Mendoza, Argentinas this ex-
cellent material has brought to light 74 new country or island

478 PB: HY. T<Oob-0.6 an Vol. 2, no. ll

records, 125 new state, province, or department records, and 17

. new county records, as well as the necessity for making certain

emendations in previous records and certain nomenclatural
changes hereinafter notede
UNITED STATES OF AMERT CAs
Massachusettss
Verbena hastata fe albiflora Moldenke (Worcester County)
Verbena urticifolia Le (Franklin & Hampshire Counties)
Comecticuts
Verbena officinalis Le (New Haven County)

New York:
Verbena simplex Lehme (New York County)
Floridas
Lantana pamaxs vare mista (Le) Le He Bailey (Charlotte
County
Illinois:
xVerbena Perriana Moldenke (Menard County)
Towa:
Verbena bracteate. Lage & Rodre (Lim County)
Missouri:

xVerbena moechina Moldenke (Jefferson County)
Texast
Change "Aloysia ligustrina (Lage) Small" to read Aloysia
lycioides Chame and add Webb County
Change “Aloysia ligustrina vare Schulzii (Standl.) Molden-
ke" to read Aloysia lycioides vare Schulzii (Stendle)
Moldenke and add Nueces County
Phyla nodiflora (Le) Greene (Jefferson County)
New Mexicos
Change "Aloysia ligustrina (Lag-) Small" to read Aloysia
lycioides Chame
Arizonas
Change "Aloysia ligustrina vare Schulzii (Standl.) Molden-
ke" to read Aloysia lycioides vare Schulzii (Standie)
Moldenke arenes a, oa
Lantana horrida HeBeKe core County)

Verbena ciliata vare ra (Greene) Perry (Apache County)
Gray eo

Vorbona Wrigntii Ae have County)
Oregons
Verbena lasiostachys vare septentrionalis Moldenke (Linn
~ County
Californias

mee 2 nodiflora vare rosea (De Don) Moldenke (Marin County)

po aes "Aloysia ligustrina (Lage) Small" to read Aloysia
lycioides Chame
Change Ahoyate ligustrina vare Schulzii (Standle) Moldenke"

to read Aloysia lycioides vare Schulzii (Standl.) Molden-
ke

1948 Moldenke, Geographic Distribution 479

Avicemia bicolor Standl- (Chispas)
Lantana Camara vare aculeata (Le) Moldenke (Veracruz)
Lantana Camara fe parvifolia Moldenke (Oaxaca)*
Lantena hispida HeBeKe (Puebla)
Phyla nodifiora (Le) Greene (Tamaulipas)
Verbena teucriifolia vare corollulata Perry (México)
BERMUDA:
Avicennia nitida Jacq. (Main)
Callicarpa americana Le (Main)
Citharexyium spinosum Le (Main)
Clerodendrum aculeatum (Le) Schlechts (Main)
Clerodendrum fragrans vare pleniflorum Schaue (Main)
Clerodendrum glabrum Ee Mey. (Mein)
Duranta repens Le (Main)
Lantana Camara Le (Main)
Lantana Camara vare aculeata (Le) Moldenie (Main)
Lantana Camara vare mista (Le) Le He Bailey (Main)
Phyla nodiflora (Le) Greene (Main) 7
Phyla nodiflora vare tans (HeBeKe) Moldenke (Main)
Stachytarpheta jemaicensis (le) Vahl (Main)
Verbena officinalis Le (Main)
Verbena rigida Sprenge (Main)
DOMINICA:
Clerodendrum umbellatum vare speciosum (Dombrain) Moldenke
COLOMBIA:
Aegiphila longifolia Turoze (Méta)

Duranta Sprucei vare columbiensis Moldenxe is tne correct
orthography of this name
Lantana Camara Le -- delete "Antioquia"
VENEZUELA:
Phyla nodiflora vare reptans (HeBeKe) Moldenke (Anzoategui)
Priva lappuiacea (Le) Perse (Anzoategui)

SURINAM:
Lantana ocujabensis Schaue
ECUADOR:

Clerodendrum fragrans vare pleniflorum Schaue (Los Rios)
Cormtia odorata (Posppe & Endl-) Poeppe (Los Rfoa)
Lippia alba (Mill-) Ne Ee Bre (Los Rfos)

Stachytarpheta cayennensis (Le Ce Riche) Vahl (Los Rios)
Verbena iitoralis HeBeKe (Santiago-Zamora)
PERU:

Verbena peruviana (Le) Britton
BRAZIL:

Aegiphila Hassleri Briqe (Rio Grande do Sul)

Aloysia ligustrina (Lage) Small should be changed to read
Aloysia iycioides Chame

Aloysia ligustrina vare paraguariensis (Briq-) Moldenke
shouid be changed to read Aloysia lycloides vare para~

480 PPE ET Ot Oe tas Vol. 2, noe il

guariensis (Briqe) Moldenke 7

Aloysia Sellowii (Briqe) Moldenke (Rio Grande do Sul)

Bouchea, fiuminensis vare pilosa Moldenke (Minas Geraes)

Duranta parvifolia Moldenke (Minas Geraes)*

Lantana canescens vare integrifolia Moldenke (Minas Geraes)

Lippia violacea Moldenke (Minas Geraes)*

Stachytarpheta australis Moldenke (Santa Catharina)

Verbena Malmii Moldenxe (Rio wrande do Sul)*

Verbena pulchra Moldenke (Rio Grande do Sul)*

Vitex amazonica Moldenxe is to be deleted

Vitex brasiliensis Steude is to ve deleted

Vitex Perriana Moldenke is to ve deleted

Vitex Perriana vare abludens Moldenxe is to be deleted

Vitex rufescens Ae le Jusse (Bahia, Parahyba, Pernambuco, &
Piauny

Vitex rufescens vare abludens (Moldenke) Moldenke (Bahia &
Pernambuco) *

Vitex spongiocarpa Ducke is to be deleted

Vitex spongiocarpa vare longidentata Moldenxe is to be delet-
ed

Vitex Sprucei vare longidentata (Moldenke) Moldenke (Amazonas)

BOLIVIA:
Aloysia ligustrina (Lage) Small is to be changed to read

Aloysia lycioides Chame

Aloysia ligustrina vare paraguariensis (Briqe) Moldenke is to
be changed to read Aloysia lycioides Vare paraguariensis
(Briqe) Moldenke

Lantana Fiebrigii Hayek (Senta Cruz)

Lantena glutinosa Poeppe (Santa Cruz) |

Lantana micrantha Briqe -- delete "Cochabamba" and add Santa
Cruz

Lantana micrantha vare armata Moldenize (Cochabamba)*

Lantana micrantha fe violacea Moldenke (Santa Cruz)

PARAGUAY:
Aloysia ligustrina (Lage) Small should be changed to read

Aloysia lycioides Chame
Aloysia ligustrina vare paraguariensis (Brige) Moldenke is to

changed to read Aloysia lycioides vare paraguariensis
(Briqe) Moldenke

Lantana micrantha fe violacea Moldenke
Stachytarpheta paraguariensis Moldenke*
URUGUAY :
Aloysia ligustrina (Lage) Small should be changed to read
Aloysia lycioides Chame
Aloysia ligustrina vars paraguariensis (Brige) Moldenke is to

be changed to read Aloysia lycioides vare paraguarioensis
(Brigqe) Moldenke |

Lantana, hypoleuca Brige

1948 Moldenke, Geographic Distribution 481

Verbena bonariensis vare conglomerata Brige
ARGENTINA:

Aloysia chacoénsis Moldenke (Catamarca)

Aloysia ligustrina (Lage) Small should be changed to read
Aloysia lycioides Chame end add Corrientes

Aloysia ligustrina vare paraguariensis (Briqe) Moldenke is to
changed to read Aloysia lycioides vare paraguariensis
(Brige) Moldenke and add Cérdoba & Tucumén

Aloysia Sellowii (Briqe) Moldenke (Corrientes & Mendoza)

Junellia chubutensis Moldenke (Chubut)*

Junellia Echegarayi vare cordifolia Moldenke (Mendoza)*

Junellia Echegarayi vare puberulenta Moldenke (Mendoza)*

Junellia erinacea (ci. & Hooke) Moldenxe (Chubut)

Junellia Lorentzii (Niederiein) Moldenke should be changed to
read Junellia ligustrina (Lage) Moldenke

Junellia O'Donelli Moldenke (Santa Cruz)*

Juneilia succulentifolia (Kuntze) Moldenke (Neuquen & Rio
Negro

caltecn aristata vare angustifolia (Kuntze) Moldenke (Chaco)

Lantana Balansae Brige (Fomosa)

Lantana Camara vare aouleata (L-) Moldenke (Tucum4n)

Lantana Fiebrigii Hayek (Fomosa) |

Lantana fucata Lindle (Cérdoba & Corrientes)

Lantana fucata fe albiflora Moldenke (Cérdoba)*

Lantana glutinosa Pospp. (Corrientes & Entre Rios)

Lantana Grisebachii Stucke (Catemarca, Entre Rfos, La Rioja,
& Salta

Lantana hypoleuca Brige (Entre Rios & Misiones)

Lantana Junelliana Moldenke (San Luis)

Lentana micrantha Briqe (Formosa)

Lantana micrantha fe violacea Moldenke (Chaco, Corrientes, &
Salta

Lantena montevidensis (Sprenge) Briqe (Entre Rios)

Lantana tiliaefolia Cheme (Misiones & Tucumfn)

Lantana xenica Moldenke (Cérdoba « Mendoza)

Lippia alba (Mille) Ne Eo Bre (Formosa & San Juan)

Lippia angustifolia Chame (Corrientes)

Lippia asperrima Cheme (Corrientes & Formosa)

Lippia tegulifera Brige (Corrientes)

Lippia turbinata Grisebe (San Juan)

Lippia turbinata fe angustifolia Osten ( Senta Fé & Santiago
del Estero

Phyia nodiflora (le) Greene (Catemarca, Chaco, Cérdoba, Entre
Rios, Jujuy, San Juan, San Luis, Mendoza, Santa Fé, &
Sentiago del Estero)

Phyla nodiflora vare canescens (HeBeKe) Moldenke (Rfo Negro)

Phyla nodiflora vare reptans (HeBeKe) Moldenke (Formosa, San

4go PHYTOLOGTIA Vol. 2, no. ll

Juan, & Santiago del Estero)
Phyia nodiflora vare rosea (De Don) Moldenke (Entre Rios &
Formosa
Stachytarpheta oayennensis (Le Ce Riche) Vahl (Corrientes)
erbena carollata Brige (Cérdoba)
Verbena moricolor Moldenke (Catamarca)
Verbena Parodii (Covas & Schnack) Moldenke (Tucumfn)
Verbena peruviana (Le) Britton (Catamarca & Tucumfn)
Verbena scrobiculata Grisebe (Santiago del Estero)
Verbena tenuisecta Briqe (Catamarca)
WALES:
Verbena officinalis Le
ANGLO -EGYPTIAN SUDAN:
Clerodendrum Wallii Moldenke (Nuer)
ERITREA: ot
Lantana vibumoides (Forske) Vahl is the correct accredition
of this name
ABYSSINIA: .
Lantana vibummoides (Forske, Vahl is the correct accredition
of this name
Lippia Radula Je Ge Baker
CAMEROONS :
Clerodendrum singwanum Thomas is the correct orthography of
this name
Lippia rugosa Ae Cheve
FRENCH EQUATORIAL AFRICA:
Lippia rugosa Ae Cheve (Middle Congo)
BELGIAN CONGO:
Clerodendrum discolor (Klotzsch) Vatke
Clerodendrun myricoides (Hochste) Re Bre -
Clerodendrum myricoides vare savanorum (DeWilde) Thomag
UGANDA:
Clerodendrum Wallii Moldenke -- delete tie "*"
Lippia efricana vare villosa Moldenke*
TANGANYIKA TERRITORY:
Lantana vibummoides (Forske) Vehl is the correct accredition
of this name
KENYA: :
Lantana viburnoides (Forske) Vahl is tne correct accredition
of this name
Lippia Radula Je Ge Baker -- delete the "*"
ANGOLA: .
Clerodendrum myricoides vare savanorum (DeWild.) Thomas
SOUTHWEST AFRICA:
Lippia Dinteri Moldenke*
BECHUANALAND PROTECTORATEs .
Lantana viburnoides (Forske) Vahl
Lippia javanica (Burme fe) Sprenge
UNION OF SOUTH AFRICA:

1948 Moldenke, Geographic Distribution 483

Clerodendrum glabrum vare ovale (Klotzsch) He He We Pearson
Transvaal )*
Lippia africana Moldenke (Cape of Good Hope & Transvaal)*
Lippia lupuliformis Moldenke (Natal)* |
Stilbe mucronata Ne Ee Bre is to be deleted
Stilbe verticillata (Ecklon & Zeyher) Moldenke (Cape of Good
Hope)*
Bethe parkioi Vata vare cuspidata (He He We Pearson) Molden-
ke (Cape of Good Hope)*
Stilbe Zeyheri Ganloger is to be deleted
Xeroplana Zeyheri Brige (Cape of Good Hope)
ARABIA:
Lantana viburnoides (Forske) Vahl is tho correct accredition
of this name
LEBANON:
Vitex Agnus-castus L.
INDIA:
Durante repens Le (Bombay)
Gmelina arborea Roxbe (Bombay)
Lippia alba (Mille) Ne Ee Bre (Assam)
Stachytarpheta urticaefolia (Salisbe) Sims (Bombay)
CHINA:
Caryopteris ningpoensis Hems]-*

Caryopteris parvifolia Batalin*

FORMOSA:
Clerodendrum Ohwii Kanehira & Hatusima*
JAPAN:

Callicarpa dichotoma (Loure) Ke Koch (Honshiu)
Callicarpa japonica vare luxurians Rehde (Honshiu)
Callicarpa mollis Siebe & Zucce (Honshiu)
Caryopteris divarioata (Siebe & Zucce) Maxim. -- delete "Mu-
. sashi"
Clerodendrunm trichotomm Thunb. -- delete "Ise", add Hokkaido
Vitex sete Vare camnabifolia (Siebe & Zucce) Hande-Mazz.
Honshiu
Vitex trifolia vare simplicifolia Cham -- delete "Ise"
Vitex trifolia vare Seana fe albiflora (Ye Matsumra)
Moldenke (Honshiu *
RYUKYO ISLANDS:
Premna microphylla Tureze (Okinawa)
PHILIPPINE ISLANDS:
Callicarpa subcandida Elme (Luzon)*
HAWAIIAN ISLANDS:
Clerodendrum fragrens (Vents) Re Bre (Oahu)
Vitex trifolia vare simplicifolia Chame (Maui)
CULTIVATED:
Aen, oioides vare paraguariensis (Briqe) Moldenke (Bra-
zil

CAPPARIS BREVIS SPRENGEL IS A GLYPHAEA
Joseph Ve Monachino

I have examined an apparently authentic flowering specimen
of Capparis brevis Sprengel, deposited in the De Candolle Her
barium a nevae The plant is identical with the widely dis-
tributed tropical African tiliaceous species excellently illus-
trated and = yoyo as Glyphaea, grewioides Hooke fe (Ice Pl.
te 7606 1848 e

The Geneva specimen was annotated in 1852 by Ae De Candolle
who noted that it seemed to be a Clematis, but that, in any e-
vent, it was not a member of the Capparidaceaee In 1853 Benthem

identified it as "Carpotroche?" renge elf in Lim.
Syste Vege 2: 576 refers Ce brevis to Capparis eustachi-
ana Jacges but his original description in 1807, which agrees

very closely with the Geneva specimen, obviously precludes any
such dispositione Ce sustachiana has entire, not trinerved,
leaves, and is glabrous, while C. brevis has subserrate triner-
ved leaves, pubescent petioles, and scabridulous stellate-
pubescent branchlets.

On the Geneva sheet De Candolle suggested that the specimen
robably was sent to Sprengel from the Antilles, a notion in
armmony with the belief that Ce brevis is referable to the West

Indian Ce eustechianae The type, however, was obtained from the
Razoumofisky garden, and Sprengel's species is listed in Fe E.
Le von Fischer's Catalogue du Jardin des Plantes a Gorenki (ed.
25 pe 526 18123 ede 1, pe 85e 1808, fide Ae DCe Prodre 1s 253)-
Ce brevis is placed in the synonymy of Ce eustachiana by the
Tndex Kewensis; otherwise it has been given little attentions

Hooker in his Flora Nigritiana (pe 238 1849) oited the
prior Grewia lateriflora Don as a synonym of G ea grewioi-
des. This synonymy is accepted by Oliver (Fl. a Afr. is
Zo7e 1868), who treats likewise Glyphaea Monteiroi Hooke fe
The latter species was noted to be only doubtfully distinct by
Je De Hooker in the original publication. With my discovery of
the identity of Capparis brevis and ohyphaee rewioides, and
accepting the findings of previous workers, complete syno=-
nymy of the species is as follows:

GLYPHAEA BREVIS (Sprengel) Monachino, combe nove
Capparis brevis Sprenge, Fle Hale Mante Prima 43-6 1807e
Grooie Ta ora Ge Don, Gene Syste 1s 549¢ 1831e
6a grewioides Hooke fe, Ice Ple te 760¢ 18486
G. Monteiroi Hooke f. in Curtis, Bote Mage te 5610. 1866.
Ge lateritlora Hutche & Je Me Dalze, Fl. West Trope Afre 1%

Only one other species of Glyphaea is known, Ge tomentosa
Mast. ex Olivers Ge Boivini Baill. and G. chalybaea Baillie, at-
tributed to the genus Glyphaea in the Index Kewensis, were act-
ually published in Baste ty Eailion (Bulle Soce Linn. Paris 13
550. 1886). prin as kh

DAVID DOUGLAS' NEW SPECIES OF CONIFERS

Elbert L. Little, Jr.

A list of the new species of conifers collected by David
Douglas in Northwestern America and California between the
years 1824 and 1832 has been assembled from the original pub
lished sources. This compilation was made in connection with
a study of Aylmer Bourke Lambert's “A Description of the Genus
Pinus,” which contained several of Douglas' discoveries. Pre-
viously, Suringar (4) had published most of these names in a
list with some other species of Douglas. The recently pub-
lished biography, “Douglas of the Fir," by Harvey (2), which
contains much information hitherto unpublished, doubtless will
arouse interest in his work, A summary of Douglas' discoveries
of conifers, therefore, may be appropriate.

David Douglas (1798-1834) (1, 2, 3), a native of Scotland,
was a botanical explorer for the Horticultural Scciety of Lon-
don (now the Royal Horticulturel Society). His first trip, in
1823, was to northeastern United States to obtain propagating
materiel of cultiveted fruits. The following year, on his
next expedition he went by ship around the Horn to the north-
western coast of North America, then returned to England in
1827 by crossing Canada overlard to Hudson Bay. He left
England for the lest time in 1629 on another ocean voyage to
the Columbia River region. In 1831 and 1832 he collected in
California. Then in the latter year he made a brief trip to
Hawaii before returning to the Columbia River. Ee was killed
in Fawaii in 1834 on his second journey there. Several impor-
tant species of forest and ornamental trees are included among
the many plants introcuced by him to horticulture.

Douglas gave herbarium names to 12 species of conifers and
collected specimens of 5 more which later were named as new by
others. However, of these Dougles published before his death
the descriptions of only 2 species of Pinus: Finus Jlambert-
iana Dougl. (Linn. Soc. London Trans. 15: 500. 1827), the
sugar pine, the largest pine in the world, and F. sabiniana
Dougl. (Linn. Soc. London Trans. 16: 749. 1633).

Several of Douglas’ discoveries were described in his manmr
script, "Some American Pines," written eviaently after he left
England the last time and published eighty years after his
death as an appendix of his journal (1, p. 338-348). After
this journal was prepared for the press, two slightly different
copies of this manuscript in cae handwriting, containing

2

486 PHYTOLOGIA Vol. 2, no. ll

17 species of conifers (northern as well as western), with de-
scriptive and geographic notes, were found. Cne species with-
out author or citation, Pinus monticola, wes described here by
Douglas apparently as new. Six species then unpublished, P.
douglasii,P. menziesii, P. nobilis, F. amabilis, P. ponderosa,
and P. contorta, had brief Latin diagnoses followed by the in-
complete citation “Sabine in Trans. Hort. Soc. Vol." However,
Joseph Sabine, secretary of the Horticultural Society of London
and Douglas’ friend and patron, did not publish these new spe-
cies in the Transactions (1, p. 338). The upheaval in the
Society in 1830, followed by Sabine's resignation and in 1832
also by Douglas’ resignation upon getting the news, and Doug-
las' absence from England together ray account for the failure
of these names to be published (2, p. 149-150, 190-192).

Five names of Douglas were published with descriptions by
David Ton in the third edition of Lambert's monographic work
“A Description of the Genus Pinus" (octavo, 2 v., illus. 1832)
among the extra pages inserted as an appendix between pages
144 and 145 in most copies of volume 2. These rames published
"in order to secure to Mr. Douglas the credit of these inter-
esting discoveries" were: Pinus sabiniana, P. monticola, P.
nobilis, P. grandis, and F. menziesii. Here appeared also P.
douglasii “Sabine as a new name for P. taxifolia Lamb., the
valuable timber tree introduced to horticulture by Douglas and
appropriately given the English common name Douglas-fir from
the specific epithet.

Two nomina nuda of Douglas, Pinus amabilis and F. insignis,
appeared in 1835 ina list of plants raised from seed he sent
to the Horticultural Spciety of London, published in the report
of the new secretary, George Bentham (Hort. Soc, London Trans.,
ser. 2, 1: 404. 1835). Douglas' names were merely mentioned,
because the living plants were too young for description.

In the extracts from Douglas’ journal and letters to his
teacher, W. J. Hooker (3), published by the latter in 1836 as a
sort of biography were Pinus venusta Dougl, (Comp. Bot. Mag. 2:
152. 1836), described from | memory in a letter, and two nomina
nuda, P, amabilis Dougl. (p. 93) and F. ponderosa (p. 111, 141).

Douglas’ authorship of another very important lumber tree,
ponderosa pine, was lost, though both the Latin and common names
still retain Douglas’ descriptive epithet for the heavy wood.
As early as 1830 there was published almost as a nomen nudum,
Pinus ponderosa Dougl. ex Loud. (Hort. Brit. 387. 1830). The
name is cited P. ponderosa Laws. (Agr. Man. 354. 1836), though
lewson's nontechnical English description was based upon young
trees without cones and was far infericr to Douglas’ own tech-

1948 Little, Douglas' Conifers 487

nical description in his manuscript. The mere mention by Lawson
that Douglas introduced the species is not sufficient to credit
him as author of his epithet. Two years later a botanical de-
ecription was published es P. ponderosa Dougl. ex Loud. (Arb.
Frut. Brit. 4: 2292, fig. 2210-2211. 1838).

Loudon published in the same volume descriptions and illus-
trations of two additional species of pines named by Douglas:
P. contorta Dougl. ex Loud. (Arb. Frut. Frit. 4: 2292, fig.
2210-2211. 1838) and P. insignis Dougl. ex Loud. (4: 2265,
fig. 2170-2172. 1838). Here also was Ficea amabilis Dougl.

.ex Loud. (4: 2342, fig. 2247-2248. 1838), based upon Pinus

amabilis LDougl. The next year the name now in use, Abies
amabilis (Dougl.) Forb. (Pinet. Woburn. 125, pl. 44. 1639)
was published, perhaps irregular as a new combination, since
Douglas’ manuscript name, a nomen nudum, was cited but Loudon's
description was not mentioned.

David Don published different names for two of Douglas’ new
coniferous species in an article describing five species of
Pinus collected by Dr. Thomas Coulter in California (Linn. Soc.
London Trans. 17: 439-444. 1836). As Dr. Coulter and Douglas
both were in California in 1831 and 1832, they may have col-
lected together or exchanged specimens. Finus bracteata D.Don
(Linn. Soc. London Trans. 17: 442. 18636) competes with P.
venusta Dougl. (1636) for the bristlecone fir. Though exact
priority has not been determined, Abies venusta (Dougl.) K. Koch
generally is adopted by custom (Little, Amer. Jour. Bot. 31:
592. 1944). Pinus radieta D. Don (Linn. Soc. London Trans.
17: 442. 1836) has priority over P. insignis Dougl. (1838) for
the Monterey pine. Douglas proposed no name for P. coulteri
D. Don (Linn. Soc. London Trans. 17: 440. 1836), regarding
it merely as a variety of his F. sabiniana when he sent speci-
mens and seeds beck to England.

Dougles' conifers and other collections from the Northwest
were cited in 1839 in the "Flora Boreali-Americana” by W. J.
Hooker (23 161-167), who received a set of specimens from his
former student. Here were. an ianaiee two more new species based
upon Douglas’ plants: Pinus lasiocarpa Hook. (Fl. Bor.-Amer.
2: 163. 1839) now Abies 3 lasiocarpa (Hook.) Nutt., and Juni-
perus occidentalis Hook. (p. 166), previously named as J. excelsa
Pursh, not Bieb, Thuja menziesii Dougl. (p. 165) was published
in synonymy under T. gigantea gantea Nutt., now T. plicata Donn. How-
ever, in his journal Douglas used T. plicata. The manuscript
name Pinus distorta Dougl. (p. 161) was cited by Hooker as a
synonym ym of P. inops, though F. contorta Dougl., the name now

“in use, had been published by Loudon the year before. Hooker
placed P. monticola Dougl. as a synonym of P. strobus L.

488 PHYTOLOGTIA Vol. 2, no. ll

Douglas' specimen of the species afterwards segregated as Taxus
brevifolia Nutt. (No. Amer. Sylva 3: 86, pl. 1€8. 1849) was
combined by Eooker with T. baccata L., of the Old World.

The 12 species of conifers to which Dougles gave manuscript
names are summarized here under the names now accepted, with
his names, where different, added in synonymy. JDougles still
is cited as author of 7 of the specific epithets now in use.
Only 3 names lack priority, 1 was given the same name by the
publishing author, and 1 is invalid under present rules as a
later homonym. ;

Abies amabilis (Dougl.) Forbes PACIFIC SILVER FIR
Pinus amabilis Dougl., nomen nudum
Picea amabilis Dougl. ex Loud.

Abies grandis (Dougl.) Lindl. GRAND FIR
Pinus grandis Dougl. ex D. Don in Lamb.

Abies procera Rehd. NOBLE FIR
Pinus nobilis Dougl. ex D. Don in Lamb.
Abies nobilis (Dougl.) Lindl., non A. Dietr.

Abies venusta (Dougl.) K. Koch BRISTLECONE FIR
Pinus venusta Dougl. (Dec. 1, 1836)
Pinus bracteata D. Don (1836)

Picea sitchensis (Eong.) Carr. SITKA SPRUCE
Pinus menziesii Dougl. ex D. Don in Lamb. (1832)
Pinus sitchensis Bong. (Aug. 1832)

Pinus contorta Dougl. ex Loud. SHORE PINE
Pinus distorta Dougl. ex Hook., pro syn.

Pinus lambertiana Dougl. SUGAR PINE .

Pinus monticola Dougl. ex D. Don in Lamb. WESTERN WHITE PINE

Pinus ponderosa Laws. PONDEROSA PINE
Pinus ponderosa Dougl. ex Loud., nomen nudum

Pinus radista D. Don MONTEREY PINE
Pinus insignis Dougl. ex Loud.

Pinus sabiniana Dougl. DIGGER PINE

a. hee

Thuja plicata Donn ex D. Don in Lamb. WESTERN REDCEDAR
Thuja menziesii Dougl., pro syn.

1948 Little, Douglas’ Conifers 489

The 5 species of Douglas’ conifers which he did not name wt
which werenamed by others are:

Abies lasiocarpa (Hook.) Nutt. ALPINE FIR
Pinus lasiocarpa Hook.

Juniperus occidentalis Hook. SIERRA JUNIPER

Pinus coulteri D. Don COULTER PINE
Pinus sabiniana Dougl. var., Dougl.
Pinus macrocarpa Lindl.

Taxus brevifolia Nutt. PACIFIC YEW
Taxus baccata Hook. partim, non L.

Pseudotsuga texifolia (Poir.) Britton DOUGLAS-FIR
Pinus douglasii Sabine ex D. Don in Lamb.

Pseudotsuga douglesii (Sabine) Carr.

Even today many Europeans retain the name Fseudotsuga doug-
lesii for the Louglas-fir with some justification, as the no-
menclature is involved and allows more than one interpretation.

Of the 17 species of conifers listed above,Douglas is cred-
ited with the introduction to horticulture of 11 (1, p. 334;
2, p. 254-26C): Abies amabilis, A. grandis, A. procera, Picea
sitchensis, Pinus coulteri, P. lambertiana, P. monticola, F.

onderosa, P. radiata, P.sabiniana, and Pseudotsuga taxifolia.
His specimens of the 6 remaining species, introduced after-
wards, probably did not contain viable seeds. Of course, he
collected specimens of other conifers which were not new. Ammg
these was the redwood, Sequoia sempervirens (D. Don) Endl.
(3,,.<rs. 150; lowell, John Thomas. Leaflets West. Bot. 2: 96.
1936), which was discovered earlier and introduced later.

It is unfortunate that Douglas did not properly publish
descriptions of all the new conifers he named and introduced
and that publication of these new species was spread among the
works of severel authors. His manuscript names were retained
by the authors who supplied descriptions. The association of
his name with the English name Douglas-fir honors his memory
far more effectively than scientific names could.

Forest Service,
United Stetes Department of Agriculture,
Washington, D. C. ;

490 PHYTOLOGIA Vol. 2, no. 12

LITERATURE CITED

1. Douglas, David. Journal kept by David Douglas during
his travels in North America 1823-1627. 364 p., illus.
London, 1914.

2. Harvey, Althelstan George. Douglas of the fir, a _ Dbio-
graphy of David Douglas, botanist. 290 p., illus. Cambridge,
Mass., 1947.

3. H] ooker_|, W. J. A brief memoir of the life of Mr.
David Douglas, with extracts from his letters. Comp. Bot. Mag.
2: 79-162, illus. 1836.

4. Suringar, J. Valckenier. Flantenverzamelaars. I, David
Douglas de plantenverzamelaar in het verre westen ten tijde der
Indianen- en buffelheerschappij. Nederland. Dendrol. Ver.
Jeaarb. 2 (1926): 69-97, illus. 1926 [1927].

THE KNOWN GEOGRAPHIC DISTRIBUTION OF THE MEMBERS OF THE
ERIOCAULACEAE. SUPPLEMENT 3

Harold N. Moldenke

Since the publication of the previous Supplement to this
list several thousand additional specimens of this group have
been examined and annotated from the herbaria of the University
of Colorado at Boulder, the Facultad Nacional de Agronomia at
Medellin, Colombia, the University of Massachusetts at Amherst,
the Chicago Natural History Museum, the California Academy of
Sciences at San Francisco, the Instituto Miguel Lillo at Tucu-
min, Argentina, Oregon State College at Corvallis, the State
College of Washington at Pullman, Rancho Senta Ana Botanic
Garden at Anaheim, California, the University of Washington at
Seattle, the Instituto Darwinion at San Isidro, Argentina, the
Southern Methodist University at Dallas, Texas, Oklahoma Agri-
cultural and Mechanical College at Stillwater, the United
States National Herbarium at Washington, end the Britton Herbar-
ium at the New York Botanical Garden. These specimens have
brought to light 47 new country or island records, 48 new state,
province, or department records, and 30 new county or parish
records. Also, 151 new binomials or trinomials, or corrections
of previous entries, must be added to the alphabetic list of
scientific names proposed in this group.

1948 Moldenke, Ertoceulacese AOL

ISLE 8T. IGNACE:
Eriocaulon septangulare With.
UNITED STATES OF AMERICA:
New York:
Eriocaulon septangulare With. (Clinton County)
New Jersey:
Eriocaulon septangulere With. (Middlesex County)
Pennsylvania:
Eriocaulon septangulare With. (Carbon & Sullivan Counties)
Maryland:
Eriocaulon Parkeri B. L. Robinson (Caroline, Cecil, Dor-
chester, & Harford Counties)
Virginia:
Eriocaulon Parkeri B. L. Robinson (Fairfax County)
North Carolina:
Eriocaulon decangulare L. (Craven County)
South Carolina: —
Eriocaulon decangulare L. (Clarendon County)
Georgia:
Eriocaulon compressum Lam. (Baldwin County)
Lachnocaulon anceps (Walt.) Morong (Charltom County)
Syngonanthus flavidulus (Michx.) Ruhl. (Ware County)
Florida:
Eriocaulon compressum Lam. (Alachua & Pinellas Counties )
Eriocaulon lineare Small (Putnam County)
Eriocaulon Ravenelii Chapm. (Okeechobee County)
Lachnocaulon anceps (Walt.) Morong (Franklin County)
Lachmocaulon Engleri Ruhl. (Polk County)
Lachnocaulon glabrum Kérn. (Collier, Duval, Manatee, & Okee-
chobee Counties
Lachnocaulon minus (Chapm.) Small (Hillsborough County)
Alabama:
Eriocaulon compressum Lam. (Coffee County)
Lachnocaulon digynum Kérn. (Mobile County)
Mississippi:
Eriocaulon decangulare L. (Hancock County)
Lachmocaulon digynum Kérn. (Harrison County)
Wisconsin:
Eriocaulon septangulare With. (Oconto County)
MEX TCO:
Eriocaulon microcephalum H.BeK. (Tamaulipas)
VENEZUELA: .
Eriocaulon Steyermarkii Moldenke -- delete the "*"
Paepalantius dichotomus Klotzech (Amazonas )
Syngonanthus Steyermarkii Moldenke (Tdchira)*
COLOMBIA:
Paepalenthus andicola var. villosus Moldenke (Cundinamarca )*
BRITISH GUIANA:
Comenthera Linderi L. B. Sm.*

492 PHYTOLOGIA Vol. 2, no. ll

SURINAM:

Eriocaulon Steyermarkii Moldenke
PERU:

Paepalanthus Karstenii Ruhl. (Cuzco)
BRAZIL:

Eriocaulon Glaziovii Ruhl. (Goyaz)

Eriocaulon tenuifolium Klotzsch (Bahia)

Leiothrix ourvifolia var. microphylla Alv. Silv. (Minas Ger-
aes )*

Leiothrix Dieleii Ruhl. (S&o Paulo)

Leiothrix hirsuta var. Magalhd&esii Alv. Silv. -- to be delet-
ed

Leiothrix polystemma var. robusta Alv. Silv. (Parang)

Paepalanthus amoenus var. curralensis Alv. Silv. (Minas Ger-
aes )*

Paepalanthus armeria Mart. (Minas Geraes )

Paepalanthus batocephalus Ruhl. (Minas Geraes)

Paepalanthus Benedicti Alv. Silv. (Minas Geraes) -~ this is
the preferred orthography of this binomial

Paepalanthus caracensis Alv. Silv. (Minas Geraes )*

Paepalanthus cearaensis Ruhl. (Minas Geraes)

Paepalanthus decipiens Ruhl. (Minas Geraes)

Paepalanthus dupatya Mart. (Minas Geraes)

Paepalanthus elongatus var. ciliatus Kérn. (Minas Geraes )*

Paepalanthus glabrifolius Ruhl. (Minas Geraes )

Paepalanthus grfo-mogoliensis Alv. Silv. -- this is the prefer-
red orthography of this binomial.

Paepalanthus Hilarei var. piauhyensis Ruhl. (Minas Geraes)

Paepalanthus itatiaiensis Ruhl. (Minas Geraes)

Paepalanthus jordanensis Alv. Silv. (S&o Paulo)

Paepalanthus Lundii K6rn. (Minas Geraes )

Paepalanthus multicostatus Ruhl. (8&o Paulo)

Paepalanthus myocephalus var. minor Kérn. (Bahia)

Paepalanthus myriophyllus Alv. Silv. (Minas Geraes )

Paepalanthus parvus Ruhl. (Pernambuco)

Paepalanthus planifolius var. alpestris Ktrn. (Minas Geraes)
Paepalanthus rigidulus Mart. (Minas Geraes) |

Paepalanthus Sellowianus Kérn. (Minas Geraes )

Paepalanthus Standleyi Moldenke (Minas Gcraes )*

Paepalanthus stellaris (Guill.) Kunth (Minas Geraes)*

Paepalanthus suffruticans var. angustifolius Alv. Sily. (Min-
as Geraes )*

Paepalanthus Warmingianus (Ktrn.) Kérn. -- this is the correct
orthography of this binomial.

Syngonanthus aquaticus Alv. Silv. (Minas Geraes)

Syngonanthus compactus Ruhl. (Amazonas )

Syngonanthus densus (Kérn.) Ruhl. (Piauhy)

Syngonanthus gracilis (Ktrn.) Ruhl. (Minas Geraes)
Syngonahthus gracilis var. setaceus Ruhl. (Bahia)

1948 Moldenke, Eriocaulaceae 493

Syngonanthus graéo-mogolensis Alv. Silv. -- this is the correct
orthography of this binomial

Syngonanthus habro Ruhl. (Minas Geraes )
Syngonanthus nitens es Ruhl. (Minas Geraes)

Syngonanthus oblongus (Kérn.) Ruhl. (Goyaz)
Syngonanthus oblongus var. aequinoctialis Ruhl. (Bahia)
Syngonanthus Schwackei Ruhl. (oahie)
Syngonanthus umbellatus var. Liebmannianus (Ktrn.) Ruhl.
(Mat togrosso
MARACA ISLAND:
Paepalanthus polytrichoides Kunth
Syngonanthus bulbifer (Huber) Ruhl.
MARAJO ISLAND:
Paespalanthus Lamarckii Kunth
Philodice Hoffmannseggii Mart.
Syngonanthus gracilis var. amazonicus Ruhl.
Syngonanthus Huberi Ruhl.
Syngonanthus umbellatus (Lam.) Ruhl.

NEPAL:

Eriocaulon oryzetorum Mart.
INDIA:

Eriocaulon nepalense Prescott (Bombay)
MANC HUKUO:

Eriocaulon robustius (Maxim.) Mak.
CHINA: :

Eriocaulon formosanum Hayata (Kwangtung )
KOREA!

Eriocaulon robustius (Maxim.) Mak.
FORMOSA:

Eriocaulon cinereum R. Br.
Eriocaulon formosanum Hayata -~ delete the "*"
Eriocaulon petrospermum Hayata -~ to be deleted
Eriocaulon pterospermum Hayata*
JAPAN:
Eriocaulon alpestre Hook. f. & Thoms. (Hokkaido)
Eriocaulon cinereum R. Br. (Honshiu) -- delete "Musashi"
Eriocaulon hondoense Satake (Hokkaido, Honshiu, Kiushiu, &
Yezo)*
Eriocaulon hondoense var. pilosum Satake (Honshiu)*
Eriocaulon hondoense var. stellatum Satake (Honshiu)*
Eriocaulon perplexum Satake & Hara (Hokkaida)*
Eriocaulon piliphorum Satake (Honshiu)*
Eriocaulon robustius (Maxim.) Mak. (Honshiu, Kiushiu, Shiko-
ku, & Yezo
Eriocaulon sikokianum Maxim. (Hokkaido)
Eriocaulon truncatum Hamilt. (Kiushiu)
Eriocaulon Zyotanii Satake (Honshiu)*
AUSTRALIA:
Eriocaulon scariosum J. Sm. -- delete the "*"

4o4 PHEDOiO GT & Vol. 2, no. ll

BRIBIE ISLAND:
Eriocaulon australe R. Br.
Eriocaulon scariosum J. Sm.
DOUBLE ISLAND:
Eriocaulon australe R. Br.
Eriocaulon scariosum J. Sm.

Addenda and errata to the alphabetic list of scientific
names proposed in the Eriocaulaceae, including mis~spell-
ings and mis-accreditions

Kunth
Blastocaulon rupestris (Gard.) Ruhl. = Blastocaulon rupestre
Gard.) Ruhl.
Blastocaulum Ruhl. = Blastocaulon Ruhl. .
Carphocephalus caulescens Kunth = Syngonanthus caulescens
(Poir.) Ruhl.
Eriocaulon decangulare Lightf. = Eriocaulon septangulare With.
Eriocaulon eleocharoides Chapm. = Lachnocaulon Engleri Ruhl.
Eriocaulon floridanum Chapm. = Eriocaulon decangulare L.
Eriocaulon hondoense Satake
Eriocaulon hondoense var. pilosum Satake
Eriocaulon hondoense var. stellatum Satake
Eriocaulon iaponicum Kérn.
Eriocaulon japonicum Kérn. = Eriocaulon iaponicum Ktrn.
Eriocaulon Kunthii var. j Kérn. = Eriocaulon Kumthii Ktrn.
Eriocaulon longifolium var. Wallichianum Burbidge = Eriocaulon
longifolium Nees
Eriocaulon longirostrum Alv. Silv. = Eriocaulon longirostrum
Alv. Silv. & Ruhl.
Eriocaulon Miquelianum Auct. Jap. = Briocaulon hondoense Satake
Eriocaulon Miquelianum Koeck. = Eriocaulon Miquelianum Kérn.
Eriocaulon nipponicum Tatew. = Eriocaulon perplexum Satake &
Hara
Eriocaulon perplexum Satake & Hara
Eriocaulon petrospermum Hayata = Eriocaulon pterospermum Hayata
Eriocaulon pterospermum Hayata
Eriocaulon piliphorum Satake
Eriocaulon pumilum Chapm. = Lachnocaulon Engleri Ruhl.
Eriocaulon Ravenelli Chapm. = Eriocaulon Ravenelii Chapm.
Eriocaulon Ravenellii Chapm. = Eriocaulon Ravenelii Chapm.
Eriocaulon Sellovianum Kunth = Eriocaulon Sellowianum Kunth
Eriocaulon spongiosum Alv. Silv. = Eriocaulon spongiosifolium

Actinocephalus polyanthus Kunth = Paepalanthus polyanthus
(Bong .)

Alv. Silv.
Eriocaulon stellare Guill. = Paepalanthus stellaris (Guill.)
Kunth

Eriocaulon truncatum Buch.~Ham. =» Eriocaulon truncatum Hami lt.

|

q
-

1948 Moldenke, Paepalanthus 495

Eriocaulon Zyotanii Satake
Eupaepalanthus Freyreissii Kérn. = Paepalanthus Freyreissii
' (Thunb.) Kérn.
Eupaepalanthus minutulus Mart. = Paepalantims minutulus Mart.
Eupaepalanthus Oerstedianus Kérn. = Paepalanthus Oerstedianus

Kérn.
Eupaepalanthus plantagineus K6rn. = Paepalanthus plantaginous
Bong « Korn.

Eupaepa lanthus Schenckii V. Ae Pouls. = Paepalanthus Schenckii
Ve Ae Pouls.
Eupaepalanthus Schraderi Ktrn. = Paepalanthus bifidus (Schrad.)

Kunth

Eupaepalanthus tortilis Kirn. = Paepalanthus tortilis (Bong.)
Mart.

Eupaepalanthus Warmingianus Kérn. = Paepalanthus Warmingianus
”(Kérn.) Kérn.

Leiothrix afinis Alv. Silv. = Leiothrix affinis Alv. Silv.
Leiothrizx araxaensis Alv. Silv. = Leiothrix araxa@nsis Alv.
Silv. z
Leiothrix curvifolia var. microphylla Alv. Silv.
Leiothrix flavescens (Kérn.) Ruhl. = Lelothrix flavescens
: (Bong «) Ruhl. :
Leiothrix hirsuta var. Blanchetiana Ruhl. = Leiothrix hirsuta
5 vare Blanchetiana (Ktrn. ) Ruhl.
Leiothrix hirsuta var. Magalhfesii Alv. Silv. = Leiothrix Go-
mesii Alv. Silv.
Leiothrix lanuginosa Bong. = Leiothrix curvifolia var. lanugin-
ose Bong « Ruhl.
Leiothrix Michaeli Alv. Silv. # Leiothrix Michaelii Alv. Silv.
Leiothrix trichophyllus Alv. Silv. = Leiothrix trichopus Alv.

Silv.
Leiothrix vivipara (Mart.) Ruhl. = Leiothrix vivipara (Bong.)
Ruhl.

Leptocephali Ruhl. -- a group of Paepalanthus Mart.
Lophophyllum Itatiaiae Kérn. = Leiothrix Beckii (Szysz.) Ruhl.

Nasmythia angustifola Chapm. = Eriocaulon compressum Lam.

Paepelanthus amoenus Kérn. = Paepalanthus amoenus (Bong.) Ktrn.

Paepalanthus amoenus var. curralensis Alv. Silv.

Paepalanthus andicola var. villosus Moldenke

Paepalanthus Arniesii Ruhl. = Paepalanthus stellaris (Guill.)
Kunth

Paepalanthus Benedicti Alv. Silv.

Paepalanthus bryoides Kunth = Paepalanthus bryoides (Bong. )

Kunth
Paepalanthus canescens (Bong.) Ruhl. = Paepalanthus canescens
(Bong.) Kérn.

Paepalanthus canescens var. angustifolia Ruhl. = Paepalanthus
canescens f. angustifolius Ruhl.

Paepalanthus caparoensis Ruhl. = Paepalanthus caparo§nsis Ruhl.

496 PHYTOLOGIA Vol. 2, no. ll

Paepalanthus caracensis Alv. Silv.
Paepalanthus cearensis Ruhl. = Paepalanthus cearaensis Ruhl.
Paepalanthus chloronema Ruhl. = Paepalanthus chloronema Alv.

Silv.
Paepalanthus ciliatus (Bong.) Ruhl. = Paepalanthus ciliatus
(Bong.) Kunth .

Paepalanthus cillioatus Ruhl. = Paepalanthus ciliolatus Ruhl.
Paepalanthus Clausenianus Kérn. = Paepalanthus Claussenianus

Kérn.
Paepalanthus corymbosus (Bong.) Ruhl. = Paepalantius corymbosus

Paepalanthus diplobator Ruhl.

Paepalanthus diplobector Ruhl. = Paepalanthus diplobator Ruhl.

Paepalanthus diplobetor Ruhl. = Paepalanthus diplobator Ruhl.

Paepalanthus diuaricatus (Bong.) Ruhl. = Paepalanthus divarica-
tus (Bong.) Kunth

Paepalanthus dominguensis Ruhl. = Paepalanthus domingensis Ruhl.
Paepalanthus Edwallii Alv. Silve = Leiothrix Edwallii Alv.Silv.

Paepalanthus elongatus Kérn. = Paepalanthus elongatus (Bong.)
Kérn.

Paepalanthus elongatus var. ciliata Kirn.=Paepalanthus elonga-
tus var. ciliatus Ktrn.
Paepalanthus elongatus var. ciliatus Kérn.

Paepalanthus erectilolius var. grandifolia Alv. Silv. = Paepal- .

anthus erectifolius var. grandifolius Alv. Silv.

Paepalanthus exiguus Kirn. = Paepalanthus exiguus (Bong.) Ktrn.
Paepalanthus falcifolius Ruhl. = Paepalanthus falcifolius Kérn.

Paepalanthus flacidus Kunth = Paepalanthus flaccidus (Bong.)

Kunth
Paepalanthus flavidulis Kunth = Syngonanthus flavidulus (Michx.)
Ruhl,

Paepalanthus Freyreissi Kérn. = Paepalanthus Freyreissii
(Thunb. ) Ktrn.

Paepalanthus grao-mogolensis Alv. Silv. = Paepalanthus grifo-
mogolensis Alv. Silv.
Paepalanthus grfo-mogolensis Alv. Silv.

Paepalanthus Hilairei var. pihauhyensis Ruhl. = Paepalanthus
Hilairei var. piauhyensis Ruhl.

Paepalanthus incanus Kunth = Paepalanthus incanus (Bong.) Ktrn.

Paepalanthus Langsdorfii Kérn. = Paepalanthus Langsdorffii
(Bong.) Kérn.

Paepalanthus Leisseringii Ruhl. = Paepalanthus Leiseringii Ruhl.

Paepalanthus lepidus Alv. Silv.

Paepalanthus lycopodiifolius Alv. Silv. = Paepalanthus lycopod=
Loides Alv. Silv. ;

Paepalanthus macrorhizus Kunth = Paepalanthus macrorrhizus

Paepalanthus microphyllus (Guill.) Ruhl. = Paepalanthus micro-
phyllus (Guill.) Kunth

=
™ 4.

ee ee ee ee

1948 Moldenke, Eriocaulaceae 497

Paepalanthus myocephalus var. major K8rn. = Paepalanthus myo-
cephalus (Mart.) Kérn.

teagan myocephalus var. minor Ktrn.
Paepalanthus nacrothrichus Alv. Silv. = Paepalanthus macrotri-

chus Alv. Silv.
Paepa lanthus negregens Alv. Silv. = Paepalanthus nigrescens

Alv. Silv.

Paepalanthus planifolius Kirn. = Paepalanthus planifolius
(Bong -) Kérn.

Pae aia plantagineus Korn. = Paepslanthus plantagineus

Bong.) Kérn.

Paepalanthus plumosus Ktérn. = Paepalanthus plumosys ( Bong.)
Kérn.

Paepalanthus polyandros Alv. Silv. = Paepalanthus polyandrus
Alv. Silv.

Paepalanthus polyanthus Kunth = Paepalanthus polyanthus (Bong -)
Kunth

Paepalanthus polyanthus var. Kirn. = Paepalanthus polyanthus
(Bong.) Kunth

Paepalanthus preadensatus Alv. Silv. = Paepalanthus praedensat-
us cue aeiiir. Silv.

Paepalanthus pubescens var. chapadensis Alv. Silv. = Paepalan-
thus pubescens var. chapadensis Ruhl.

Paepa lanthus eee vare longepilosa Alv. Silv. = Paepalanthus

pullus var. longepilosue Alv. Silv.
Paepalanthus ramosus Kunth = Paepalenthus ramosus (Wikstr.)

Kunth

Paepalanthus Riedelianus var. macrocephala Alv. Silv. = Paepal-
anthus Riedelianus (Bong.) Kérn.

Paepalanthus saxatilis Kirn. = Paepalanthus saxatilis (Bong.)
Kérn.

Paepalanthus Schlechteri (Ruhl.) Macbr. = Syngonanthus Schlech-
teri Ruhl.

Paepalanthus Schwackeanus var. glabrecens Alv. Silv. = Paepal-
anthus Schwackeanus var. glabrescens Alv. Silv.

Paepalanthus sellovianus Kirn. = Paepalanthus Sellowianus KSrn.

Paepalanthus spyrophorus Alv. Silv. = Paepalanthus spirophorus
Alv. Silv.

Paepalanthus stellaris (Guill.) Kunth

Peepalanthus stellaris Kunth = Paspalanthue stelleris (Guill.)
Kunth api lntAeck te Phe

Paepa lant hus suffruticans var. angustifolia Alv. Silv. = Pae-
palanthus suffruticans var. angustifolius Alv. Silv.

Paepalanthus suffruticans var. angustifolius Alv. Silv.

Paepa lanthus trichopetalus Alv. Silv. = Paepalanthus trichopet~
alus Ktrn.

Paepalanthus t trichophyllus Ktrn. = Paepalanthus trichophyllus

Bong.) Kérn.
Paepalanthus vellosioides Kérn. = Paepalanthus vellozioides

498 P Be¥eB O50 :6°T.A Vol. 2, noe ll

Kérn.
Paepalanthus virides Kérn. = Paepalanthus viridis Kérn.
Paepalanthus Warmingianus Kérn. = Paepalanthus Warmingianus
(Kérn.) Kérn.
Paepalanthus Warmingianus (Kérn.) Kérn.

Paepalanthus Warmingii Kérn. = Paepalanthus Warmingianus (Ktrn.)
Kérn .

Philodice Hoffmansegii Mart. = Philodice Hoffmannsegeii Mart.

Platycaulon consanguineum Kérn. = Paepalanthus planifolius var.
consanguineus (Kérn.) Ruhl.

Psilocephalus nitens Kunth = Syngonanthus nitens (Bong.) Ruhl.

Syngonanthus aciphyllus (Kérn.) Ruhl. = Syngonanthus aciphyllus

Bong - ) Ruhl.

Syngonanthus anthemidiflorus var. £ K6rn. = Syngonanthus anth-

emiflorus (Bong.) Ruhl.

Syngonanthus caracensis var. glablescens Alv. Silv. = Syng onan=
thus caracensis var. glabrescens Alv. Silv.

ec eer ee ee

Syngonanthus caulescens (Kunth) Ruhl. = Syngonanthus caulescens
(Poir.) Ruhl.

Syngonanthus caulescens var. bello-horizontina Alv. Silv. =
Syngonanthus caulescens var. bellohorizontinus Alv. Silv.

Syngonanthus cipoensis Alv. Silv. = Syngonanthus cipotnsis Alv.

Silv.

Syngonanthus goyazensis (Bong.) Ruhl. = Syngonanthus goyazensis .
(Kérn.) Ruhl.

Syngonanthus gracilis (Kunth) Ruhl. = Syngonanthus gracilis

Kérn.) Ruhl.
Syngonanthus gracilis var. & Kunth) Ruhl. = Syngonanthus grac-
ilis (Kérn. Ruhl.

Syngonanthus gracilis var. microphylla Alv. Silv. = Syngonanth-
us gracilis (Kérn.) Ruhl.

Syngonanthus grao-mogolensis Alv. Silv. = Syngonanthus grfo-
mogolensis Alv. Silv.

Syngonanthus grfo-mogolensis Alv. Silv.

Syngonanthus helminthorhizus (Mart.) Ruhl. = Syngonanthus hel-
minthorrhizus (Mart. ) Ruhl.

Syngonanthus lanceolotus Alv. Silv. = Syngonanthus lanceolatus

Alv. Silv.
Syngonanthus nigrescens Alv. Silv. = Syngonanthus niger Alv.
Silv.

Syngonanthus nitens var. Koernickeana Ruhl. = Syngonanthus
nitens var. Koernickei Ruhl.

Syngonanthus niveus (Kunth) Ruhl. = Syngonanthus niveus ( Bong.)
Ruhl.

Syngonanthus oblongus var. aequinoxialis Ruhl. = Syngonanthus
oblongus var. aequinoctialis Ruhl.

Syngonanthus pallens Alv. Silv.

Syngonanthus pallidus Alv. Silv. = Syngonanthus pallens Alv.
Silv.

1948 Moldenke, Eriocaulaceae 499

Syngonanthus Steyermarkii Moldenke
Syngonanthus umbellatus var. Liebmanniana Ruhl. = Syngonanthus

umbellatus var. Liebmannianus (Kérn.) Ruhl.
Trichocalyx rufulus Kunth = Leiothrix rufula (A. St. Hil.) Ruhl.

Addenda

Paepalanthus Hilairei var. paiuhyensis Ruhl. = Paepalanthus
Hilairei var. piauhyensis Ruhl.

8 anthus atro-virens Ruhl. = Syngonanthus atrovirens
(Kérn.) Ruhl.

ADDITIGNAL NOTES ON THE GENUS PETREA. IV

Harold N. Moldenke

PETREA Houst.

References: Irm&o Augusto, Flora do Rio Grande do Sul 227
fas "Petraeas Le"]. 19463 E. H. Walker, Contrib. U. Se Nat.
Herb. 30 (1): 402. 1947.

According to Dr. L. H. Bailey's list of nurserymen handling
various species of cultivated plants, the Everglades and the
Royal Palm nurseries in Florida handle P. volubilis.

The Glaziou 16320 [Macbride photos 24636] distributed as
"Petrea Glaziovii Briq." is solanaceous.

PETREA ANDREI Moldenke
Additional citations: ECUADOR: Loja: R. Espinosa 1171 (N).

PETREA ASPERA Turcz.
Additional citations: VENEZUZLA: Yaracuy: Killip 37068 (3).

PETREA BRAQTEATA Steud.

Additional citations: BRITISH GUIANA: A- C. Smith 2626 (8).
PETREA MACROSTACHYA Benth.

The species is called "sandpaper vine" in British Guiana and
is described as a tough gray "rope" with stems about 1 om. in
diameter, not reaching to the crown of trees, leaves stiffly
leathery and scabrous, flowers in long terminal drooping spikes,
the calyx-lobes enlarged, membranous, purple, and the "flowers"
tubular and deep—purple.

Additional citations: BRITISH GUIANA: Fanshawe 4854 [F.2118]
(N); Forest Dept. British Guiana 4854 [F.2116] (N); A. OC. Smith

shor (3).

— es

500 PHYTOLOGIA Vol. 2, no» ll

PETREA MAYNENSIS Huber

The species has been collected on high shores of streams,
in anthesis in February.

Additional citations: BRAZIL: Amazonas: Ducke s.n. [Herb.
Rio de Janeiro 35657] (8).

PETREA PERUVIANA var. ACUMINATA Moldenke

Seibert describes this plant as a beautiful iiana which
could well be used as an ornamental, the flowers very sweetly
odorous, the corolla lavender. He collected it at an altitude
of 150 m., blooming in November.

Additional citations: PERU: Loreto: Seibert 1880 (W--
1909049); Mexia 6498 (Gg--288808 ).

PETREA PUBESCENS Turcz.

Seibert describes this species as a beautiful tree to 12 m.
tall, worthy of cultivation, with bluish-lavender flowers in
September, growing at an altitude of 330 m.

Additional citations: PERU: Madre de Dfos: Seibert 2164 (W--
1909065).

PETREA RACEMOSA Nees ;
The species is described as a “weeping” shrub, growing at an
altitude of 1100 m., blooming in September and October.
Additional citations: BRAZIL: Minas Geraes: Williams & Assis
7602 (G, N). So Paulo: Heiner 206 (8).

PETREA RUGOSA H.B.K.
Additional citations: COLOMBIA: Caldas: Haught 2103 (S).

PETREA VOLUBILIS L.

References: Seymour, Host. Ind. Fumgi N. Am. 588-589. 1929;
E. A. Menninger, Introductory Offer Flow. Trees Coll. [2].

1946; E. A. Menninger, 1947 Cat. Flow. Trees 43. 1946; Irmfo
Augusto, Flora do Rio Grande do Sul 227 [as "Petraeae volubilis
Le" J. 1946.

Menninger calls this plant "Queen's wreath" and "purplewreath
petrea" and offers plants for sale at $2 each. He describes it
thus: "This woody vine to 35 feet from northern South America,
with its rough leaves and lovely blue flowers three times a
year, is one of our finest climbers [in Florida]. The profuse
flower sprays are two-colored for the corolla is dark blue, the
sepal is light blue. When the corolla drops, the sepal persists,
changing to a dull gray and finally serving as a wing for the
single seed embedded in its axis." Irm&o Augusto records the
common name "touca de viuva", Eastwood records “sandpaper vine"
ang Mexia records "veracruza". Matuda gtates that the species
grows also in Chiapas, Mexico. The Kelly 248 and Eastwood s.n.
from Honolulu do not state on their labels that the plants were.

1948 Moldenke, Notes on Petrea . 501

in cultivation, but I assume that they were. Dr. L. H. Bailey,
in his list of nurseries handling material of various cultivat-
ed genera, says that the Everglades and the Royal Palm nurser-
ies in Florida handle this species.

Additional citations: MEXICO: Guerrero: Née 33 (Q). Oaxaca:
Schultes & Reko 569 (0a-~8289). Sinaloa: Mexia 1933a (Gg--
157102). Tamaulipas: Edw. Palmer 317 (Gg--31222). Veracruz:
Matuda 1478 (Mh). Yucatdn: G. F. Gaumer 379 (Gg--160526).
BRITISH HONDURAS: Gentle 2366 (Mh). COSTA RIOA: Guanacaste:
Brenes s.n. [Frontera Norte, 1910] (N). CULTIVATED: Florida:
Simmonds son. [U. S. Pe I. 36024] (Ca--9144). Hawaiian Islands:
Eastwood s.n. [Honolulu, August 1--16, 1924] (Gg--34502); Kelly
248 (Gg--51221). Mexico: C. Conzatti 5305 (Mh). Peru: Souku
tae Me LOCALITY OF COLLECTION UNDESIGNATED: Née 34 (Q), 30
Q);

3 (Q)-

PETREA VOLUBILIS var. PUBESCENS Moldenke

Additional citations: MEXICO: Oaxaca: Seler & Seler 2777
(Gg-~245875). Tamaulipas: F. A. Barkley 17M174 (N). COSTA RICA:
Alajuela: Brenes eae [133 279; is19LI N, Si), 14290 [5;
144110] (N), s.n. [San Gerardo de San Ramon, 1903] (N, Si, Si).

PETREA Houst.

The Baker and Baker s.n. from Cambodia, distributed as
"Petraea” is actually something in the Nyctaginaceae. It is
represented by sheet no. 31220 in the California Academy of
Sciences herbarium. The Thorp 29695, sheet 299022 in the same
herbarium, distributed as Petraea zanquebarica J. Gay, is actu-
ally Dicerocaryum zanguebarium ee Merr. in the Pedaliaceae.

PETREA KOHAUTIANA Presl
Additional citations: MARTINIQUE: Sieber Fl. Mart. 374 [Herb.
Prager 18663] (Gg--31219).

PETREA MACROSTACHYA Benth.

References: A. R. McIntyre, Curare 31. 1947 [quoting Robert
Schomburgk].

Additional citations: BRITISH GUIANA: Herb. Forest Dept. Br.
Guiana 4854 [F.2118] (K).

PETREA MAYNENSIS Huber

Sandeman describes this plant as "a small tree growing in
semi-shade, with very beautiful flowers, making the effect of
Parma violet petals surrounded by pointed periwinkle blue
bracts. Ovate scabrous opposite leaves. The inflorescence has
rather the appearance of a Triplex [=Triplaris]". He found it
blooming in November at an altitude of 700 feet.

Additional citations: PERU: Loreto: Sandeman 3474 (K).

502 | P BY ToOsk OG. Ia Vol. 2, no. ll

PETREA PUBESCENS Turez.

Hanbury-Tracy describes this species as a "tree 20 to 40
feet tall; flowers dull purple", collected at an altitude of
5000 feet.
grapes citations: VENEZUELA: Mérida: Hanbury-Tracy 136
K, K ° 3

ADDITIONAL NOTES ON THE GENUS AMASQCNIA. III

Harold N. Moldenke

AMASQNIA ANGUSTIFOLIA Mart. & Schau.

Additional citations: BRAZIL: Goyaz: G. Gardner 3411 [Herb.
Monac. 924; Macbride photos 20345 & 28390] (N--photo of type,
N--photo of isotype).

AMASONIA CAMPESTRIS (Aubl.) Moldenke

References: Pynaert & Gentil, Rev. Hort. Belg. 22: 211 [as
A. punicea]. 1896.

Additional citations: TRINIDAD: Ryan s.n. [Macbride photos
22773] (N-=photo). BRITISH GUIANA: A. C. Smith 2441 (s). FRENCH
GUIANA: Aublet s.n. [Herb. Montintt] (F--photo of isotype, N--
photo of isotype, S--isotype, Si--photo of isotype, Z--photo of
isotype). BRAZIL: Bahia: Blanchet 3156 [Macbride photos 7887 &
30184] (N--photo, N--photo). Maranh&o: Frées 11779 (8).

AMASCNIA HIRTA Benth.

Additional citations: BRAZIL: Mattogrosso: Martius 5835
(Herb. De Candolle 827; Herb. Monac. 929; Macbride photos 7886
& 20346] (N--photo, N--photo). So Paulo: L. Riedel 814 (N).

AMASONTA SPRUCEANA Moldenke —

Additional citations: VENEZUELA: Amazonas: Spruce 3288
[Macbride photos 28391] (N-~photo of isotype).

= F.

SPECIFIC NAMES IN GRATIOLA
He Ae Gleason

Of the six species of Gratiola which enter the Manual Range,
only three are at all common or well distributed; a fourth oc-
curs only from Delaware southward and again in Ohio and Ken-
tucky, and a fifth is known from a single collection only. A
sixth species is attributed to the genus by Gray's Manual and
various recent authors, but has been assigned to a segregate
gemus by Pennell and appears under two other generic nemes in
current Manualse Of the six, only one, and that one the rarest
in our area, has been able to continue with its traditional
meme unchanged through the recent epidemic of name-changing,
which impresses me as a lamentable condition completely
foreign to the explicitly stated basis of the International
Code: "The éssential points in nomenclature ares (1) to aim at
fixity of names; (2) to avoid or to reject the use of foms
and names which may cause error or ambiguity or throw science
into oe oaeeret

Gratiola pilosa Michx- appears under that name in Gray,
seventh edition, as Sophronanthe pilosa (Michx.) Small in
Britton & Brown, as Tragiola ilosa (Michxe) Small & Pemell
in Small‘s Mamial and in Pemell's recent monographe There is
no competition with the specific epithet pilosae Generic seg-
regation or aggregation is a matter of studied scientific
opinion (we hope) and the International Code neither encour
ages nor discourages ite

Gratiola remosa Walter is unchallengede ~

abundant plant of the Manual Range was long known as

Gratiola sphaerocarpa Elle Since 1918 it has been generally

wn to American botanists that the Linnaean name & virgini-
ema belongs to this plant; this name has been used in the re-
cent manuals of Small and Rydberg and in various local floras,
such as those of Kentucky, Indiana, and Illinoise There is no
doubt that the use of one name for two plants "tends to throw
science into confusion": each of these three local floras
finds it necessary to quote synonyms to make their meaning in-
telligible, and Fernald in a discussion of Gratiola-problems
also had to use both names to insure that his meaning would be
understoode Nevertheless, rules are rules, and there seems to
be no way to avoid this regrettable changee

Gratiola viscosa Schwe of Gray's Mammal and the Illustrated
Flora is a homonym of Ge viscosa Horneme Again nothing can be
done about it and the plant has been re-named G. viscidula
Pennelle

Now we come to the really important casese The most abund-
ant and widely distributed species of the gemus was long known
as Ge virginiana Le When that name was transferred to another
species, as recounted above, this common plant was left with-
out a neme in usagee Pennell resurrected G- neglecta Torre

505

504 P HY 0 L0.6 1°54 Vol. 2, no» ll

(1819) and this name has since come into general usage in most —
recent literaturee Gratiola aurea, usually accredited to Muhl- —
enberg but actually published by Pursh in 1814, is the most
conspicuous local member of the genus and abundant along the
Atlantic seaboard. Both of these names are antedated by Ge
lutea Rafe (1811)- Fernald says Ge lutea applies to G. “virgin-
Tana”; Pennell says it applies to Ge aureae Both have examine
the type; each insists that his identification of it is cor-
rect, but Pemnell bolsters his position by stating that Refin-
esque himself, in a later publication, announced that Pursh's
Ge aurea had already been named Ge lutea. }
““~ Wo matter which man is correct, the results are unfortunate. —
If Pennell is right, the well known Ge aurea gets a new name
If Fernald is right, the equally well known "Ge virginiana", —
just getting accustomed to one new name, must start out anew 3
under a third namee

There is only one thing that can be done in this dilemma,
and that is to exclude Ge lutea from all consideratione That
can be done under the rulese The use of Ge lutea for two dif-
ferent species by two competent botanists, each of whom insists
on the validity of his opinion and will presumably continue to
use the name indefinitely into the future, will certainly be a
"nermanent source of confusion or error" (International Code,
Arte 62). "A name of a taxonomic group must be rejected when
its application is uncertain" (Article 63), which is surely the
case when competent men disagree on the identification of the

Ce

Our species of Gratiola will then be (1) G aurea Pursh,
1814, not Ge lutea Raf. 1811, nomen dubiu; Ge ramosa
Walte; (3) Ge Visoidula Pennell, 1919, not Ge viscida Schw.
1824, homonym; (4) Ge neglecta Torre 1819, not Ge Virginiana of

recent literature or G lutea Ref. 1811, neers oe -
Ge

Virginiana Le 1753, not Ge sphaerocarpa Ell. 1816; (6 “a
pitosa 6.

Author Index to Volume Two

Brenckle, J. F., 169, 402

Cornman, J. Fe, 401

Gleason, H. Ae, 201, 279, 281, 291, 294, 428, 503

Little, E. L., Jr., 451, 457, 485

Lundell, O. L., l

McNair, Je Bey 353

Merrill, E. D.;, 5

Moldenke, H. N., 6, 50, 57, 65, 89, 123, 129, 152, 171, 195,
198, 213, 242, 246, 248, 3506, 325, 349, 363, 372, 582, 387,
408, 433, 464, 477, 490, 499, 502

Monachino, Je Ve, 212, 406, 432, 484

Van Melle, P. J., 185, 249, 353

1948

Index

505

Subject Index to Volume Two

Abelia, 252

Abies, 252, 451, 487, 488, 489

Acanthaceas, 37

Acantholippia, 85; 865905. 556;
538

Acanthopanax, 252

Acer, 206, 207, 208, 209, 252,
45 57; 460, 461, 462, 463

Aconitum, 35, 36

Actaea, 211

Aegiphala, 588

Aegiphila, 7, 8, 9, 58, 59, ee.
ai. 62, 63, 64, 65, 68, 69,
70; 72, 79; 81, 83, Bs, 85,
88, 90, 91, 129, 214, 306,
3335, 334, 335, 384, 387, 388,
589, 3590, 591, 393, 394,
395, 396, 399, 400,
433, 4 437, 438,
459, 440, Aho, 445, 444,
445, 450,
479

-Aegiphyla, 388, 440

Aegiphylla, 397

Aegyphylla, 57
Aesculus, 252, 253

Aethionema, 255

Agaricini, 417

Alangium, 253

alcohols, 39

alkaloids, 35, 46, 47, 48,

Alnus, 253

Alonsoa, 307

Aloysia, 9, 10, 11, 12, 50, 66,

9, 70, 74, 755 80, 81,

82:,.835,7 84, 86, 875. 88, 89;
91, 124, 153, 154, 507, 308,
5309, 510, 327, 530, 355, 3536,
551s 3305 346, 363, 383, 3585,
386, 408, 464, 478, 479, 480,
481, 483

Amaryllis, 42

Amasonia, 74, 91, 198, 199,
200, 246, 247, 334, 336, 502

Amazonia, 247

Amblyarrhena, 8) oe be 304

Ampelopsis, 204, 205, 253

49

Amygdalus, 267, 268
Anacardiaceae, 40

Anastraphia, 214

Andromeda, 253, 255, 266, 278
Anonaceae, 40

Apocynacese, 40

Aralia, 253

Araliaceae, 40

Arbutus, 25%

Arctostaphylos, 253
Arctous, 253

Arcyphyllum, 212
Ardisia,

Aristolochia, 253
Aronia, 2

Artemisia, 254
Asclepiadaceae, 37, 40
Asparegus, 25
Astragalus, 254, 324

Atreaphaxis, 254

Atriplex, 254
Avicermia, 65, 66, 67, 68, 69,

Tay 139 Ths 73s. Ths T99-8Rs
81, 8&3, 85, 86, 88, 91, 92;
93, 124, 153, 345, 479

Avicenniaceae, 65, 89, 123,
124, 125 525, 562, + 908; 477

Avicularia, 169, 170, 171, 402

Azalea, 263, 270

Baillonia, 50

Bambusa, 268

Basistemon, 244

Berberis, 254, 265

Betula, 254

Bignonia, 205

Bignoniaceae, 40

Bixacese, Lo

Blakea, 279, 280

Bombacaceae, 40

Bouchea, 93, 154, 530; 346, 387,
08, 480

Brevaisia, 247

Broussonetia, 254 -

Bruckenthalia, 254

Brunfelsia, 390
Bucida, 456
Buddleia, 254, 363

Bunias, 210 Chaenomeles, 256
Burroughsia, 93, 330 Chamaecerasus, 263, 264
Burseraceae, 40 Chamaecyparis, 256
Buxus, 255 Chascanum, 95, 339, 386
Cakile, 210 Chimonanthus, 256
Calendula, 50 Chiogenes, 256
Calliandra, 1 . Chloanthes, 310, 345, 387
Callicarpa, 66, 67, 68, 69, 70, Chrysanthemum, 151

72, 73, 74, 78, 82, 83, 85, Cinnamomum, 245

88, 93, 94, 95, 124, 154, Cissampelos, 215

155, 255, 310, 326, 327, 334, Cissus, 253
342, 345, 344, 345, 346, 387, Clatus, 216, 256, 261

3

3
q
4
;

479, Citharexylum, 13, 14, 15, 51,
Calligonum, 255 67, O85 69% 10>. 7ls aeeeaae
Calpidisca, 215 74, 76, 77, 78, 79, 80, 81;
Calycanthus, 255 82, 84, 85, 87, 95; 96,.9%5
Calycotome, 255 124," 125, 155, 3275: 330p aaa
Calyptranthes, 215 334, 346, 363, 383, 384, 385,
Calyptrella, 300, 301, 428, 429 387, 433,479
Camphorosma, 255 Citrus, 267
Campsis, 255 Cladrastis, 256
Capparidaceae, 58 Clematis, 241, 256, 257
Capparis, Aah Cleome, 58
Caprifolium, 264 Clerodendron, 257
Carapana, 255 Clerodendrum, 65, 66, 67, 68,
Caricaceae, 40 69, 70, Tl, 725° 5an0 eee
Carpinus, 255 765°77; -183’79;° 00,00 eee
Carpobolei, 418 83, 84, 85; 86, 87, 88, 89,
Carptotepala, 351, 377 ‘ 97, 98, 99, 100, 101, 102,
Caryocaraceae, 40 155 5-326, 332, 3554; Do0s1 ee
Caryo teris, Ls; 68, 70; 7o; 345, 346, 348, 384, 386, 387,

79, 64, 87, 95, 343, 346, 408, 409, 479, 482, 483

387, 483 Clethra, 257 |
Casearieae, 432 climate, 34
Casselia, 243, 244, 245, 246 Cochlospermaceae, 40
Casselieae, 173 Coffea, 248
Cassiope, 255 Colutea, 257
Castelia, 86, 95 Combretaceae, 40
Catalpa, 255 Condalia, 456
Cattleya, 4d <onEre 102, 311, 312, 343, 346,
Ceanothus, 255 3
Cedrus, 456 Conostegia, 429
Celastrus, 255 Contortae, 37
Celtis, 255, 256 Cordia, 130, 442
Centradenia, 428 Coriandrum, 37
Cerasus, 267, 268 Cornus, 257
Zoratostiene, 256 Cornutia, 80, 85, 102, 131,
Cercis, ros Dal; 334, 346, 585, 479

Cestrum, 63 Coronilla, 257

1948

Corylopsis, 257
Cotinus, 452
Cotoneaster, 257
Crataerus, 257, 258
Cremanium, 305, 429
Cruciferae, 36

Cryptomeria, 258
Cryptomyces, 243
Cryptophyta, 417
Cryptostegia, 131
Cupressus, 42, 258
Cyathoidei, 418
Cylindrocephala, 407
Cyperaceae, 376
Cypripedium, 42
Cytisus, 258, 263
Daphne, 258

Daphnopsis, 3, 212, 213
Datura, 34

Decumaria, 258
Dermatocalyx, 1351
Desfontainia, 216, 217, 218
Desmodium, 211, 263
Deutzia, 258, 259
Dicerocaryum, 1735
Dichromena, 378
Dimidiati, 418
Diospyros, 259
Diostea, 102, 337
Dipterocalyx, 53
Dipterocarpeceae, 40
Distylum, 259

Drosophile, 34
Dupatya, 377

Dupotya, 377

Duranta, 165%175:118, 67.68;
FO3 ei ly Tey 135° 745 78579;
80, 825; 84, 85, 86, 87, 88,
£02, 2055 1255: 352525934 354,
336, 337, 345, 346, 364, 385,
409, 464, 479, 480, 483

Ebenaceae, 40

Ehretia, 259

Elaeagnus, 259

Elsholtzia, 259

energy, 33, 34, 39, 48, 49

Enkianthus, 259

Ephedra, qo, 259

Epiphyti, 417

Index

S07

Eriaucolon, 377

Eriocaulaceae, 6, 123, 152, 349,
Dies: DID» Aloe lak Dl panos

Eriocaulon, 124, 132, 133, 134,
142, 155; 218; 21Gss2205 221,
222, 549," 3505 FOL 5. Jaeat DOr,
5055. 373, 3745 3150/3765 Sita
378, 379, 380, 381» 491--495

Erythroxylaceae, 40

Eucalyptus, 37

Eugenia,

Eumeriania, 294

Euonymus, 259, 260

Eupatorium, 406, 407

Euphorbia, 202, 474

Euscaphis, 260

evolution, 34, 39

Fagus, 260

Faradaya, 80, 87, 88, 103, 345

Feramea, 388

Flacourtiaceae, 40, 432

Forsythia, 260

Fraxinus, 260

Fungi, 417

Galanthus, 42

Galium, 210

Galpinsia, 134
Gastromycetes, 417, 418

’ Gaultheria, 260

Genista, 258, 260

Geunsia, fl; 135 T7> 78, 195
81, 83, 84, 103

Ghinia, 105, 173, 243, 384

Glandularia, 146, 148, 149,
150, 238

Gleditsia, 260

glycerides, 36, 46, 47, 48, 49

Glycosmis, 463

Glyphaea, 484

Gmelina, 67, 68, 69, 71, 74, 75,
(6s: 10g BOR G2-o55 84, &5,
86, 88, 103, 104, 340, 342,
544, 345, 346, 485

Graphardisia, 4

Gratiola, 503, 504

Grewia, 4&4

Guttiferae, 40

Gypsophila, 409
Halerpestes, 134

508

Halesia, 456
Halimium, 260
Hedera, 260, 261
Hedysarum, 211
Helianthemum, 261
Heltetta, 223

Heliocarpus, ei 3
Helipterum, 312

Hernandiaceae, 40

Heterocentron, 428

Hibiscus, 261

Holmskioldia, 70, 76, 78; 79;
81, 83, 104, 341, 342, 344,
346, 386

Holodiscus; 261

Hosea, 71, 104

Hopea, 275

Hudsonia, 261

Humiriaceae, 40

Hydrangea, 261

Hyperbaena, 134, 224

Hypericon, 289

Hypericum, 261, 288, 289

Hyphydra, 379

Iberis, 261

Tlex, 261, 262

Imbricata, 407

Tpomoea, 224

Tris, 211

Jasminum, 262

Juglans, 451, 457, 459, 463

Junellia, 51, 72, 79, 86; 104,
199 ¢ 156; 1575-35595 557s B98
385, 409, 465, 466, 481

Juniperus, 185, 186, 187, 188,
189, 190, 191, 192, 193, 194,
195): 262,265 5-593 5°354 5.355,
556, 357, 358, 559, 360, 361,
362, 363, 401, 402, 487, 489

Kadsura, 263

Kerria, 263

Kochia, 263

Labiatae, 2435, 246, 248, 388

Laburnum, 263

Lachnanthes, 379

Lachmnocaulon, 153, 350, 351,
373, 379, 491, 494

Lachnostechys, 104

Lampaya, 51, 52, 77; 104, 537 »

PHYT OL:GG rt bh

Vol. 25 no. li %

538
Landolphia, 291
Lantana; 18, 19, 52;0Sd5e@a.
3 1s 68, 69; 70, 71; T2,
735 T4s 75s 755 Tia 185.4%
80, 82; 83, 84, 855 86, 87;
88, 89, 104, 105, 106, 125, 5
137, 138, 155, 156, 225; 226500
244, 312, 313, 526, 527, ©
529, 330, 5515-3925 9908
535, 536, 557, 5385 5995
341, 342, 2? 5, 3
365, 382, 383, 384 , 385,
410, 411, 412, 413, 467,
478, 479, 480, 481, 482,

483
Larix, 263
Lathyrus, 209, 210, 211, 212
Lauraceae, 40

Lecythidaceae, 40

Lecythis, 497 |
Leiothrix, 351, 374 »379,492-499

Lespedeza, 263

Ligustrum, 263

Limonia, 463

Lindera, 263

Lippia, 10, 19, 20, 2l;.22je00,
33; 54, 65; 67; 69, 79, 71;
129 135 PA TOP Es
SL, O25 'O5; 84, 8S:
106, -107 5-125, 156;
244, 310, 313, 314,
517s 518, 350, 551s
336, 337, 338, 339, 340, 34
347, 365, 383, 384,
387, 408, 414, 415,
470, 479, 480, 481,

Liriodendron, 263

Lisianthus, 235

Loganiaceae, 40

Loiseleuria, 263

Lonicera, 263, 264

Loranthaceas, 40

Lychnis, 319

Lycium, 264

Lycoperdei, 418
Magnolia, 264

Mahoberberis, 264
Mahonia, 265

1948

Malus, 265
Malva, 138
Malvales, 37
Manmnilaria, 255
Manabea, 388, 397, 399
Mapourea, 388
Maripa, 159
Marlea, 263 ;
Megapterium, 139
Melasanthus, 22
Meliaceae, 40
Menispermaceae, 40
Menispermum, 265
Mentha, 319
Meriania, 294, 295, 296, 297;
298, 299, 300
Mespilus, 258
Miconia, 501,502 5 503; 304,
~ 305, 306, 428, 409, 430, 431
Miconieae, 431
Moneses, 42
Monimiaceae, 40
Monochilus, 387
Moraceae, 40
Moringaceae, 40
Morus, 265
Mozartia, 226
Myrica, 265
Myristicaceae, 36, 40
Myrtaceae, 37, 40, 41

Myrtiflorae, BM
Nashia, 53,,-.54 ’ 107; 384

Nelumbo, 205

Nemasporei, 418

Neosparton, 70, 107
Nicotiana, 319

Nigella, 319

Nitraria, 265

Nolana, 320

nomen extinctum, 454, 455

nomina conservanda, 454, 4

nomina extincta, 456

nomina generica conservanda,453

aenins rejicienda, 456
haea, 205

ees, 409

Oenothera, 34, 134, 140

Ochnaceae, 40

Olacaceae, 40

Index

509

Oleaceae, 40

Gneidium, 42

Orixa, 265

Osmanthus, 265

Osmia, +407

Oxalis, 324

Oxera, 345, 566, 367 5 386, 387

Pachydesmia, 4
Pachymeriae, 296
Pachysandra, 265

Padus, 228

Paederia, 265

Paeonia, 265

Paepa lanthus, 140, 228, 229,
20, Bly 2Ias.e52% 351, 552 5
364, 365, 367, 373, 374, 375,
Nls FSs, 3795. 580,, 584, 416,
419, 472, 473, 491--499

Palmae, 36, 40

Paris, 4a

Parthenocissus, 204, 205, 265,

266

Pavia, 252

Pedaliacese, 175

Pehoia, 172

Penstemon, 266

Peraea, 172

Perebea, 432

Peronema, 107

Persicaria, 405, 406

Petitia, 69, 70, 107, 108

Petraca, 172, 176

Petraea, 172,.173, 174, 175,
176, 177, 178, 179, 180, 181,
182, 183, 184, 196, 197, 198

Petraseae, 173

Petraeovitex, 108, 347

Petrea, 74, 11 4::829' 635 G65) GD,
105; 171, 1725. 3755: Agta
176, 177, 178, 179,
162, 183,164.19,
198, 248, 331, 334, 336, 385

Petria, 172

Petroca, 172

Phalloidei, 417

Philadelphus, 266

Phillyria, 266

Philodice, 352, 493, 498

Photinia, 266

510 PHY T OLOG24

Ph le, a2; 69, 72; te
8 3 5, 86, 88, 89,
125, 126, 140, 141,

74;
108,

157,

159, 2335 326, 5275 528,
330, 331, 332, 333, 3345
337, 338, 339, 3540, 341,
343, 344, 347, 383, 354,

478, 479, 481, 482

Phyllodoce, 266
Phys ocarpus, 266

Phytolaccaceae, 40
Picea, 206, 488

Pinaceae, 41

Pinus, 266, 257, 457, 458,

G86, 487, 488, 489
Pisum, 209, 210, 2ll
Pitrea, 172
Platanus, 267
Podocarpus, 267
Polygala, 368
Polygonaceae, 180

81,

109,
158,
3295
3355
342,
385,

485,

Polygonum, 169, 170, 254, 267,
TB 03, 404, 405

Poncirus, 267
Populus, 267
Portulaca, 368
Poterium, 267

Prema, 5, 6, 54, 65, 66, 67,
68, 10, 71; 73, 74, 75; 76;
79, 80, 82, 85, 84, 85, 86,
87, 88, 109, 110, 11, 343,

345, 347, 386, 483

Premos, 5, 6

Priva, 73, 82, 87, 111, 126,
142, 159, 331, 332, 335, 384

Proteaceae, +0

Prunus, 210, 211, 267, 268

Pseudocarpidium, 111
Pseudosasa, 2
Peeudotsuga, 489

Psychotria, 388, 440
Pteridium, 233

Pterostyrax, 268

Ptychocarpus, 432, 433
Sisaraee

Pygmaeopremna, 54, 68, 74, 75s

a0) TLE
Pyrus, 265, 268
Quercus, 202, 269, 270

Ranunculus, 134

eee index, 38, 46, 47,

Rehdera, lll, 333
Rhacoma, 456

Rhamus, 270, 273, 287, 288

Rhaphithamus, 111
Rhizophoraceae, 40
Rhododendron, 270
Rhodothamus, 270
Rhodotypos, 270
Rhondonanthus, 381
Rhus, 142, 270, 271
Rhynchosia, 212
Ribes, 271

Robinia, 255, 271

Rondonanthus, 352, 381

Rosa, 2/7l, 272, 320
Rubiaceae, 40, 248
Rubus, 2/2, 273
Rudbeckia, 320
Ruscus, 2/3

Sabal, 125, 126, 127, 128, 165,
166

Sabina, 473
Sagaretia, 2735
Salicornia, 273
Salix, 2/75, 274
Salvadoraceae, 40
Sambucus, 274
Santolina, 274
Sanvitalia, 320

Sapindaceae, 40
Sapindus, 274
Sapium, 274
Sapotacese, 4o, 432
Sarcomycetes, 417
Sarracenia, 291
Sassafras, 263
Schisandra, 275
Sclerotiaceae, 417
Scrophulariaceae, 244
Sickingia, 497
Sida, 281, 282, 283
Simaruba, 397
Simarubaceae, 40

Sium, 283, 284, 285, 286,

Sequoia, 489
Smilax, 275

Vol. 2, NOsumm

1948

Solanaceae, 63

Solanum, 275

Sophora, 275

Sophronanthe, 503

Sorbus, 265, 275

ei gravities, 37, 46, 47,
8

Sphaeriacei, 417
Sphenodesme, 65, 66, 69, 71,
Pee7?, Ol, OF, 111, 112, 344
Spiraea, 261, 266, 275
Sporigastrei, 418
Stachytarpheta, 22, 54, 55, 65,
oy on,. 09, 70,71, 75, 75;
76; ae 78, 9s 80, 81, 82,
ee 65, 186,67, 88, 112, 113,
234, 320, 332, 535;
338, 340, 341, 34 347,
570, 571, 386 ;
417, 473, 482,
Stachyurus,
Stanhopea, 42
Staphyleacese, 40
Stauntonia, 275
Stephenandra, 275
Stewartia, 275

Stilbaceae, 325, 382, 417, 418

~~

77
Stilbacei, 417; 418
Sterculiaceae, 37, 40
Stilbe, » 113, 474, 483
Stillingia, 274, 275
Struthanthus, 1
Styrax, 275
Svenhedinia, 142
Svensonia, 113
Symbolanthus, 235
Symphorema, 78, 79, 81, 113

Symphoremacese, 142, 325, 382,
(7

i} horicarpos, 2/5 -
Symp locaceae, ho
Symplocos, 275
Syngonanthus, 6, 142, 350, 351,

Index

id

Tapirira, 497
Tatea, 54

Taxus, 488, 489

Tectona, 59, 78, 83, 113, 114,
521, 342, 344, 345, 347

Tei jsmanniodendron, 65, 66, 71,
T5x fos (93 155.79; Oly ee,
84, 85, 114

Tetraclea, 126, 159, 328, 329

Teucridium, 346

Teucrium, 276

Thuja, 276, 487, 488

Thujopsis, 276

Thymus , 276

Tiliaceae, 40

Timotocia, 173, 242, 243, 244,
245, 2

Tithymalopsis, 321, 474

Tonina, 379

Topobea, 279, 280, 281

Toxicodendron, 142

Trachelospermum, 276

Trachyspermum, 37

Tradescantia, 41

Tragiola, 503

Tremelloidei, 417

Triadenum, 288, 289, 290, 291

Trillium, 42

Triplaris, 180, 5Ol

Tripterygium, 276

Trochodendron, 277

tropical acids, 39

Tsoongia, 343, 344

Tuberacei, 418

Tubiflorae, 37

Ulmus, 277, 278, 451, 452

Umbellatae, 294, 298

Umbelliferae, 36

Urginea, 143

Utricularia, 215

~ Vaccinium, 4, 275, 277

Vehea, 292, 293
Varronia, 143, 144, 145

Joes 211s 319 374, 375, 377, Velloziaceae, 365

378, 380, 581, 418, 491--499
Syringa, 260, 275

Taligalea, 246
Tamarix, 275

Verbena, 22, 23, 24, 25, 26,
Sry Con Dl, 55; 56, SF. fon
67, 68, 69, 70, 71, 72, 73s
74, 75s 75, 77s 78s 79s 805

512 PHYTOLOG

81, 82, 85, 84, 85, 87, 88,

89, 114, 115, 116, 117, 126,

127, 128, 135, 137, 142, 145,

146, 147, 148, 149, 150, 159,

160, 161, 162, 163, 164, 165,

1665 2355, 236, 237»

238, 241, 3521, 522,

523 326, 327, 528,

332, 3535, 536,

340, 341, 342,

348, 382, 3835,

419, 420, 421,

425, 426, 427,

476; 477, 478,

479, » 482

Verbenaceae, ys "hO, 65, 89,
Ug a 1 124, 152, 320g.00e's
388, 417, 477

Verbeneae, 243, 246, 248, ap

Veronica, 151

Viburnum, 277

Vinca, Vinca, 277

Viola, 241

Viorna, 241

Virola, 597

Vitaceae, 40

Vitex, 29, 30, 31, 65, 66, 675

psn 10s Fixes las dete

75, 163 Tis 18s 192 80; Si,
82, 83, 84, 85, 86, 87, 88,
89, 117; 118, 119, 120, 121,
122, 123, 168, 277, 329, 334,
339, 341, 342, 343, 344, rakia
348, 382, 385, 386, 387, 477,
480, 483

Publication dates of Volums 2

No. 1 — August 26, 1941
No. 2 — December 8, 1941
No. 3 — January 4, 1945

4 — July 8, 1943

5 -- September 20, 1946
6 == October 18, 1946

7 -- April 22, 1947

No.
No.
No.
No.
No. 8 == June 30, 1947
No. 10 == April 22, 1948

\

"3 S pew mete
s é. = ‘ee

. % Von

i ‘+e

Vol. 2, no. I

IA "
Vitictpremna, 69» T+, 75. 77am
783 795 87, 125 4

Vitis, 204, 205, 277
Vochysiaceae, 40 :
volatile <ORy 37» 38; 39; 65
47, 48, 49 3
Volkemeria, 25/
Weigeltia, 242
Welwitechia, 42
Winteranacese, 40
Wisteria, 2/77
Xanthoceras, 277
Xanthoxalis, 524

Xeroplana, 483
Xylophacos, 324
Xylosteum, 263
Zanthoxylum, 278
Zelkova, 2/78
Zenobia, 278
Zygophyllaceae, 40
Zygophyllum, 276

No. 9 -- December 3, 1947
No. ll -- July 9, 1948

Fe

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