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MONTANA 


This  "cover"  page  added  by  the  Internet  Archive  for  formatting  purposes 


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Preliminary 
Amphibian  and  Reptile  Survey 

of  the 
Helena  National  Forest:  1995 

STATE  DOCUMENTS  COLLECTION 

.!UN  IP  1998 

MONTANA  STATE  LIBRARY 
.,^,    1515  E.  6th  AVE. 
HELENA,  MONTANA  59S2Q 

A  Report  to: 

USDA  Forest  Service 

Helena  National  Forest 

2880  Skyway  Drive 
^,^elena,MT  59601 

•#  i%J§ M    M  irf  |t^ 


Submitted  by 

JM-IES  D.  REICHEL 

March  1996 

Montana  Natural  Heritage  Program 
1515  East  Sixth  Avenue 

P.O.  Box  201800 
Helena.  MT  59620-1800 


©  1996  Montana  Natural  Heritage  Program 


This  document  should  be  cited  as  follows: 

Reichel,  J.  D.  1996.  Preliminary  amphibian  and  reptile  survey  of  the  Helena  National  Forest:  1995.  Montana 
Natural  Heritage  Program.  Helena,  MT.  87  pp. 


11 


ABSTRACT 

A  total  of  44  sun'eys  and  several  additional  sightings  were  made  in  the  Helena  National 
Forest  (HNF)  between  May  and  August  1995.  Localized  areas  across  the  entire  forest  were 
covered  in  the  survey.  Surveys  of  ponds,  lakes,  seeps,  streams  or  other  wetlands,  made  by  1  or  2 
individuals.  Each  survey  took  10-150  person-minutes  and  consisted  of  a  thorough  search  of  the 
wetland  perimeter  and  netting  of  near  shore  aquatic  habitats  for  adults,  eggs,  larvae,  and  tadpoles. 
Stream  sampling  was  done  by  hand  and  dipnet.  Seeps  were  checked  by  rolling  over  rocks  and 
logs  in  and  near  wet  areas.  In  addition  to  surs'eys,  sightings  were  made  from  road  kills,  vocal 
identifications,  or  fortuitous  sightings  by  other  reliable  individuals. 

Four  amphibians  are  present  on  the  HNF:  Long-toed  Salamander  (Ambystoma 
macrodactylum).  Tailed  Frog  {Ascaphus  truei).  Western  Toad  {Bufo  boreas),  and  Spotted  Frog 
{Rana  pretiosa).  The  Spotted  Frog  was  the  most  widespread  amphibian  tliroughout  the  forest. 
The  Tailed  Frog  has  been  reported  from  a  single  location  on  the  Lincoln  District.  Long-toed 
Salamanders  were  found  throughout  the  main  Rocky  Mountain  chain  and  in  the  Elkliom 
Mountains.  The  Western  Toad  was  found  in  very  few  locations  on  the  HNF  in  1995,  all  in  the 
main  Rocky  Mountain  chain.  Historically  it  has  been  reported  in  the  Big  Belt  Mountains; 
however  it  was  not  found  there  during  our  surveys.  This  is  consistent  with  the  apparent  region- 
wide  declines  in  this  species.  Four  other  prairie-inliabiting  amphibians  have  been  reported  in  the 
area,  though  in  some  cases  well  away  from  HNF  lands;  these  include  the  Western  Chorus  Frog 
{Pseudacris  trisehatd),  Woodhouse's  Toad  {Bufo  woodhousii),  Plains  Spadefoot  {Scaphiopus 
bombifrons),  and  Northern  Leopard  Frog  {Rana  pipiens).  The  Western  Chorus  Frog  is  common 
in  prairie  ponds  to  the  north  and  east  of  HNF  lands;  two  reports  were  received  for  the  Helena 
National  Forest,  but  need  confimiation.  A  tadpole  reported  to  be  a  Woodhouse's  Toad  is  present 
at  the  Montana  State  University  Museum;  given  the  difficulty  in  identifying  toad  tadpoles  and 
distance  from  known  sites,  this  should  be  treated  as  hypothetical  until  verified.  The  Plains 
Spadefoot  is  known  from  the  Helena  Valley,  but  has  yet  to  be  found  in  the  ITNF.     A  report  was 
received  of  a  Northern  Leopard  Frog  from  the  vicinity  of  McDonald  Pass;  given  the  distance 
from  other  known  locations  and  unusual  habitat,  this  report  should  be  treated  as  hypothetical 
until  verified.  The  Northern  Leopard  Frog  was  also  reported  historically  from  several  prairie 
areas,  outside  and  at  lower  elevations  than  HNF  lands.  The  Deepdale  Fishing  Access  Site  had 
frogs  as  recently  as  1994,  however  three  surveys  in  1995  failed  to  relocate  them.  Northern 
Leopard  Frogs  are  nearly  extirpated  from  western  Montana,  and  recent  evidence  indicates  a 
decline  elsewhere  in  Montana  (except  perhaps  the  southeast  comer). 

Ten  reptiles  have  been  reported  from  near  the  HNF,  but  only  tliree  have  been  definitely 
reported  from  on  the  forest:  the  Racer  {Coluber  constrictor),  Western  Terrestrial  Garter  Snake 
{Thamnophis  elegans)  and  Conimon  Garter  Snake  {TJwmnophis  sirtalis).  All  were  reported  in 
the  main  Rocky  Mountains.  The  Racer  and  Western  Terrestrial  Garter  Snake  also  were  found  in 
the  Big  Belt  Mountains  and  there  is  a  record  of  the  Common  Garter  Snake  from  the  Elkliom 
Mountains.  The  following  reptiles  have  been  reported  in  the  area  and  may  eventually  be  found 
on  lower  elevation  HNF  lands:  Painted  Turtle  {Chrysemys  picta).  Spiny  Softshell  {Trionyx 
spinifera).  Short-homed  Lizard  {Phrynosoina  douglasi).  Rubber  Boa  {Charina  bottae).  Milk 
Snake  {Lcmpropeltis  triangidum),  Gopher  Snake  {Pituophis  catenifer),  and  Western  Rattlesnake 


ni 


{Crotalus  viridis).  The  Painted  Turtle  has  been  recorded  just  off  the  HNF  on  the  east  side  of  the 
Elkhom  Mountains.  The  Spiny  Softshell  is  present  in  large  rivers  at  lower  elevations;  it  has  been 
reported  from  Canyon  Ferry  Reservoir,  but  there  is  no  recent  confirmation.  Both  the  Short- 
homed  Lizard  and  Milk  Snake  are  present  in  the  area  near  Three  Forks,  south  of  the  HNF.  The 
Rubber  Boa  has  been  recorded  just  off  the  HNF  south  of  Helena  and  near  Granite  Butte;  it  surely 
occurs  on  the  HNF.  The  Gopher  Snake  has  been  reported  from  the  intermountain  valleys,  as 
close  as  Va  mile  from  the  HNF.  The  Western  Rattlesnake  also  has  been  recorded  just  off  the 
HNF;  v/iih  several  records  at  lower  elevations,  it  probably  will  eventually  be  found  on  the  HNF 
lands. 


IV 


TABLE  OF  CONTENTS 

ABSTRACT iii 

ACKNOWLEDGMENTS vii 

INTRODUCTION 1 

METHODS  AND  MATERIALS 2 

RESULTS  AND  DISCUSSION   3 

Species  kno\vn  to  be  present  on  the  Helena  National  Forest 6 

Long-toed  Salamander  {Ambystoma  macrodactylum) 6 

Tailed  Frog  {Ascaphus  truei) 8 

Western  Toad  {Bufo  boreas)   10 

Spotted  Frog  {Rana  pretiosa) 12 

Racer  {Coluber  constrictor) 14 

Western  Terrestrial  Garter  Snake  {Thamnophis  elegans)   16 

Common  Garter  Snake  {Thamnophis  sirtalis) 18 

Species  Potentially  Present  on  the  Helena  National  Forest 20 

Western  Chorus  Frog  {Pseiidacris  triseriata)  20 

Woodhouse's  Toad  {Bufo  woodhousii)    22 

Plains  Spadefoot  {Scaphiopus  [=SpeaJ  bombifrons) 24 

Northern  Leopard  Frog  {Rana  pipiens)  26 

Painted  Turtle  {Chrysemys picta) 28 

Spiny  Softshell  {Trionyx  spiniferus)  {=Apa!one  spiniferd) 30 

Short-homed  Lizard  {Phrynosoma  douglasi)    32 

Rubber  Boa  {Charina  bottae)    34 

Milk  Snake  {Lampropeltis  trianguhun) 36 

Gopher  Snake  {Pituophis  catenifer  [=melanoleucusJ)  38 

Western  Rattlesnake  {Crotahis  viridis)  40 

Regional  Information 41 

Rocky  Mountain  chain 41 

Elkhom  Mountains 41 

Big  Belt  Mountains   42 

Dry  Range   42 

RECOMMENDATIONS 43 

Surveys,  Monitoring  and  Research   43 

Management 44 

BIBLIOGRAPHY 45 


V 


Appendix  1.  Data  Sheets  used  for  Reptiles  and  Amphibian  Surveys  and  Observations    59 

Appendix  2.  Sites  surveyed  during  1995  amphibian  and  reptile  surveys   61 

Appendix  3.  Amphibians  and  reptiles  observed  during  surveys  on  or  near  the  Helena  National 
Forest  in  1995    64 

Appendix  4.  Amphibians  and  reptiles  reported  from  in  and  around  the  Helena  National  Forest 
66 

Appendix  5.  Notes  on  harlequin  surveys  and  examination  of  potential  northern  bog  lemming 
habitat  on  the  Helena  National  Forest 83 

Appendix  6.  Heritage  program  species  ranking  definitions  85 


VI 


ACKNOWLEDGMENTS 

I  would  like  to  thank  the  staff  from  the  Helena  National  Forest  for  their  assistance  in 
determining  the  location  of  possible  sur\'ey  sites,  information  on  heip  obser\'ations,  field 
assistance,  and  other  support;  they  included  Quinn  Carver,  Brent  Costain,  Doug  Grupenhoff, 
Archie  Haiper,  Shane  Hendrickson,  Connie  Jacobs,  Barry  Paulson,  Melanie  Scott,  and  Len 
Walch.  D.  D.  Dover,  J.  Hinshaw,  C.  Jones,  and  K.  Jurist  assisted  with  field  work,  data  entry,  and 
map  preparation.  Financial  support  for  the  project  came  from  the  Helena  National  Forest  (U.S. 
Forest  Service,  Northern  Region)  and  the  Montana  Natural  Heritage  Program  (Montana  State 
Library,  Natural  Resources  Information  System  and  The  Nature  Consers'ancy). 

Museum  records  were  received  from:  American  Museum  of  Natural  History,  Academy  of 
Natural  Science,  Bingham  Young  University,  California  Academy  of  Science,  Carnegie  Museum, 
University  of  Puget  Sound  Museum,  Field  Museum  of  Natural  History,  Glacier  National  Park 
Museum,  Illinois  Natural  History  Survey,  University  of  Kansas,  Los  Angeles  County  Museum, 
Louisiana  State  University  Museum  of  Zoology,  Museum  of  Comparative  Zoology  -  Harvard, 
Milwaukee  Public  Museum,  Montana  State  University  Museum,  Michigan  State  University 
Museum,  North  Carolina  State  Museum  of  Natural  History,  Northern  Louisiana  University 
Museum,  University  of  Colorado  Museum,  University  of  Georgia  Museum  of  Natural  History, 
University  of  Idaho  Museum,  University  of  Michigan  Museum,  University  of  Montana  Museum, 
University  of  South  Dakota,  United  States  National  Museum  of  Natural  History,  University  of 
Texas  -  Arlington,  University  of  Texas  -  El  Paso,  Peabody  Museum  -  Yale,  University  of 
California-Berkeley  Museum  of  Vertebrate  Zoology,  and  Mid-continental  Ecological  Sciences 
Center  at  University  of  New  Mexico  Museum  of  Southwestern  Biology.  Much  of  the  museum 
data  was  received  with  the  help  of  Dr.  Charles  Peterson,  Idaho  State  University,  Pocatello. 


's'il 


INTRODUCTION 

Many  amphibians  are  apparently  declining  in  the  western  U.S.  and  world-wide  (Com  and 
Fogelman  1984,  Phillips  1994,  Yoffe  1992).  Acid  rain,  ozone  depletion,  pollution  by  toxic 
chemicals  and  heavy  metals,  predation  and/or  competition  by  exotic  species,  habitat  alteration, 
climate  change,  disease,  immune  system  problems,  and  some  combination  of  these  factors  have 
all  been  suggested  as  possible  causes  (Blaustein  et  al.  1994a,  1994b;  Corn  and  Fogelman  1984; 
Phillips  1994;  Yoffe  1992). 

Bass  and  non-native  trout  have  been  introduced  into  waters  on  or  near  the  Helena  National 
Forest  (HNF)  and  have  been  implicated  in  declines  of  native  amphibian  populations  in  some 
areas.  Past  forestry  practices  and  large  scale  logging  continue  to  be  detrimental  to  resident 
herpetofauna  (Bury  et  al.  1991).  The  Tailed  Frog  (Ascaphiis  truei),  present  on  the  HNF,  is 
thought  to  be  one  of  the  most  sensitive  indicators  of  stream-side  and  aquatic  community  health  in 
forested  landscapes  (R.  B.  Bury,  pers.  comm.).  Preliminary  data  indicate  the  Northern  Leopard 
Frog  {Rana  pipiens)  has  disappeared  over  much  of  its  former  range  in  western  Montana  and  is 
declining  in  at  least  some  areas  of  eastern  Montana  (Hendricks  and  Reichel  in  review;  Reichel 
1995a,  1995b;  Werner  and  Reichel  1994,  1996).  The  US  Fish  and  Wildlife  Service  now  lists  the 
Western  Toad  {Biifo  boreas)  as  a  Candidate  (C-1)  species  in  Colorado,  Wyoming  and  New 
Mexico.  Apparent  declines  have  recently  been  reported  in  northern  Idaho  (C.  Peterson  pers. 
comm.),  northwest  Montana  (Reichel  and  Flath  1995;  Werner  and  Plumber  1995;  Werner  and 
Reichel  1994,  1996),  Yellowstone  National  Park  (Koch  and  Peterson  1995),  Wyoming,  and 
Colorado  (Carey  1993). 

The  U.S.  Fish  and  Wildlife  Service  listed  two  Montana  amphibians  and  two  reptiles  as 
Candidate  (C2)  species:  the  Spotted  Frog  {Rana pretiosa).  Tailed  Frog,  Short-homed  Lizard 
{Phrynosoma  douglasi)  and  Northern  Sagebmsh  Lizard  {Sceloporus  graciosus  graciosus).  The 
U.S.  Forest  Service  Region  1  lists  the  Coeur  d'Alene  Salamander  {Plethodon  idahoemis)  as 
"Sensitive"  and  is  considering  adding  several  other  amphibians.  The  Montana  Natural  Heritage 
Program  and  the  Montana  Department  of  Fish,  Wildlife  and  Parks  list  6  amphibians  [Coeur 
d'Alene  Salamander,  Idaho  Giant  Salamander  {Dicamptodon  aterrimns).  Tailed  Frog,  Canadian 
Toad  {Bufo  hemiophrys).  Spotted  Frog,  Wood  Frog  {Rana  sylvatica)]  and  7  reptiles  [Snapping 
Turtle  {Chelydra  serpentina).  Spiny  Softshell  {Trionyx  spiniferus).  Short-homed  Lizard, 
Sagebrush  Lizard,  Western  Hognose  Snake  [Heterodon  nasicus).  Smooth  Green  Snake 
{Opheodrys  vemalis).  Milk  Snake  {Lampropeltis  triangulum)]  as  species  of  special  concem  in 
the  state.  The  Northern  Leopard  Frog  and  Western  Toad  are  being  considered  for  addition  to  the 
species  of  special  concem  list;  currently  they  on  the  watch  list.  Seven  of  these  species,  the  Tailed 
Frog,  Western  Toad,  Spotted  Frog,  Northern  Leopard  Frog,  Spiny  Softshell,  Short-homed  Lizard, 
and  Milk  Snake  occur  or  potentially  occur  on  the  HNF. 


METHODS  AND  MATERIALS 

Historic  locations  of  amphibians  and  reptiles  were  recorded  from  literature  (see 
Bibliography)  and  museum  specimen  records.    Records  were  received  from  over  20  major 
museum  collections  in  North  America  (see  Acknowledgments).  Locations  derived  from  these 
sources  have  been  entered  into  a  database  and  digitized. 

Survey  sites  were  chosen  based  on  4  criteria:  1)  high  priority  sites  as  determined  by  the  ILNF; 
2)  location  of  streams,  seeps  and  wetlands  on  topographic  maps;  3)  accessibility  of  the  wetlands 
by  roads  or  hiking  trails;  and  4)  conversations  with  district  biologists  regarding  stream-seep- 
wetland  locations.  Based  on  the  above,  2-8  sites  were  chosen  daily  for  surveys.  A  total  of  10- 
150  person-minutes  were  spent  at  each  site,  depending  upon  the  size  of  the  area  and  what  was 
found.  Initially,  the  entire  shoreline,  or  a  major  part  thereof,  was  searched  by  walking  slowly 
along  the  edge  and  up  into  the  surrounding  vegetation,  including  rolling  over  rocks  and  logs.  At 
regular  intervals,  the  aquatic  habitat  was  sampled  for  tadpoles  or  larvae  using  dipnets.  If  the 
initial  sampling  showed  amphibian/reptile  species  present,  further  effort  was  expended  in  order 
to  get  some  idea  of  abundance  and  distribution. 

An  attempt  was  made  to  capture  at  least  the  first  few  individuals  of  a  species  seen  at  a  survey 
site.  The  species  name  was  recorded  along  with  developmental  stage  and  sex  (if  possible);  the 
animals  were  then  released.  Representative  samples  of  the  more  common  species  in  an  area  were 
preserved  for  permanent  museum  records  and  will  be  deposited  at  the  Idaho  State  University 
Museum.  Water  temperature,  air  temperature,  pH,  a  general  description  of  the  area,  and  other 
parameters  were  recorded.  Standard  data  sheets  used  during  this  project  are  given  in  Appendix 
1;  the  amphibian  survey  data  sheet  was  developed  by  U.S.  Fish  and  Wildlife  Service  and  is  used 
extensively  by  a  variety  of  researchers  in  the  western  U.S.  Much  site-specific  data  was  gathered 
during  these  surveys;  not  all  data  has  been  analyzed  or  is  presented  in  this  report,  but  is  available 
from  the  Montana  Natural  Heritage  Program. 

Natural  Heritage  Program  species  status  ranking  definitions  and  explanations  are  given  in 
Appendix  6. 


RESULTS  AND  DISCUSSION 

A  total  of  47  sites  were  surveyed  of  which  29  had  one  or  more  amphibian  or  reptile  species 
present  (Figure  1,  Appendices  2  and  3);  one  site  was  surveyed  two  times.  Although  no  species 
were  found  at  1 8  sites,  their  absence  may  have  been  due  to  the  time  of  day,  weather  conditions, 
or  other  factors  at  the  time  of  sampling.  With  three  exceptions,  all  of  the  sites  were  on  HNF 
land. 

In  addition  to  the  48  sur\'eys,  there  were  a  number  of  sightings  (i.e.  road  kills,  chance 
observations)  for  which  data  are  available  and  the  sightings  considered  reliable.  Species  location 
data  from  surveys,  chance  encounters,  and  historic  records  (from  the  literature  and  museum 
specimens)  are  listed  in  Appendix  4.  Distribution  maps  were  created  using  survey  and  sighting 
data  and  historical  records;  inset  statewide  maps  for  each  species  are  based  on  sight  and 
specimen  records,  both  recent  and  historic. 

No  previous  publications  or  reports  on  reptiles  or  amphibians  concentrate  on  the  HNF  area. 
Based  on  museum  specimens,  publications,  surveys  and  incidental  obsers'ations,  four 
amphibians  and  three  reptiles  have  been  located  on  the  HNF;  an  additional  four  amphibians  and 
seven  reptiles  may  eventually  be  found  to  occur  there.  Tliree  amphibian  and  one  reptile  species 
were  actually  obser\'ed  during  the  study.  The  following  results  are  presented  as  individual 
species  summaries  for  the  Forest  as  a  whole,  followed  by  specific  infonnation  on  each  mountain 
range. 

In  the  following  species  accounts,  the  section  on  "Similar  Species"  covers  species  only  which 
are  known  or  suspected  to  occur  in  Montana;  outside  Montana  other  confusing  species  may  occur 
which  are  not  covered  in  this  report.  Photos  of  all  Montana  amphibians  and  reptiles  may  be 
found  in  Reichel  and  Flath  (1995).  Keys  to  amphibian  eggs  (Livezey  and  Wright  1947)  and 
tadpoles  (Altig  1970)  are  a\'ailable  in  the  literature,  but  are  difficult  to  use,  and  for  many  species 
are  not  satisfactory  for  field  identification. 


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Species  known  to  be  present  on  the  Helena  National  Forest 

Long-toed  Salamander  {Ambystoma  macrodactyhim) 

Description:  Adults  are  dart:  gray  to  black  with  an  irregular  (and  sometimes  broken)  green  to 
yellow  stripe  down  the  middle  of  the  back.  Adult  snout-vent  length  varies  from  2  to  3.25". 
All  salamanders  have  smooth  moist  skin  without  scales. 

Eggs  and  Larvae:  Egg  masses  are  typically  laid  in  small  clusters  of  5-100  eggs  but  may  be 
laid  singly  (Nussbaum  et  al.  1983);  egg  masses  are  typically  attached  to  underwater 
vegetation  or  submerged  branches.  Within  the  clear  gelatinous  eggs,  the  embryos  are 
somewhat  light-colored,  while  frog  and  toad  embr>'os  are  dark  (e.xcept  in  Tailed  Frogs). 
Larval  Long-toed  Salamanders  are  typically  brown-  or  gray-colored,  are  found  in  ponds,  have 
tliree  external  gills,  and  are  relatively  small  (<1.75"  snout-vent)  and  slender.  They  are 
distinguished  from  Tiger  Salamander  larvae  by  the  9-13  gill  rakers  on  the  inside  of  the  3rd 
gill  arch  (17-22  rakers  on  the  Tiger  Salamander);  they  are  also  smaller  and  lack  the  large  head 
and  mouth. 

Similar  species:  Adult  Long-toed  Salamanders  can  be  distinguished  from  Coeur  d'Alene 

Salamanders  by  the  longest  toe  on  the  hind  foot  which  is  longer  than  the  sole  and  a  yellow 
tliroat  patch.  Long-toed  Salamanders  lack  a  groove  running  vertically  from  nostril  to  mouth. 

Habitat  and  Habits:  Long-toed  Salamanders  are  found  in  a  wide  variety  of  habitats  from 
sagebrush  to  nearly  alpine.  They  breed  in  ponds  or  lakes  (very  rarely  in  slow  moving 
streams),  usually  those  without  fish  present;  on  the  HNF  they  were  found  in  temporary  and 
permanent  ponds/lakes  and  inactive  beaver  ponds.  Adults  go  to  the  breeding  ponds 
immediately  after  snow-melt  and  are  usually  the  earliest  breeding  amphibians  in  western 
Montana.  In  the  Pacific  Northwest,  eggs  hatch  in  3-6  weeks  and  metamorphosis  occurs  after 
2-14  months  (Nussbaum  e/o/.  1983,  Leonard  e/o/.  1993).  Long-toed  Salamanders  were 
found  in  14  locations  on  the  HNF.  The  earliest  surveys  on  15-25  May  1995  found  only  egg 
masses,  from  newly  laid  to  nearly  ready  to  hatch.  July  surveys  found  primarily  small  larva, 
however,  a  pond  near  the  head  of  Austin  Creek  had  2  small  larva  and  1  juvenile  which 
appeared  nearly  fully  transformed;  this  would  indicate  at  least  some  larva  overwinter  and 
transfonn  when  over  a  year  old.  At  an  o.xbow  on  the  Blackfoot  River  a  single  transforming 
juvenile  war  found  on  25  August,  showing  transformation  may  take  only  one  season  at  lower 
elevation  sites.    Individuals  were  found  in  the  Rocky  Mountains  and  the  Elkhorn  Mountains 
from  4350  -  7050  ft.  elevation.  Spotted  Frogs  co-occurred  at  all  sites  and  Western  Toads  at  2 
sites. 

Surveying:  Larvae  can  readily  be  seen  in  ponds  during  the  day  and  sampled  with  a  dipnet;  egg 
masses  are  somewhat  harder  to  see.  During  the  breeding  season,  adults  may  also  be  seen  in 
the  water,  particularly  during  night  surveys.  During  the  rest  of  the  spring,  summer  and  fall, 
adults  may  occasionally  be  found  in  and  under  logs  on  the  forest  floor.  Metamorphosed 
individuals  are  active  at  night,  particularly  when  it  is  wann  and  rainy;  they  may  be  captured  at 
tlais  time  by  either  night  searches  or  pitfall  traps. 

Status:  The  Long-toed  Salamander  is  the  most  common  salamander  in  western  Montana.  The 
Long-toed  Salamander  is  also  common  on  the  HNF  in  the  Rocky  Mountains.  The  Long-toed 
Salamander  is  also  found  in  the  northwestern  part  of  the  Elklioms,  the  farthest  east  reported 


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location  in  its  range.  The  Eikhoms  are  the  only  isolated  mountain  range  east  of  the 
Continental  Divide  where  this  species  is  found.  However,  none  were  found  in  the  southern 
Elklioms  or  Big  Belt  Mountains.  In  the  Lewis  and  Clark  National  Forest  (L&CNF)  it  was 
found  to  be  very  local  in  distribution  east  of  the  Continental  Divide  and  was  not  found  in  the 
isolated  ranges  to  the  east  (Reichel  1995a).  Given  the  unique  position  of  the  Elkliorn 
populations  more  surveys  should  be  done  to  better  understand  and  document  the  distribution 
there. 
Montana  Natural  Heritage  Program  rank:  G5  S5. 

Tailed  Frog  {Ascaphus  truei) 

Description:  Adults  are  gray  or  brown  with  gray,  brown,  or  occasionally  yellow  blotches;  the 
skin  has  a  distinctly  bumpy  texture.  The  adult  has  a  snout-vent  length  of  1.5-2"  and  lacks  a 
tympanum.  The  outer  toe  of  the  hind  foot  is  broader  than  the  other  toes.  The  male  has  a 
bulbous  "tail"  which  acts  as  a  penis. 

Eggs  and  Larvae:  Approximately  50  eggs  are  laid  in  rosary-like  strings  attached  to  the 
underside  of  rocks.  The  tadpole  (up  to  2"  long)  is  unique  in  that  it  has  a  large  mouth 
modified  into  a  sucker;  the  color  is  quite  variable. 

Similar  species:  No  other  frog  or  toad  has  the  outer  toe  of  the  hind  foot  broader  than  the  other 
toes;  all  other  frogs  and  toads  have  a  tympanum  behind  each  eye. 

Habitat  and  Habits:  Tailed  Frogs  are  found  in  and  along  small,  swift,  cold  mountain  streams.  In 
the  Cascade  Mountains  of  Washington  and  Oregon,  the  Tailed  Frog  appears  to  be  very 
sensitive  to  siltation  and  frequently  disappears  in  and  downstream  from  clearcuts  and  water 
diversions  (Bury,  pers.  comm.).  Preliminary  findings  do  not  indicate  that  this  is  the  case  in 
Montana.  Eggs  are  laid  during  the  late  summer  and  take  approximately  4  weeks  to  hatch. 
Tadpoles  take  1-4  years  to  metamorphose,  depending  on  water  temperature  (Nussbaum  et  al. 
1983;  Metter  1967).  Sexual  maturity  in  Montana  is  attained  at  ages  6-7,  (Daugherty  and 
Sheldon  1982)  which  is  the  latest  age  for  sexual  maturity  of  any  North  American  amphibian. 

Surveying:  Tadpoles  are  frequently  found  while  electro-shocking  fish.  They  may  also  be  found 
by  turning  over  rocks  in  rapid  water  with  a  net  held  just  downstream.  Adults  are  best  found 
by  walking  up  streams  starting  30-60  minutes  after  dark. 

Status:  The  Tailed  Frog  on  the  HNF  is  known  from  a  single  observation  from  the  late  1960s  on  a 
"tributary  of  Copper  Creek"  (Franz  1971).  While  Tailed  Frogs  should  be  considered  a 
species  with  a  very  localized  distribution  on  the  HNF,  it  may  be  more  common  and 
widespread  in  suitable  habitat  than  is  currently  known.  It  should  be  looked  for  tliroughout  the 
Forest,  especially  to  the  south  and  east  of  the  currently  known  location.  East  of  the 
Continental  Divide  on  the  L&CNF  it  was  also  found  to  be  very  local  in  distribution  (Reichel 
1995a).    It  seem  doubtful  that  the  Tailed  Frog  is  found  in  the  Elkhorn  Mountains  given  the 
intensity  of  surveys  (A.  Harper,  pers.  comm.),  and  the  surveyors  specifically  watching  for  the 
species,  during  the  summer  of  1995.  It  is  common  and  widespread  in  northwestern  Montana 
(Reichel  and  Flath  1995,  Werner  and  Reichel  1994,  1996).  Previously  it  was  a  USFWS 
Candidate  species  (C-2).  We  would  recommend  that  all  sightings  of  this  species  be  reported. 

Montana  Natural  Heritage  Program  rank:  G5  S3S4. 


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Western  Toad  {Bufo  boreas) 

Description:  Adults  are  colored  with  a  gray,  browTi,  or  olive-green  mottling  and  a  prominent 
white  or  yellowish  line  down  the  center  of  the  back;  very  young  transformed  toads  typically 
lack  the  dorsal  line,  and  the  warts  are  often  red-brown  in  color.  The  pupils  are  horizontal. 
The  adult  has  a  body  length  of  2.5-5".  There  are  no  cranial  crests  and  the  skin  is  relatively 
dry  with  many  warts  and  glands  present. 

Eggs  and  Larvae:  Eggs  are  laid  in  long,  clear,  double  strings,  and  each  has  a  black  embryo. 
Tadpoles  are  typically  jet  black,  while  all  mid-  to  large-sized  frog  tadpoles  in  Montana  are 
green  or  bronze  (except  for  some  Tailed  Frogs);  very  small  frog  tadpoles  are  also  black. 

Similar  species:    Other  Montana  toads  have  cranial  crests  between  their  eyes.  The  Plains 
Spadefoot  has  one  tubercle  on  the  sole  of  the  hind  feet,  a  vertical  pupil,  and  smoother  skin. 
NOTE:  It  is  very  difficult  to  distinguish  among  the  four  Montana  toad  species  eggs,  larvae, 
and  recently-transformed  toadlets. 

Habitat  and  Habits:  Adults  are  largely  terrestrial  and  found  in  a  variety  of  habitats  from  valley 
bottoms  to  high  elevations;  they  breed  in  lakes,  ponds,  and  slow  streams  with  a  preference  for 
shallow  areas  with  mud  bottoms.  Breeding  and  egg  laying  in  Montana  usually  takes  place  1- 
3  months  after  snow-melt,  from  April  at  lower  elevations  to  July  at  higher  sites.  Tadpoles  are 
typically  2-3  months  old  at  metamorphosis  in  Montana,  depending  on  water  temperature 
(Black  1970).  Following  metamorphosis,  hundreds  of  small  toads,  many  with  the  tails  still 
present,  can  be  found  on  the  shores  of  breeding  ponds.  Western  Toads  were  reported 
breeding  from  only  two  locations  on  the  HNF  in  1995.  We  found  thousands  of  Vi  grown 
tadpoles  in  a  pond  near  Dog  Creek  on  1 1  July  1995  and  3  metamorphs  on  an  oxbow  along  the 
Blackfoot  River  on  25  August. 

Surveying:  Tadpoles  are  easily  seen  in  ponds  during  the  day  and  can  be  sampled  with  a  dipnet. 
During  the  breeding  season,  adults  may  be  seen  in  the  water  but  at  other  times  are  found  in 
more  terrestrial  habitats. 

Status:  Tadpoles  and  metamorphs  of  the  Western  Toad  were  observed  at  only  two  sites  during 
the  1995  survey  in  the  HNF  (Appendix  2,  3).  No  adults  were  seen  during  our  surveys 
however,  B.  Spettigue  (pers.  comm.)  saw  one  adult  near  McDonald  Pass  (Appendix  4).  None 
were  seen  in  the  Elkhom  or  Big  Belt  Mountains,  although  historic  records  exist  for  sites  in  or 
near  both  (Appendix  3,4).  The  rarity  of  this  species  on  the  HNF  and  lack  of  recent  sightings 
in  the  eastern  ranges  is  of  concern.  During  1994  we  found  no  Western  Toads  in  the  Little 
Belt,  Highwood,  or  Crazy  Mountain  of  the  L&CNF  although  historic  records  exist  for  all 
three  ranges;  additionally  only  a  single  breeding  site  was  located  on  the  main  Rocky 
Mountains  of  the  L&CNF  (Reichel  1995a).  Brunson  (1952)  regarded  the  Western  Toad  as 
one  of  the  most  common  batrachians  (frogs  and  toads)  in  western  Montana.    Black  (1970) 
supported  its  common  occurrence  not  only  in  the  west  but  in  many  counties  east  of  the 
continental  divide.  The  Western  Toad  has  declined  from  the  most  common  anuran  in 
western  Montana,  to  a  relatively  rare  one  in  the  state  in  the  past  25  years  (Reichel  and  Flath 
1995,  Werner  and  Plumber  1995,  Werner  and  Reichel  1994,  1996). 

The  U.S.  Fish  and  Wildlife  Service  now  lists  this  species  as  a  Candidate  (C-1)  species  in 
Colorado,  Wyoming,  and  New  Mexico.  Apparent  declines  have  recently  been  reported  in 
northern  Idaho  (C.  Peterson  pers.  comm.),  Yellovv'stone  National  Park  (Koch  and  Peterson 


10 


1995,  Peterson  et  al.  1992),  Wyoming,  and  Colorado  (Carey  1993).  We  would  recommend 
that  all  sightings  of  this  species  be  reported  and  that  a  monitoring  program  be  set  up  for  this 
species. 
Montana  Natural  Heritage  Program  rank:  G4  S3S4. 

Spotted  Frog  {Rana pretiosd) 

Description:  The  adult  has  a  snout-vent  length  of  2-4".  Adults  are  dark  to  light  brown,  gray,  or 
olive  green  with  dark  spots  (frequently  with  lighter  centers)  found  on  the  back,  sides  and  legs. 
The  number  and  pattern  of  spotting  is  quite  variable.  The  back  and  sides  are  often  covered 
with  small  bumps.  The  underside  of  the  legs  is  bright  red,  salmon,  or  orange;  this  bright 
color  may  extend  up  to  the  chin  or  be  replaced  by  a  light,  mottled  gray  on  the  chin,  chest, 
and/or  belly.  In  younger  subadults,  bright  leg  color  is  often  lacking  and  instead  a  light, 
lemon-colored  wash  is  present.  In  these  subadults  there  is  often  a  dark  mask  present,  with  a 
light  jaw  stripe  extending  to  the  shoulder;  both  the  mask  and  jaw  stripe  may  be  less  obvious 
in  larger,  older  animals. 

Eggs  and  Tadpoles:    Eggs  are  laid  in  large,  globular  masses  of  1 50-500  at  the  surface  of  the 
water.  The  tadpoles  are  dark  green  to  brown  on  top  with  some  gold  flecking  whereas  the 
underside  has  an  iridescent  bronze  or  silver  color.  Total  length  of  tadpoles  may  reach  3";  the 
eyes  are  located  on  top  of  the  head. 

Similar  species:    The  bright-colored  pigment  on  the  undersides  of  the  adult's  legs  distinguish  this 
species  from  all  other  fi-ogs  in  Montana.  Younger  individuals,  without  colored  legs,  may 
usually  be  distinguished  from  other  frogs  by  a  combination  of:  1)  dorsal  spots  usually  present 
but  not  surrounded  by  light-colored  halos;  2)  dorsolateral  folds  present;  3)  toes  without  pads 
at  the  tips;  and  4)  a  pale  gray,  (rather  than  white)  belly. 

Haoitat  and  Habits:    Spotted  Frogs  are  regularly  found  at  the  water's  edge  in  openings  within 
forest  habitats.  Wetlands  in  or  near  treeline  are  also  used,  but  populations  are  uncommon  in 
the  large,  open  intermountain  valleys.    Eggs  hatch  in  2-3  weeks  and  tadpoles  take  2-14 
months  to  metamorphose,  depending  on  water  temperature  (Nussbaum  et  al  1983,  Turner 
1958).  Breeding  takes  place  in  lakes,  ponds  (temporary  and  permanent),  springs,  and 
occasionally  backwaters  or  beaver  ponds  in  streams.  All  the  egg  masses  in  a  particular  pond 
are  often  found  in  the  same  location  at  the  margin  of  the  pond;  therefore,  the  eggs  are 
susceptible  to  drying  up  if  pond  levels  recede  substantially  before  hatching.  Young  and  adult 
fi-ogs  often  disperse  into  marsh  and  forest  habitats  but  are  not  usually  found  far  from  open 
water.  The  Spotted  Frog  was  commonly  found  throughout  the  HNF  from  just  above  the 
prairie  edge  at  4350  to  7050  ft.  elevation  near  timberline.  Individuals  were  found  in  every 
type  of  wetland  habitat,  although  numbers  varied  widely  from  one  to  50  or  more  per  site. 
Eggs  were  found  on  the  first  sur.'eys  on  1 5  May  95  at  two  locations  in  the  Elkhoms  and  as 
late  as  25  May  (last  spring  sur\'eys).  Hatching  tadpoles  were  seen  on  17  May  95  in  the  Big 
Belts;  mid-large  tadpoles  were  present  during  July  surveys  and  only  metamorphs  were  seen 
by  late  August.  The  largest  group  of  egg  masses  I  have  seen  was  found  on  20  May  in  an  old 
beaver  pond  on  a  tributary  of  Beaver  Creek  (Appendix  2);  I  estimated  120  egg  masses  were 
present  covering  an  area  about  6  X  8  ft. 

Sur\'eying:    Adults,  tadpoles,  and  eggs  are  easily  seen  in  and  along  the  ^^'ater  during  the  da;>'  l.-:-:: 


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can  be  sampled  with  a  dipnet;  adults  may  also  be  captured  by  hand.  Many  adults  may  leave 
the  breeding  ponds  following  egg  laying  and  move  to  nearby  feeding  areas  for  the  summer. 
Tadpoles  are  difficult  to  distinguish  from  those  of  the  Northern  Leopard  Frog  in  areas  where 
the  two  species  may  overlap. 

Status:  The  most  common  frog  on  the  ITNF  and  in  western  Montana.  It  was  observed  in  all  areas 
sur\'eyed  on  the  HNF,  and  occurred  in  all  28  sites  where  any  amphibian  was  species  found. 
However,  it  appeared  to  be  much  less  common  in  the  Big  Belt  Mountains  than  the  Elkliorns 
or  main  Rocky  Mountains.  The  species  was  previously  a  U.S.  Fish  and  Wildlife  Service 
Category  2  Candidate  species  in  Montana;  elsewhere  in  its  range  it  is  listed  as  a  C-1,  with 
Threatened/Endangered  status  warranted  but  precluded  by  work  on  higher  priority  species 
(U.S.  Fish  and  Wildlife  Service  1993).  While  significant  declines  are  known  from  the 
southern  end  of  the  range  (Nevada,  southern  Idaho,  Utah)  and  are  also  apparent  in  coastal 
Washington  (McAllister  et  al.  1993),  Oregon,  and  California,  recent  (as  yet  unpublished) 
research  indicates  that  those  populations  are  actually  different  species. 

Montana  Natural  Heritage  Program  rank:  G4  S4. 


Racer  {Coluber  constrictor) 

Description:  A  slender,  but  moderately  long  snake,  the  Racer  ranges  from  20-65  inches  in  length. 
Adult  coloration  is  uniform  across  the  dorsal  side  but  it  can  vary  from  a  greenish-gray  to 
brown  or  blue.  The  ventral  side  is  whitish  to  pale  yellow,  the  latter  color  extending  onto  the 
upper  lip  scales  and  nasal  region.  The  eyes  are  relatively  large.  The  scales  are  smooth  and 
the  nostril  is  bordered  by  two  scales. 

Young:    Snakes  (up  to  about  20")  have  a  much  different  coloration  than  the  adults  consisting 
of  a  series  of  dorsal  brown  blotches  edged  with  black  which  run  the  length  of  the  animal;  a 
row  of  blotches  is  also  found  on  each  side  of  the  animal  extending  onto  the  ventral  side. 

Similar  species:  Young  Gopher  Snakes  may  be  distinguished  by  the  keeled  rather  than  smooth 
scales  of  the  young  Racer.  Young  Western  Hognose  Snakes  have  an  upturned  nose.  Smooth 
Green  Snakes  are  smaller  and  colored  bright  grass-green  and  whitish  below;  their  nostrils  are 
centered  in  single  scales.  Also  see  Rubber  Boa. 

Habitat  and  Habits:  The  Racer  is  associated  with  open  habitats,  either  in  shortgrass,  slirub-steppe, 
or  forested  areas  (Hammerson  1982a,  Ba.xter  and  Stone  1985).  It  is  often  found  near  water 
and  rocks.  The  Racer  is  an  extremely  fast  and  agile  snake.  A  clutch  of  perhaps  3-7  eggs  is 
laid  in  the  summer  (Stebbins  1985).  It  preys  on  insects  and  small  vertebrates  such  as  mice 
and  frogs. 

Surveying:  They  may  be  sur\'eyed  for  by  slowly  walking  through  appropriate  habitat  on  warm, 
sunny  days  and  carefiiUy  watching  for  them;  this  technique  is  moderately  effective  for  the 
Racer.  However,  as  with  many  lizards  and  snakes,  they  may  easily  be  missed.  Carefully 
documented  incidental  obsen'ations  may  provide  the  best  clues  to  their  distribution.  They 
may  be  also  taken  in  funnel  traps  with  drift  fences.  Mark-recaplTjre  methods  offer  the  best 
opportunity  for  determining  population  status. 

Status:  The  Racer  was  not  seen  in  this  sur\'ey,  however,  Davis  (1963)  has  a  map  of  this  species 
which  shows  records  apparently  on  the  HNF.  A  more  recent  sighting  was  made  just  off  the 


14 


Seiben  Ranch  below  the  Forest.  They  would  be  expected  to  occur  on  the  HNF  at  low  to  mid- 
elevations;  given  the  poor  accuracy  of  the  Davis  (1963)  map,  any  sightings  should  be 
documented.  Of  particular  interest  would  be  documentation  of  any  denning  sites  located. 
Montana  Natural  Heritage  Program  Rank:  G5  S5. 


Western  Terrestrial  Garter  Snake  {Thanmophis  elegans) 

Description:  Adult  Western  Terrestrial  (or  Wandering)  Garter  Snakes  are  smaller  in  body  size 
than  the  Common  Garter  Snake,  their  length  varying  from  18-43".  Tliree  yellow  longitudinal 
stripes  are  present  (one  dorsal,  two  lateral  on  the  2nd  and  3rd  scale  rows),  but  the  dorsal 
stripe  is  much  narrower  than  that  of  the  Common  Garter  Snake.  A  distinctive  feature  of  the 
Western  Terrestrial  Garter  Snake  is  a  series  of  alternating  black  spots  which  run  the  length  of 
the  body  between,  and  somewhat  on,  the  yellow  stripes.  The  background  color  between  the 
stripes  is  a  dusky  gray,  green  or  brown,  compared  to  the  black  or  occasionally  dark  green 
found  in  the  Common  Garter  Snake.  The  ventral  surface  has  a  series  of  dark  black/bro\\'n 
blotches  which  may  cover  most  of  the  surface.  All  black,  presumably  melanistic,  individuals 
are  occasionally  found  near  Townsend.  The  dorsal  scales  are  keeled  and  there  are  normally  8 
upper  labial  scales. 
Young:  The  coloration  of  young  snakes  is  similar  to  that  of  the  adults;  young  are  live-bom. 

Similar  species:  See  Common  Garter  Snake. 

Habitat  and  Habits:  The  habitat  and  habits  of  the  Western  Terrestrial  Garter  Snake  are  similar  to 
the  Common  Garter  Snake,  i.e.,  they  are  found  in  most  habitats  but  are  particularly  common 
around  wetlands.  In  the  HNF  area  the  species  was  found  between  3450  ft  in  the  valley 
bottom  and  6200  ft  near  McDonald  Pass,  but  probably  occurs  higher.  Females  give  birth  to 
4-19  young  during  the  summer  (Stebbins  1985). 

Surveying:  Timed  sight  surveys  may  be  conducted  around  wetlands  and  riparian  feeding  areas  or 
at  denning  areas  where  higher  concentrations  of  garter  snakes  occur;  clear  mornings  are  the 
best  survey  times.  Much  distributional  information  may  come  from  recording  incidental 
sightings.    More  intensive  research  may  be  done  using  funnel  traps  in  combination  with  drift 
fences.  More  intensive  research  and  survey  projects  may  use  mark-recapture  or 
radiotelemetry  techniques. 

Status:  Western  Terrestrial  Garter  Snakes  were  found  in  the  main  Rocky  Mountains  and  Big  Belt 
Mountains  on  the  HNF;  there  are  also  historic  locations  just  off  the  forest  at  the  base  of  the 
Elkhom  Mountains.  Given  the  small  number  of  recent  records  from  tliroughout  the  area,  all 
records  should  be  documented  until  the  distribution  is  better  understood;  of  particular  interest 
would  be  documentation  of  denning  sites.  Sightings  of  elegans  from  the  Elkliom  Mountains 
should  be  documented  to  confinii  their  presence  in  that  range. 

Montana  Natural  Heritage  Program  Rank:  G5  S5. 


16 


Common  Garter  Snake  {Thamnophis  sirtalis) 

Description:  The  Common  Garter  Snake  consists  of  two  color  phases  in  western  Montana,  both 
ranging  from  1 8-52"  in  length.  Both  phases  have  tliree  yellow  longitudinal  stripes:  one 
located  dorsally  and  one  on  each  side  on  the  2nd  and  3rd  scale  rows  above  the  belly  scales. 
Between  the  yellow  stripes  is  a  black  (or  dark  green)  background,  broken  with  red  spots  in 
one  color  phase  but  lacking  red  in  the  other.  Ventral  coloration  varies  from  yellow  to  bluish, 
and  some  individuals  of  the  red-sided  color  phase  have  small  black  spots  on  the  edge  of  the 
ventral  scales.  The  dorsal  scales  are  keeled,  and  normally  there  are  7  upper  labial  scales. 
Young:  The  coloration  of  young  snakes  is  similar  to  that  of  the  adults;  young  are  live-born. 

Similar  species:  The  Western  Terrestrial  Garter  Snake  has  black  spots  overlapping  the  dorsal 
yellow  stripe;  the  background  color  between  stripes  tends  to  be  paler  dusky  green,  gray  or 
browTi.  The  Plains  Garter  Snake  has  the  side  yellow  stripe  on  the  3rd  and  4th  scale  rows 
above  the  belly  scales  and  the  dorsal  stripe  is  often  orange  or  red. 

Habitat  and  Habits:  Garter  snakes  are  found  in  all  forest  habitats  but  are  more  common  at  lower 
elevations  around  marsh-bog-pond  situations,  where  they  prey  on  young  fish,  frogs,  toads, 
mice  and  invertebrates.  They  are  sometimes  confused  with  water  snakes  because  of  their 
frequent  aquatic  exploits,  but  there  are  no  true  water  snakes  in  Montana.  Typical  of  most 
garter  snakes,  they  emit  a  noxious  secretion  when  handled  and  can  be  aggressive  when 
disturbed.  The  Common  Garter  Snake  was  reported  at  two  locations  in  1995,  both  about 
5600  feet,  on  the  HNF  by  A.  Harper  and  S.  Hendrickson;  historically  they  have  also  been 
found  at  low  elevations  in  the  intermountain  valleys.  Garter  snakes  eat  a  variety  of 
vertebrates  and  invertebrates,  with  the  Common  Garter  Snake  concentrating  more  on 
amphibians  than  the  Western  Terrestrial  Garter  Snake.  The  Common  Garter  Snake  is  a  live- 
bearer  giving  birth  to  12-18  young  during  the  summer  in  Colorado  (Hammerson  1982a). 

Surveying:  Timed-sight  surx'eys  may  be  conducted  around  wetlands  and  riparian  feeding  areas  or 
at  denning  areas  where  higher  concentrations  of  garter  snakes  occur;  clear  mornings  are  the 
best  survey  times.  Much  distributional  information  may  come  from  recording  incidental 
sightings.    More  intensive  research  may  be  done  using  funnel  traps  in  combination  with  drift 
fences.  More  intensive  research  and  survey  projects  may  use  mark-recapture  or 
radiotelemetry  tecliniques. 

Status:  Common  Garter  Snakes  were  found  in  the  Rocky  Mountains  and  Elkhom  Mountains  on 
the  HNF.  There  is  also  a  historic  record  of  them  from  the  valley  between  the  two  locations. 
Given  the  small  number  of  records  from  throughout  the  area,  all  records  should  be 
documented  until  the  distribution  is  better  understood;  of  particular  interest  would  be 
documentation  of  denning  sites.  Sightings  of  sirtalis  from  the  Big  Belt  Mountains  and  Rocky 
Mountains  north  of  McDonald  Pass  should  be  documented  to  confirm  their  presence  in  those 
areas.  Only  the  red-sided  color  phase  has  observed  in  this  area,  however  the  color  phase 
lacking  red  spots  should  be  watched  for.  The  Common  Garter  Snake  is  much  less  abundant 
than  the  Western  Terrestrial  Garter  Snake  in  this  area,  as  it  is  currently  in  northwestern 
Montana. 

Montana  Natural  Heritage  Program  Rank:  G5  S4. 


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Species  Potentially  Present  on  the  Helena  National  Forest 

Western  Chorus  Frog  (Pseudacris  thseriata) 

Description:  Adults  are  very  small  (0.75-1 .5")  and  have  tiny,  almost  unnoticeable  toe  pads. 

They  have  a  dark  line  extending  from  the  snout  thiough  the  eye  to  the  groin.  Basic  coloration 
is  quite  variable  with  the  background  color  being  green,  brown,  gray,  or  reddish.  Typically  3- 
5  dark  longitudinal  stripes  ai-e  present  on  the  head  and  back  which  may  be  broken  up  into 
spots  on  some  individuals. 

Eggs  and  Tadpoles:  Eggs  are  laid  in  small  clusters  of  10-100,  usually  less  than  1"  across  and 
attached  to  submerged  vegetation  (Wheeler  and  Wheeler  1966,  Baxter  and  Stone  1985). 
Individual  eggs  are  about  1  mni  in  diameter.  Tadpoles  are  brown/bronze  and  the  eyes  are 
located  on  the  sides  of  the  head. 

Similar  species:  Pacific  Chorus  Frogs  {Pseudacris  regilla)  have  obvious  toe  pads  and  an  eye 
stripe  ending  at  the  shoulder.  Recently  metamorphosed  Ranid  frogs  could  be  confused  with 
this  species  but  the  coloration  differs  and  the  tiny  toe  pads  are  lacking  (often  visible  only  with 
a  magnifying  glass  on  small  chorus  frogs). 

Habitat  and  Habits:  Western  Chorus  Frogs  are  regularly  found  in  the  water  only  during  the 
breeding  period  in  spring.  Their  presence  is  obvious  during  this  time  due  to  their  call  which 
is  given  frequently  at  night  and  sporadically  throughout  the  day.  Following  breeding,  these 
frogs  move  into  adjacent  uplands  and  are  rarely  seen.  In  eastern  Montana  they  breed  in 
temporary  ponds  and  small  lakes  surrounded  by  prairie;  in  some  locations  in  Montana  they 
are  also  found  in  open  forested  habitats.  Eggs  hatch  in  about  2  weeks  and  tadpoles  are  about 
2  months  old  at  metamorphosis  (Wlieeler  and  Wheeler  1966,  Nussbaum  et  al.  1983). 

Surveying:  Aduhs  are  easily  sur\'eyed  for,  using  their  calls  for  identification  during  the  breeding 
season  in  the  spring  and  early  summer.  During  the  breeding  season,  adults  may  also  be  seen 
in  the  water,  but  their  small  size  and  habit  of  freezing  or  diving  when  disturbed  makes 
observation  difficult;  night  surveys  may  be  more  productive.  Egg  masses  are  difficult  to  find. 
Tadpoles  may  be  seen  in  ponds  during  the  day  and  can  be  sampled  with  a  dipnet. 

Status:  Common  tliroughout  the  prairies  of  eastern  Montana.    Two  reports  of  Western  Chorus 
Frogs  were  received  for  the  HNF.  One  report  was  from  the  vicinity  of  McDonald  Pass  and 
the  other  from  a  cow  pond  in  the  upper  Jackson  Creek  drainage  of  the  Elkhorn  Mountains 
(Appendix  4);  given  the  distance  from  other  known  locations  and  unusual  habitat,  these 
report  should  be  treated  as  hypothetical  until  verified.  It  seems  possible  that  Western  Chorus 
Frogs  will  eventually  be  verified  from  the  lower  elevations  of  the  HNF.  The  most  likely 
locations  would  be  the  extreme  northeastern  portion  of  the  main  Rocky  Mountains  east  of 
Lincoln,  the  east  side  of  the  Big  Belts,  or  the  Dry  Range. 

Montana  Natural  Heritage  Program  rank:  05  S5. 


20 


Woodhouse's  Toad  {Bitfo  woodhousii) 

Description:  Adults  have  dry  skin  with  small  warts,  and  are  gray,  brown,  or  olive-green  with 
paler  mottling  or  spots.  A  prominent  white  or  yellowish  line  runs  down  the  center  of  the 
back;  very  young  transformed  toads  typically  lack  the  dorsal  line,  and  the  warts  are  often  red- 
brown  in  color.  Woodhouse's  Toad  has  parallel  cranial  crests  between  the  eyes  and  post- 
orbital  crests  connecting  to  them  at  a  right  angle  behind  the  eyes;  the  post-orbital  crests 
typically  touch  the  parotoid  glands.  If  a  lump-like  boss  is  present  on  the  snout,  it  does  not 
extend  back  between  the  eyes.  The  pupils  are  horizontal.  The  adult  has  two  black  tubercles 
on  the  hind  feet  and  a  body  length  of  2.5-4". 
Eggs  and  Tadpoles:  Similar  to  those  of  the  Western  Toad. 

Similar  species:  The  Western  Toad  lacks  cranial  crests.  The  Great  Plains  Toad  has  large,  white- 
bordered,  dark,  dorsal  blotches.  The  Canadian  Toad  has  a  lump  between  the  eyes;  frequently 
the  parotoid  gland  is  separated  from  the  post-orbital  crest  which  may  be  broken  or  absent. 
NOTE:  It  is  very  difficult  to  distinguish  among  the  four  Montana  toad  species  eggs,  larvae, 
and  recently-transformed  toadlets. 

Habitat  and  Habits:  Adults  are  partially  terrestrial  but  often  found  near  water.  They  are  usually 
found  in  irrigated  agricultural  areas  and  flood  plains,  rather  than  the  more  upland  areas  used 
by  Great  Plains  Toads  (Bragg  1940,  Timkin  and  Dunlap  1965,  Black  1970).  They  are  most 
active  at  night,  although  they  may  at  times  be  found  feeding  during  the  day  (Hammerson 
1982a).  They  typically  breed  in  permanent  lakes,  ponds,  reservoirs,  and  slow  streams,  with  a 
preference  for  shallow  areas  with  mud  bottoms  (Black  1970,  Hammerson  1982a,  Baxter  and 
Stone  1985).  Breeding  and  egg  laying  is  spread  out  over  the  spring  and  early  summer,  with 
knowTi  dates  from  Montana  ranging  from  4  May  to  1  July  (Black  1970). 

Surveying:  Adults  may  easily  be  found  by  using  their  loud  calls  for  identification  on  warm  (>54° 
F)  nights;  calling  peaks  during  the  first  few  hours  after  sunset  (Hammerson  1982a).  "Road 
hunting"  on  warm  nights  may  also  be  effective.  Eggs  and  tadpoles  are  seen  in  ponds  during 
the  day  and  can  be  sampled  with  a  dipnet;  however,  identification  of  toad  eggs  and  tadpoles 
is  difficult  or  impossible  in  the  field. 

Status:  Woodhouse's  Toad  is  relatively  common  in  southeastern  Montana,  however,  its  status 
elsewhere  in  the  state  is  unclear.  A  tadpole,  reported  as  Woodhouse's  Toad,  present  in  the 
Montana  State  University  Museum  was  collected  at  the  mouth  of  Trout  Creek  on  the 
Missouri  River  in  1948;  it  should  be  treated  as  hypothetical,  until  verified,  given  the 
difficulty  identifying  Bnfo  tadpoles  and  the  long  distance  to  the  nearest  record.     Geographic 
and  habitat  relationships  with  other  toads  in  Montana  are  not  well  known.  It  should  be 
watched  for  at  low  elevations  in  prairie  or  shrub-steppe  habitat  on  the  HNF;  it  could  occur 
along  the  Missouri  Ri\'er.  Any  located  on  the  HNF  should  be  well  documented  with  a 
description  indicating  how  the  species  was  differentiated. 

Montana  Natural  Heritage  Program  rank:  G5  S4. 


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Plains  Spadefoot  {Scaphiopus  [=Spea]  bombifrons) 

Description:  Adults  are  colored  gray  or  brown  with  darker  mottling  on  the  back  and  a  white 
belly.  Some  individuals  have  indistinct  longitudinal  streaking.  The  pupils  of  the  Plains 
Spadefoot  are  vertically  elliptical  and  there  is  a  high,  hard  lump  between  the  eyes.  Its  skin  is 
less  warty  than  true  toads.  The  adult  has  a  single  tubercle  on  the  hind  feet  and  has  a  body 
length  of  less  than  2.5". 

Eggs  and  Tadpoles:  Oval  egg  masses  of  10-250  eggs  are  attached  to  underwater  plants  or 
debris.  Tadpoles  are  mottled  sooty  and  olive-yellow  above  and  paler  below  with  gold 
metallic  flecking  over  all;  iris  is  gold. 

Similar  species:    Other  Montana  frogs  and  toads  have  round  or  horizontally  elliptical  pupils. 

Habitat  and  Habits:  Adults  are  found  in  grassland  and  sagebrush  areas,  particularly  in  areas  with 
sandy  or  loose  soil  (Wheeler  and  Wheeler  1966,  Hammerson  1982a,  Baxter  and  Stone  1985). 
Except  during  breeding,  they  are  seldom  found  in  the  water.  They  are  primarily  nocturnal 
and  emerge  from  their  burrows  only  following  heavy  rains.  They  breed  in  shallow  temporary 
pools  usually  following  heavy  spring  or  summer  rains  (Hammerson  1982a).  Males  call 
loudly,  with  groups  being  heard  for  up  to  a  mile.  Eggs  hatch  after  2-3  days  and  tadpoles 
transforai  in  6-10  weeks  (Wheeler  and  Wheeler  1966,  Hammerson  1982a). 

Surveying:  Adults  may  be  easily  found  by  using  their  calls  for  identification  when  breeding  at 
night  or  by  "road  hunting"  on  wann,  rainy  nights.  Calling  normally  takes  place  only  when  the 
temperature  is  >50°  F  (Hammerson  1982).  Tadpoles  are  seen  in  ponds  during  the  day  and 
can  be  sampled  with  a  dipnet.  Surveying  is  complicated  by  the  long  time  periods  which  this 
species  spends  underground,  especially  during  droughts. 

Status:  The  Plains  Spadefoot  is  not  known  from  ITNF  lands;  the  nearest  record  is  from  Helena  in 
1988.  Locally  common  in  eastern  Montana;  there  are  large  gaps  in  the  known  range.  It 
should  be  watched  for  at  low  elevations  in  prairie  or  shrub-steppe  habitat  on  the  HNF.  Any 
located  on  the  HNF  should  be  well-documented. 

Montana  Natural  Heritage  Program  rank:  G5  S4? 


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Northern  Leopard  Frog  {Rana pipiens) 

Description:  Adults  are  brown  or  green  with  large,  dark  spots  surrounded  by  light-colored  halos 
on  the  sides  and  back.  The  dorso-latcral  folds  (ridges  along  the  sides  of  the  back)  are  usually 
lighter  in  color  that  the  surrounding  background.  The  under-side  is  typically  white,  but  may 
be  cream-colored  or  yellowish.  The  adult  has  a  body  length  of  2-5".  Newly  transformed 
froglets  may  lack  spots  and  are  about  1"  in  length  (Leonard  et  al.  1993). 
Eggs  and  Tadpoles:  Eggs  are  laid  in  2-5"  globular  masses  composed  of  hundreds  to 
thousands  of  eggs  (Hammerson  1982a,  Nussbaum  et  al.  1983).  The  tadpoles  are  brown  to 
dark  browTi  on  top  with  some  metallic  flecking,  whereas  the  underside  is  often  nearly 
transparent  (Nussbaum  et  al  1983).  Total  length  of  tadpoles  may  reach  more  than  3";  the 
eyes  are  located  on  top  of  the  head. 

Similar  species:    None,  although  some  newly-transformed  froglets  may  lack  spots,  which  makes 
them  extremely  difficult  to  distinguish  from  Spotted  and  Wood  Frogs. 

Habitat  and  Habits:    Northern  Leopard  Frogs  are  found  in  or  near  water  in  non-forested  habitats. 
Vegetation  is  typically  dense,  as  in  a  cattail  marsh  or  dense  sedge-meadow.  Breeding  takes 
place  in  lakes,  ponds  (temporary  and  permanent),  springs,  and  occasionally  backwaters  or 
beaver  ponds  in  streams.  In  Colorado,  eggs  hatch  in  4-15  days  and  tadpoles  take  8-15  weeks 
to  metamorphose,  depending  on  water  temperature  (Hammerson  1982a). 

Surveying:    Both  adults,  tadpoles,  and  eggs  are  easily  seen  in  and  along  the  water  during  the  day 
and  can  be  sampled  with  a  dipnet;  adults  may  also  be  captured  by  hand.  At  very  low 
densities  adults  may  be  difficult  to  find  and  may  be  detected  using  a  call  recorder.  Tadpoles 
are  difficult  to  tell  from  those  of  the  Spotted  Frog  in  areas  where  the  two  species  may 
overlap. 

Status:    Historically,  the  Northern  Leopard  Frog  was  widespread  in  Montana  but  it  now  appears 
to  be  extinct  tliroughout  much  of  the  western  part  of  the  state.  It  is  still  common  and 
widespread  in  the  southeastern  comer  of  the  state,  but  it  may  be  declining  in  central  and 
northeastern  Montana.  It  appears  that  only  localized  populations  are  present  on  the  western 
edge  of  the  plains.  A  single  report  of  this  frog  is  known  from  the  HNF;  an  obsers'ation  of  one 
was  made  near  McDonald  Pass  in  1994.  Given  the  recent  declines  in  this  species  and  the 
unusual  habitat,  this  record  should  be  treated  as  hypothetical  until  verified.  Several  other 
records  exist  from  near  HNF  lands  at  lower  elevations.  The  most  recent  record  was  of  a  few 
individuals  seen  at  the  Deepdale  Fishing  Access  south  of  Townsend  in  1994;  however, 
despite  three  surveys,  none  were  seen  in  1995.  Due  to  its  significant  decline  and  lack  of 
current  reports  from  the  HNF,  all  sightings  of  this  species  should  be  documented. 

Northern  Leopard  Frogs  are  now  absent  from  many  other  areas  in  North  America  where 
they  were  common  a  few  decades  ago.  Widespread  extinctions  are  known  from  Alberta 
(Koonz  1993),  Wyoming  (Koch  and  Peterson  1995),  Colorado  (Hammerson  1982b,  Corn  and 
Fogelman  1984),  Idaho  (Groves  and  Peterson  1992),  Washington,  and  Oregon  (Leonard  et  al. 
1993).  Bullfrog  and  fish  introductions,  acid  rain,  ozone  depletion,  immune  system 
suppression,  and  "Postmetamorphic  Death  Syndrome"  have  all  been  suggested  as  causes  for 
frog  extirpations  in  other  areas  (Com  and  Fogelman  1984,  Hammerson  1982b,  Carey  1993, 
Leonard  e/o/.  1993). 

Montana  Natural  Heritage  Program  ranJ<::  G4  S3S4. 


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Painted  Turtle  {Chrysemys picta) 

Description:  Adult  Painted  Turtles  have  a  relatively  flat  dorsal  shell,  or  carapace,  the  length  of 
which  may  reach  9"  in  females  and  7"  in  males.  The  background  color  of  the  shell  may  be 
dark  brown,  olive,  or  black.  A  series  of  short,  irregular  yellow  lines  are  often  scattered  across 
the  shell,  and  a  red  and  black  border  forms  the  outer  edge.  The  ventral  shell,  or  plastron,  is 
red  with  a  centrally-located  yellow  and  black  blotch  with  edges  flaring  out  along  the  border 
of  the  scutes.  The  edge  of  the  plastron  also  has  a  series  of  black  and  yellow  blotches.  The 
head,  neck,  and  legs  are  marked  with  yellow  lines  and  a  red  spot  appears  behind  the  eye. 
Very  dark  colored  individuals  are  occasionally  found.  Males  are  distinguished  by  longer 
front  claws  and  longer  tails  with  the  anus  posterior  to  the  margin  of  the  carapace  (Ernst  et  al. 
1994). 

Eggs  and  Young:  The  elliptical,  white,  soft-shelled  eggs  are  about  28-35  mm  in  length  and 
16-23  mm  in  width  (Ernst  et  al.  1994).  They  typically  number  6-23  per  clutch.  Coloration  of 
young  Painted  Turtles  is  more  vibrant  and  the  shell  is  not  quite  as  flattened  as  adults. 

Similar  Species:  None. 

Habitat  and  Habits:  Painted  Turtles  are  active  during  the  day  and  are  rarely  seen  far  from  ponds, 
lakes,  or  the  slow-moving  water  of  streams.  Adults  are  primarily  herbivorous,  feeding  on  a 
variety  of  aquatic  plants,  but  will  also  scavenge  on  animal  remains.  Eggs  are  usually  laid 
within  10-20  feet  of  the  water's  edge,  although  some  individuals  will  travel  up  to  600  m 
seeking  a  suitable  site.  During  egg-laying,  the  female  excavates  a  hole  with  her  hind  feet  and 
deposits  the  eggs,  which  are  then  covered  by  several  inches  of  dirt.  Predation  on  turtle  eggs 
by  raccoons,  skunks,  etc.  is  common,  and  shell  fragments  are  evidence  of  such  activity. 
Female  Painted  Turtles  may  lay  more  than  one  clutch  of  eggs  each  summer.  Young  borne  of 
late  egg  depositions  overwinter  in  the  nest  and  do  not  emerge  until  the  following  spring 
(Ernst  et  al.  1994).  Once  females  lay  their  eggs,  they  return  to  the  pond,  where  they  can  often 
be  seen  basking  on  logs  or  rocks  along  with  juveniles  and  males.  Painted  Turtles  are  sexually 
mature  at  3-5  years  of  age  and  may  live  to  be  30  years  or  older  (Ernst  et  al.  1994). 

Surveying:  Although  various  turtle  traps  can  be  used  for  surveys,  visual  identification  is  suitable 
for  presence/absence  studies  since  the  three  turtle  species  in  Montana  are  easily 
distinguished.  Basking  peaks  at  different  times  during  the  day,  depending  on  season  and 
location;  in  the  northern  states  and  Canada  it  generally  peaks  in  the  morning.  Surveys  should 
be  done  on  sunny  days  with  a  pair  of  binoculars.  During  cold  or  cloudy  weather,  turtles  tend 
to  remain  underwater  for  long  periods  and  can  be  missed  on  a  walk-through  survey. 

Status:  Painted  Turtles  are  locally  quite  common  in  Montana  at  lower  elevations.  They  were  not 
found  on  the  HNF.  However  they  are  known  from  lower  elevation  areas  on  the  plains 
adjacent  to  the  forest,  and  were  reported  within  Va  mile  of  the  forest  boundary  by  A.  Harper 
and  S.  Hendrickson  (pers.  comm.)  in  the  western  Elkhom  Mountains.  They  probably  occur 
on  the  HNF  at  low  elevations;  any  sightings  should  be  documented.  There  has  been  some 
concern  about  Painted  Turtle  populations  nationally;  whether  declines  have  occurred  in 
Montana  is  unknown 

Montana  Natural  Heritage  Program  Rank:  G5  S5. 


28 


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Spiny  Softshell  {Trionyrx  spiniferus)  (=Apalone  spinifera) 

Description:  Spiny  Soft-shells  have  flexible,  leathery  shells.  The  carapace  is  olive-gray,  marked 
with  dark  spots.  The  plastron  is  white  or  light  cream-colored.  Female  carapace  length  is  up 
to  18  inches  or  more,  whereas  males  are  typically  6-8  inches.  The  nostrils  are  terminal, 
allowing  this  turtle  to  remain  entirely  beneath  the  surface  and  take  air  through  its  "snorkel." 
Eggs  and  Young:    The  nest  is  a  flask-shaped  excavation  containing  4-39  (typically  12-18) 
hard-shelled,  spherical,  white  eggs.  The  individual  eggs  range  in  size  from  24-32  mm  in 
diameter  and  average  about  28  mm.  Hatchlings  resemble  adults  and  are  30-40  mm  in  shell 
length  (Ernst  e/ a/.  1994). 

Similar  Species:  None. 

Habits  and  Habitat:  Spiny  Softshells  are  active  during  the  day.  This  highly  aquatic  turtle  is 
found  in  rivers  or  their  connecting  backwaters  with  muddy  or  sandy  bottoms.  Unlike  other 
Montana  turtles,  they  do  not  move  overland  from  one  water  body  to  another.  Mud  and  sand 
banks  and  bars  are  used  for  both  basking  and  nesting.  Hibernation  takes  place  beneath  the 
water,  usually  beneath  5-10  cm  of  bottom  substrate  (Ernst  et  al.  1994).  The  retracted  head 
and  neck  combines  with  the  profile  of  the  shell  to  produce  a  wedge  shape,  which  allows  this 
turtle  to  escape  by  literally  diving  into  the  bottom  mud.  If  necessaiy,  additional  strokes  of  the 
legs  will  completely  bury  it  in  the  substrate,  hidden  from  view.  Food  items  include  fish, 
crayfish,  frogs,  toads,  aquatic  insects,  and  carrion.  Spiny  Soft-shells  have  a  surprisingly  long, 
agile  neck  and  can  inflict  a  painful  bite.  They  can  be  safely  handled  by  grasping  the  shell  on 
each  side  between  the  front  and  rear  legs  with  the  head  pointing  away  from  the  captor. 

Surveying:  Although  various  turtle  traps  can  be  used  for  surveys,  visual  identification  is  suitable 
since  the  three  turtle  species  in  Montana  are  easily  distinguished  .  A  pair  of  binoculars  is 
helpful  and  surveys  should  be  done  on  warm  sumiy  days;  basking  seldom  takes  place  before 
10:00  a.m.  (Ernst  e/ a/.  1994).  During  cold  or  cloudy  weather,  turtles  tend  to  remain 
underwater  for  long  periods  and  can  be  missed  on  a  walk-through  surv^ey.  Care  should  be 
taken  to  watch  for  the  snorkel-like  nostrils  projecting  just  above  the  surface  of  the  water. 

Status  and  Distribution:  Found  mainly  in  the  Yellowstone  and  Missouri  Rivers  and  their  major 
tributaries.  These  populations  may  be  separated  from  each  other  and  are  believed  to  be 
disjunct  form  the  population  in  South  Dakota  (Ernst  et  al.  1994);  they  have  not  been  reported 
from  North  Dakota  (Wlieeler  and  Wheeler  1966).  The  Missouri  River  population  is  known 
from  the  tail  of  Fort  Peck  Reservoir  upstream  to  the  first  dam  above,  and  from  most  of  the 
Mussellshell  River;  their  presence  on  other  tributaries  is  presently  unknown.  A  specimen 
was  reported  by  Black  (1970)  from  Canyon  Ferry  Resers'oir,  however,  no  museum  has 
reported  having  one,  nor  have  recent  biologists  in  the  area  reported  them  here;  it  should  be 
treated  as  hypothetical  given  the  long  distance  to  the  nearest  record.  Any  located  on  the  HNF 
should  be  documented  and  reported.  Considered  a  Species  of  Special  Concern  in  Montana. 

Montana  Natural  Heritage  Program  Rank:  G5  S3.  Species  of  Special  Concern. 


30 


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Short-horned  Lizard  {Phrynosoma  donglasi) 

Description:  The  Short-homed  lizard  has  a  broad,  somewhat  flattened  body  and  relatively  short 
limbs  and  tail.  It  is  generally  tan  to  gray  with  dark  and  light  spots  and  blotches;  the  belly  is 
white.  There  is  a  distinctive  line  of  pointed  scales  along  each  side,  and  the  head  has  short, 
blunt  "horns"  pointing  backward.  Adult  lizards  range  from  1.7  -  5.5"  in  length. 
Young:  Young  are  live-bom  and  resemble  small  adults. 

Similar  species:  None. 

Habitat  and  Habits:  The  Short-homed  lizard  is  found  in  a  variety  of  habitats,  including  dry  open 
forests,  grasslands,  and  sagebmsh;  the  soil  is  usually  loose  or  sandy.  In  fimier  soil  situations, 
it  may  use  the  burrows  of  other  animals.  It  is  active  during  the  day,  typically  with  the  peak  of 
activity  in  mid-late  moming.  A  Short-homed  Lizard  may  squirt  blood  from  its  eyes  when 
disturbed.  Little  is  known  about  reproduction  in  this  part  of  the  range;  young  are  bom  in  late 
summer.  Ants  are  the  primary  food  of  the  species. 

Surveying:  They  may  be  surveyed  for  by  slowly  walking  tlirough  appropriate  habitat  and 

watching  carefijlly  for  them;  look  carefully  near  ant  mounds;  this  teclinique  has  low  success 
with  Short-homed  Lizards  however.  As  with  many  lizards  and  snakes,  they  are  easily 
missed.  Carefully  documented  incidental  observations  may  provide  the  best  clues  to  their 
distribution.  They  may  be  also  taken  in  pitfall  or  funnel  traps  in  combination  with  drift 
fences. 

Status:    The  Short-homed  Lizard  subspecies  found  in  Montana  {P.  d.  brevirostra)  is  currently  a 
U.S.  Fish  and  Wildlife  Service  Category  2  Candidate  species  (U.S.  Fish  and  Wildlife  Service 
1994).  It  is  widely  distributed  (but  apparently  localized)  in  eastem  Montana.  There  are  no 
records  from  the  Helena  National  Forest,  but  there  are  records  from  the  Three  Forks  area  to 
the  south  (Reichel  and  Flath  1995).  The  most  likely  places  to  find  this  species  would  be  in 
near  the  south  ends  of  the  Big  Belt  and  Elkhom  Mountains  at  low  elevations.  This  species 
may  be  vulnerable  to  collecting  for  the  pet  trade  and  agricultural  conversion  of  native 
habitats.  It  should  be  watched  for  in  open  pine,  prairie,  or  shrub-steppe  habitat  with  loose  or 
sandy  soils;  all  sightings  should  be  documented. 

Montana  Natural  Heritage  Program  Rank:  G5  S4.  A  Species  of  Special  Concem. 


Rubber  Boa  {Char via  boltae) 

Description:  The  Rubber  Boa  looks  and  feels  like  rubber,  hence  its  name.  It  is  a  small  snake 
(14-33"  length),  stout,  and  uniformly-colored  either  brown  or  green  on  the  dorsal  side.  The 
ventral  surface  is  cream  to  tan  in  color.  The  scales  are  small  and  smooth,  except  for  those  on 
the  head  which  are  enlarged.  The  tail  is  short  and  blunt  and  the  eyes  are  very  small.  It  is  a 
very  slow  moving  snake  which  can  easily  be  caught  if  detected. 

Young:  Rubber  Boas  are  born  alive  and  young  are  more  tan  (or  even  pinkish)  than  the  adults 
on  both  the  dorsal  and  ventral  surfaces. 

Similar  species:  The  Racer  is  much  quicker  and  more  active,  has  larger  eyes,  and  a  thin,  tapered 
(not  blunt)  tail. 

Habitat  and  Habits:  The  Rubber  Boa  is  a  secretive,  slow-moving,  docile  snake,  usually  found 
under  logs  and  rocks  in  either  moist  or  dry  forest  habitats,  but  rarely  in  marsh  or  bog 
situations.  Denning  locations  are  typically  in  areas  with  fractured  rock  on  south  facing 
slopes;  recent  data  indicates  it  rarely  moves  more  than  a  short  distances  from  its  den 
(Peterson  pers.  comm).  Occasionally  this  snake  is  seen  sunning  itself  on  a  road,  trail,  or  open 
area,  but  it  is  primarily  nocturnal.  Feeding  is  primarily  on  small  mice,  but  also  on  slirews, 
salamanders,  snakes  and  lizards.  Two  to  eight  young  are  bom  alive  in  late  summer  or  early 
fall. 

Surveying:  There  are  no  practical  methods  for  surveying  other  than  systematic  searches  of  a 
given  area  rolling  over  rocks,  logs,  etc.  Driving  roads  at  night,  particularly  after  a  rain  when 
the  temperature  is  >  10°  C,  may  be  more  effective,  especially  on  roads  which  follow  a  stream. 
Previous  sightings  are  of  value  in  locating  general  areas  of  activity  and  demiing  sites.  Funnel 
traps  may  be  effective. 

Status:  Sightings  of  Rubber  Boas  are  infrequent,  but  they  are  widely  distributed  and  probably 
common  throughout  western  Montana.  They  were  not  found  during  this  surv-ey,  nor  are  there 
historic  records  of  their  presence  definitely  located  on  the  HNF.  However,  a  record  from 
Grizzly  Gulch  in  1994  may  have  been  on  the  forest.  They  probably  occur  tliroughout  the 
Rocky  Mountains  of  the  HNF  at  low  to  mid-elevations.  They  should  be  watched  for  on  the 
HNF;  any  sightings  should  be  documented.  Of  particular  interest  would  be  any  reports  of  this 
species  from  the  Elkhom  or  Big  Belt  Mountains  or  documentation  of  any  denning  sites 
located. 

Montana  Natural  Heritage  Program  Rank:  G5  S4. 


34 


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Milk  Snake  (Lampropeltis  triangidum) 

Description:  The  Milk  Snake  is  a  slender  and  medium-sized  snake  (to  42  inches  in  length  or 
more),  with  smooth  scales.  It  has  a  highly  recognizable  series  of  red  to  orange  saddles  or 
rings  that  are  bordered  by  black  bands  and  separated  by  white  or  yellow  bands.  Width  of 
dark  and  light  bands  can  \axy  widely.  The  subspecies  in  Montana  {L.  t.  gentilis)  tends  to  be 
paler,  with  orange  bands  replacing  red,  and  a  light  belly  with  few  or  no  black  spots. 

Similar  species:  None  in  Montana. 

Habitat  and  Habits:  Little  is  known  of  Milk  Snakes  in  Montana  because  only  a  few  have  been 
reported.  In  Wyoming  and  elsewhere  they  are  usually  found  near  cliffs,  talus,  outcrops,  and 
rocky  hillsides  in  forested  and  open  country.  They  can  be  found  in  or  under  rotten  logs.  Milk 
Snakes  are  secretive  and  most  active  at  night.  They  eat  a  variety  of  vertebrates,  including 
other  snakes,  lizards,  eggs,  small  mammals,  and  sometimes  invertebrates  such  as  earthworms 
and  insects.  Eggs  are  laid  in  mid-summer.  Milk  Snakes  sometimes  vibrate  their  tails  when 
disturbed.  Their  name  stems  from  an  old  tale  alleging  that  these  snakes  milk  cows. 

Surveying:  Timed-sight  sur\'eys  may  be  conducted  around  cliff  bases  and  outcrops;  turning  over 
rocks  in  these  areas  may  be  very  effective  in  May  and  early  June  (L.  Vitt,  pers.  comm.).  Most 
distributional  information  will  likely  come  from  recording  incidental  sightings..  More 
intensive  research  may  be  done  using  funnel  traps  in  combination  with  drift  fences.  The  most 
intensive  research  and  sur\'ey  projects  may  use  mark-recapture  or  radio-telemetry  techniques. 

Status:  Milk  Snakes  are  very  rare  and  local  in  Montana.  There  are  no  records  from  the  Helena 
National  Forest,  but  there  are  records  from  the  Three  Forks  area  to  the  south  (Reichel  and 
Flath  1995).  The  most  likely  places  to  find  this  species  would  be  in  rocky  canyons  on  the 
south  ends  of  the  Big  Belt  and  Elkhom  Mountains.  The  subspecies  found  in  Montana  is 
highly  sought  for  the  pet  trade.  All  records  of  Milk  Snakes  should  be  fully  documented;  of 
particular  interest  would  be  documentation  of  denning  sites. 

Montana  Natural  Heritage  Program  Rank:  G5  S2.  A  Species  of  Special  Concern. 


36 


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Gopher  Snake  {PiUwphis  catenifer  f^DielanoleucusJ) 

Description:    Montana's  largest  snake,  the  adult  Gopher  Snake  (also  called  Bullsnake  or  Pine 
Snake)  can  reach  a  total  length  of  7  feet,  but  most  specimens  seen  in  western  Montana  range 
between  3-5  feet.  It  is  readily  recognized  by  a  series  of  large  black  to  brown  blotches  which 
run  down  the  back,  and  another  series  along  the  sides.  The  blotches,  which  are  set  on  a 
yellow  background,  become  more  widely  spaced  and  darker  towards  the  tail.  The  dorsal 
scales  are  keeled.  There  is  usually  a  black  band  on  the  head  located  in  front  of  and  extending 
below  the  eyes.  The  ventral  coloration  is  yellow  to  white,  often  spotted  with  black,  and  the 
anal  plate  is  undivided. 

Eggs  and  Young:  Gopher  Snakes  lay  bet\\'een  2-24  eggs  during  the  summer  months 
(Hammerson  1982a),  and  the  young  resemble  the  adults  in  coloration. 

Similar  species:  Young  Racers  have  a  black  border  on  dark  blotches  and  the  scales  are  not 

keeled.  Young  Western  Hognose  Snakes  have  an  upturned  nose.  Western  Rattlesnakes  have 
a  rattle  on  their  tail  and  triangular  shaped  heads. 

Habitat  and  Habits:  Gopher  Snakes  are  associated  with  dry,  arid  habitats  including  grassland, 
slirub-steppe,  and  open  pine  forest.  They  feed  on  rodents,  rabbits  and  ground  dwelling  birds, 
and  to  a  lesser  extent  on  frogs,  toads,  etc.,  found  around  stock  ponds  and  other  wetlands. 
They  have  a  habit  of  hissing  and  vibrating  the  tail  when  alarmed,  often  sounding  like 
rattlesnakes.  They  occasionally  climb  trees,  hence  the  common  name  "Pine  Snake." 

Surveying:  Walk-through  surveys,  done  on  a  regular  basis  in  warm,  sunny  weather  probably 
give  the  best  results  without  resorting  to  trapping  techniques.  They  are  most  easily  found 
near  dens  in  the  spring  and  fall.  Funnel  trapping  is  effective  and  they  may  occasionally  be 
found  by  night  driving  during  the  mid-summer.    Data  can  be  enhanced  by  mark-recapture 
techniques. 

Status:  The  Gopher  Snake  was  not  seen  in  this  survey  nor  are  there  historic  records  from  the 
HNF.  However  they  are  known  from  lower  elevation  areas  on  the  plains  adjacent  to  the 
forest,  and  were  reported  within  V*  mile  of  the  forest  boundary  by  A.  Harper  and  S. 
Hendrickson  (pers.  comm.)  in  the  western  Big  Belt  Mountains.  They  almost  certainly  occur 
on  the  HNF  at  low  to  mid-elevations;  any  sightings  should  be  documented.  Of  particular 
interest  would  be  documentation  of  any  denning  sites  located. 

Montana  Natural  Heritage  Program  Rank:  G5  S5. 


38 


Western  Rattlesnake  {Crotalus  viridis) 

Description:  Rattlesnakes  have  a  heat-sensing  pit  located  between  the  nostril  and  the  eye.  The 
fangs  are  hollow  and  hinged,  allowing  them  to  be  folded  back  against  the  roof  of  the  mouth. 
The  head  is  triangular  in  shape  and  blunt-nosed.  The  eyes  are  slightly  elevated.  There  are 
several  white  lines  which  run  along  the  side  of  the  head.  Adult  Western  Rattlesnakes  have  a 
narrow  neck  but  a  stout  body  with  total  length  ranging  from  15-60  inches.  The  dorsal 
background  color  varies  from  pale  green  to  brown  with  a  series  of  brown  or  black  blotches 
edged  with  a  dark  and  then  light  line  extending  the  length  of  the  body.  The  blotches  often 
merge  into  rings  on  the  tail.  There  are  also  blotches  on  the  sides  of  the  body.  The  ventral 
side  is  pale  yellow  to  white  and  without  blotches.  The  scales  are  keeled.  The  tail  ends  in  a 
rattle  which  helps  to  warn  potential  predators  of  the  snake's  presence.  The  young  have  the 
same  color  pattern,  but  are  brighter  in  color  than  adults. 

Similar  species:  No  other  snake  in  Montana  has  rattles,  but  see  Racer,  Gopher  Snake  and 
Western  Hognose  Snake  which  may  have  similar  color  patterns. 

Habitat  and  Habits:  The  Western  Rattlesnake  is  an  inhabitant  of  more  open  and  arid  country  but 
it  is  also  found  in  Ponderosa  pine  stands  or  mixed  grass-coniferous  forests.  It  is  more  likely 
to  be  encountered  on  south-facing  slopes  and  areas  of  rock  outcrops.  It  is  feared  and  often 
needlessly  killed  due  to  its  poisonous  bite.  Rattlesnakes  may  den  in  large  numbers,  moving 
up  to  7  miles  out  from  the  dens  during  the  summer  (Peterson,  pers.  comm.);  den  sites  are 
most  common  in  south-facing  talus  slopes.    In  Wyoming,  it  is  found  up  to  elevations  of  over 
8500  feet  (Baxter  and  Stone  1985).  Rattlesnakes  prey  on  a  variety  of  animals  including  mice, 
ground  squirrels,  rabbits,  amphibians,  and  other  snakes.  In  Colorado,  females  give  birth  to  4- 
21  young  during  the  summer  (Hammerson  1982a). 

Surveying:  A  walk-through  survey  on  a  warm  sunny  day  is  probably  the  best  method  for 
determining  presence/absence;  it  is  easiest  to  find  near  den  sites  in  spring  and  fall.  Funnel 
traps  and  night  driving  are  both  effective  techniques.    Mark-recapture  methods  can  be  used 
to  determine  more  precise  numbers. 

Status:  The  Western  Rattlesnake  was  not  been  found  in  the  HNF  but  is  known  from  lower 
elevation  areas  to  the  north,  south,  east  and  the  valleys  between  the  mountain  ranges.  It 
would  most  likely  be  encountered  at  lower  elevations  in  open  habitats.  The  habit  of  denning 
at  traditional  sites  in  large  numbers  makes  rattlesnakes  vulnerable  to  commercial  collecting 
or  simply  killing  by  fearfial  people.  Observations  of  Western  Rattlesnakes  should  be  reported 
to  document  the  presence  of  this  species  on  the  HNF;  of  particular  interest  would  be 
documentation  of  any  denning  sites  located. 

Montana  Natural  Heritage  Program  Rank:  G5  S4. 


40 


Regional  Information 

Rocky  Mountain  chain:  The  Tailed  frog  was  only  reported  from  the  main  Rocky  Mountain 
range  of  the  HNF  in  the  Lincoln  District.  It  appears  to  be  more  localized  on  the  HNF  than  in 
areas  to  the  west  and  north.  It  is  not  known  on  the  HNF  to  the  south  of  the  Blackfoot  River. 
Electro-shocking  fish  surveys  should  be  used  to  determine  how  widespread  this  species  is;  all 
incidental  observations  should  be  recorded. 

Other  species  found  during  sur\'eys,  or  for  which  historic  locations  are  known,  on  the  Rocky 
Mountains  of  the  HNF  include  the:  Long-toed  Salamander,  Western  Toad,  Spotted  Frog,  Racer, 
Western  Terrestrial  Garter  Snake,  and  Common  Garter  Snake.  All  of  these  species  are  also 
present  in  the  Elkliom  and/or  the  Big  Belt  Mountains.  The  Western  Toad  was  only  found 
breeding  at  two  locations  on  the  forest;  this  may  be  a  function  of  the  small  amount  of  sampling  or 
the  apparent  decline  in  much  of  western  Montana.  Both  breeding  locations  should  be  considered 
for  monitoring  to  see  if  Western  Toads  continue  to  breed  at  them  and  are  successfully  reaching 
metamorphosis.  Substantial  populations  of  the  Spotted  Frog  were  found  throughout  the  range;  it 
was  more  commonly  encountered  than  any  other  amphibian  or  reptile,  though  ironically  it  is  one 
of  the  two  C-2  Candidate  species  present  on  the  HNF.  A  beaver  pond  area  on  a  tributary  of 
Beaver  Creek  might  be  considered  for  long-term  monitoring  of  Spotted  Frogs  and  Long-toed 
Salamanders;  it  currently  has  very  large  populations  of  both  species.  Concerned  citizens  might  be 
willing  to  participate  in  long-term  surveys.  The  Racer  was  not  found  during  our  surveys;  the 
only  historic  records  for  the  Rocky  Mountains  on  the  HNF  are  from  Davis  (1963),  which  is  a 
large-scale  state  map  with  dots  which  appear  to  fall  on  the  Forest.  The  Western  Ten-estrial 
Garter  Snake  appears  to  be  the  most  common  reptile  on  the  forest.  It  is  perhaps  2-5  times  as 
conmion  as  the  Common  Garter  Snake,  which  may  be  undergoing  a  decline  in  Montana,  Idaho, 
and  elsewhere. 

Given  the  low  numbers  of  locations  for  any  amphibians  or  reptiles  on  the  PINF,  all  sightings 
of  any  species  should  be  recorded.  A  possible  exception  would  be  for  Spotted  Frogs;  however, 
breeding  locations  found  should  be  recorded  even  for  this  species  until  more  are  known  and 
mapped.  Of  particular  interest  would  be  records  of  the  following  species  which  have  not  yet 
been  recorded  on  the  Forest:  Western  Chorus  Frog.  Plains  Spadefoot,  Northern  Leopard  Frog, 
Painted  Turtle,  Rubber  Boa,  Gopher  Snake,  and  Western  Rattlesnake.  All  of  these  potentially 
present  species,  except  the  Rubber  Boa,  are  most  likely  to  be  seen  at  low  elevations  in  open 
habitat.  Additionally,  the  reported  sites  for  Northern  Leopard  Frog  and  Western  Chorus  Frog 
near  McDonald  Pass  should  be  revisited. 

Elkhorn  Mountains:  No  species  were  restricted  to  the  Elkhom  Mountains.  The  Long-toed 
Salamander  is  found  in  the  northv.estem  part  of  the  Elkhoms,  the  farthest  east  reported  location 
in  its  range.  The  Elkliorns  are  the  only  isolated  mountain  range  east  of  the  Continental  Divide 
where  this  species  is  found.  Given  this  unique  situation,  the  distribution  should  be  carefully 
mapped. 

Other  species  found  during  surv'eys,  or  for  which  historic  locations  are  known,  in  the  Elkliom 
Mountains  on  the  HNF  include  the  Spotted  Frog  and  Common  Garter  Snake.  Both  of  these 
species  are  also  present  in  the  niain  R.ock>'  and/or  the  Big  Belt  Mountains.  Substantial 

41 


populations  of  the  Spotted  Frog  were  found  throughout  the  range;  it  was  more  commonly 
encountered  than  any  other  amphibian  or  reptile,  though  ironically  it  is  one  of  the  two  C-2 
Candidate  species  present  on  the  HNF.  The  Corrmion  Garter  Snake  is  known  from  a  single 
location. 

The  Western  Toad  was  collected  along  Prickley  Pear  Creek  in  Jefferson  County  in  1951. 
\Vhether  or  not  this  was  located  on  the  HNF  is  unknown;  it  was  not  located  during  our  surveys. 
Perhaps  the  best  chance  of  finding  the  Milk  Snake  on  the  HNF  occurs  in  the  southeastern 
Elkhorns,  given  a  known  site  to  the  south  near  Tlaree  Forks  and  its  secretive  habits.  It  seem 
doubtful  that  the  Tailed  Frog  is  found  in  the  Elkliom  Mountains  given  the  intensity  of  surveys 
(A.  Harper,  pers.  comm.),  and  the  surveyors  specifically  watching  for  the  species,  during  the 
summer  of  1995. 

Given  the  meager  information  available  from  this  range,  and  the  Western  Toad  (known  to  be 
declining)  recorded  from  historic  records  but  not  relocated,  all  sightings  of  amphibians  and 
reptiles  are  of  interest  from  this  range  and  should  be  recorded.  More  baseline  distribution 
infonnation,  particularly  for  species  not  yet  recorded  and  breeding  sites  for  known  species,  is 
necessary  before  monitoring  sites  are  chosen. 

Big  Belt  Mountains:    The  only  species  reported  from  the  Big  Belt  Mountains  on  the  HNF  were 
the  Spotted  Frog,  Racer,  and  Western  Terrestrial  Garter  Snake.  While  Spotted  Frogs  appeared  to 
be  common  and  widespread  in  this  range,  they  did  appear  more  localized  than  in  either  the 
Elkhorns  or  main  Rocky  Mountains.  The  Racer  was  not  found  during  our  surveys;  the  only 
historic  records  for  the  Rocky  Mountains  on  the  HNF  are  from  Davis  (1963),  which  is  a  large- 
scale  state  map  with  dots  which  appear  to  fall  on  the  Forest.  The  Western  Terrestrial  Garter 
Snake  was  found  at  two  locations  and  is  probably  wide-spread  in  the  range. 

The  Western  Toad  was  collected  historically  just  east  of  the  HNF  in  the  Big  Belts,  but  was 
not  located  during  our  sur\'eys.  Perhaps  the  best  chance  of  finding  the  Spiny  Softshell  or 
Woodhouse's  Toad  on  the  HNF  occurs  in  the  Missouri  River;  both  have  questionable  historic 
records  there.  Given  the  meager  information  available  from  this  range,  and  at  least  one  species 
known  from  historic  records  but  not  relocated,  all  sightings  of  amphibians  (except  perhaps 
Spotted  Frogs)  and  reptiles  are  of  interest  and  should  be  recorded.  Even  for  Spotted  Frogs  any 
breeding  locations  found  should  be  recorded.  Such  baseline  distribution  information  is  necessary 
before  monitoring  sites  are  chosen. 

Dry  Range:  No  herps  are  known  from  this  range;  there  are  no  historic  specimen  records  and  we 
did  not  sur\'ey  in  the  range  in  1995.  Given  the  total  lack  of  information  available  from  this  range, 
all  sightings  of  amphibians  and  reptiles  are  of  interest  and  should  be  recorded.  Baseline 
distribution  information  is  necessary  before  monitoring  sites  are  chosen. 


42 


RECOMMENDATIONS 

Surveys.  Monitoring  and  Research 

1)  All  incidental  sightings  of  amphibians  and  reptiles  from  the  HNF  should  be  recorded  and 
forwarded  to  the  Natural  Heritage  Program,  which  is  the  central  depository  for  amphibian  survey 
and  monitoring  data  from  Montana  in  cooperation  with  the  regional  Declining  Amphibian  Task 
Force.  For  the  Spotted  Frog,  only  breeding  locations  are  necessary  to  record.  A  half-day  training 
session  for  biologists  (including  seasonal  employees)  and  other  interested  field  people  in  May 
would  raise  awareness  of  this  data  need,  and  provide  the  training  required  for  accurate 
identification  of  animals  observed. 

2)  Special  efforts  should  be  made  to  increase  our  knowledge  of  the  range  and  biology  of  those 
species  which  are  either  uncommon  (Tailed  Frog,  Western  Toad)  or  for  which  no  records  exist 
within  the  Forest  (Western  Chorus  Frog,  Woodhouse's  Toad,  Plains  Spadefoot,  Northern 
Leopard  Frog,  Painted  Turtle,  Spiny  Softshell,  Short-homed  Lizard,  Rubber  Boa,  Milk  Snake, 
Gopher  Snake,  and  Prairie  Rattlesnake).  One  of  the  most  efficient  ways  of  accomplishing  these 
objectives  is  to  encourage  individuals  to  fill  out  incidental  sighting  reports  and  submit  them  to 
the  Montana  Natural  Heritage  Program  for  entry  into  the  species  database.  Certainly  all  Tailed 
Frogs  (larvae  and  adults)  found  during  fisheries  surveys  should  be  recorded;  this  is  the  most 
efficient  way  to  get  data  on  this  species. 

3)  Due  to  the  time  constraints  and  the  large  area  covered  in  the  1995  survey,  it  should  not  be 
regarded  as  a  definitive  index  of  all  the  herptiles  or  their  distribution  on  the  HNF.  The  secretive 
habits  of  many  amphibians  and  reptiles,  and  our  lack  of  knowledge  regarding  their  reproductive 
behavior  makes  it  difficult  to  assess  their  overall  status.  We  recommend  that  additional  surveys 
be  conducted,  concentrating  on:  A)  potential  Western  Toad  and  Northern  Leopard  Frog  breeding 
sites;  B)  low-elevation,  xeric  habitats  (including  wetlands  within  this  matrix)  for  reptiles  and 
plains-dwelling  amphibians;  and  C)  gathering  additional  distribution  information  from  the  Big 
Belt,  Elkhom  and  Dry  ranges. 

4)  Begin  a  monitoring  program,  surs'eying  the  two  known  Western  Frog  breeding  sites,  and  the 
beaver  pond  on  a  tributary  of  Beaver  Creek  where  large  numbers  of  Spotted  Frogs  and  Long-toed 
Salamanders  breed.  This  should  be  a  twice-yearly  monitoring,  once  in  late  spring  to  determine  if 
breeding  took  place  and  once  in  mid-late  summer  to  check  if  larva  are  successfully  transforming. 
Monitoring  of  Tailed  Frogs  could  be  done  in  conjunction  with  fish  monitoring  by  electro- 
shocking,  when  sites  are  more  well  kno\Mi.  When  more  breeding  locations  for  amphibians  are 
known  in  the  Elkhorns  and  Big  Belts,  long-term  monitoring  of  typical  marsh-pond  habitats 
should  be  set  up  at  several  sites  in  order  to  evaluate  relative  numbers  and  breeding  success  of  the 
more  common  species:  Long-toed  Salamander,  Spotted  Frog,  Western  Toad,  Western  Terrestrial 
Garter  Snake,  and  Common  Garter  Snake.  Particular  attention  needs  to  be  given  to  any  Western 
Toad  and  Northern  Leopard  Frog  breeding  sites  found. 


43 


5)  Life  history  and  ecology  of  the  amphibians  in  Montana  is  poorly  known  for  most  species. 
Long-term  monitoring  will  provide  information  on  timing  of,  and  habitat  requirements  needed 
for,  successflil  breeding. 

6)  We  recommend  that  any  areas  that  are  under  consideration  for  mining,  road  building, 
extensive  logging  operations,  or  other  large-scale  habitat  altering  activities  be  surveyed 
thoroughly  for  amphibian  presence  and  breeding  activity.  Particular  attention  should  be  given 
documenting  the  presence  of  either  Western  Toads  or  Northern  Leopard  Frogs.    This  is 
particularly  important  for  toads  in  higher  elevation  areas  where  the  potential  for  UV  damage  is 
greater. 

Management 

1)  With  an  increasing  number  of  amphibian  species  declining  for  various  reasons,  it  seems 
reasonable  to  pro-actively  manage  habitat  to  support  them.  While  not  all  ways  of  preserving 
these  species  are  currently  known,  several  management  activities  could  certainly  negatively 
impact  them.  Without  adequate  breeding  areas,  amphibians  cannot  survive,  and  the  types  of 
water  used  is  often  species-specific. 

a)  Fish  stocking  in  currently  fishless  lakes  and  ponds  in  which  amphibians  breed  should  be 
carefully  evaluated.  Fish  introductions  are  thought  to  be  a  major  factor  in  frog  declines  in 
the  Sierra  Nevada  Mountains  and  probably  elsewhere  as  well  (Hayes  and  Jennings  1986). 

b)  When  "improving"  springs  or  seeps  for  livestock,  leave  a  portion  of  the  area  suitable  for 
amphibian  reproduction.  This  could  include  a  small  fenced-off  area  above  where  water  is 
diverted  for  storage  in  a  watering  tank. 

c)  Springs,  seeps,  and  both  permanent  and  temporary  ponds  should  be  considered  when 
analyzing  effects  of  land  management  activities  such  as  grazing,  logging,  and  road 
building. 

2)  A  critical  component  of  the  life  cycle  in  snakes  is  the  wintering  den.  Many  species  hibernate 
in  large  aggregations  in  traditionally-used  sites.  Often  these  hibemacula  are  used  by  several 
species,  and  mating  for  many  species  takes  place  at  the  den  site.  Snakes  then  move  out  for  up  to 
7  miles  for  the  summer,  returning  in  the  fall.  These  sites  are  typically  in  areas  where  snakes  can 
get  well  down  into  an  area  of  fractured  rock  on  south-facing  slopes,  often  near  cliffs  or  in  talus. 
While  these  sites  are  robust,  road  building  or  mining  may  nonetheless  destroy  them.  Den  sites 
should  be  protected  and  data  relating  to  their  locations  kept  where  successive  biologists  have 
access  to  them. 


44 


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U.S.  Fish  and  Wildlife  Service.  1993.  Endangered  and  threatened  wildlife  and  plants:  Finding  on 

petition  to  list  the  spotted  frog.  Fed.  Register  58(87):27260-27263. 
U.S.  Fish  and  Wildlife  Service.  1994.  Endangered  and  Threatened  Wildlife  and  Plants;  Animal 

candidate  review  for  listing  as  Endangered  or  Threatened  species.  Fed.  Reg.  59(219):58982- 

59028. 
Ward,  J.  P.  1978.  TERRAPENE  ORNATA.  Cat.  Amer.  Amph.  and  Rept.  217.1-4. 
Webb,  R.  G.  1962.  North  American  soft-shelled  turtles  (Family  Trionychidae).  Univ.  Kan.  Publ., 

Mus.  Nat.  Hist.  13:429-611. 
Webb,  R.  G.  1973.  TRIONYX  SPINIFERUS.  Cat.  Amer.  Amph.  and  Rept.  140.1-4. 
Werner,  J.  K.  1974.  Phrynosoma  douglassi  brevirostre.  Herpetological  Review  5(1):  20. 
Werner,  J.  K.,  and  T.  Plummer.  1995.  Amphibian  and  reptile  survey  of  the  Flathead  Indian 

Reservation  1993-1994.  Unpubl.  rep.,  Salish  Kootenai  College,  Pablo,  MT.  55  pp. 
Werner,  K.  and  J.  D.  Reichel.  1994.  Amphibian  and  reptile  survey  of  the  Kootenai  National 

Forest:  1994.  Montana  Natural  Heritage  Program.  105  pp. 
Werner,  K.  and  J.  D.  Reichel.  1996.  Amphibian  and  Reptile  Monitoring/Survey  of  the  Kootenai 

National  Forest:  1995  Montana  Natural  Heritage  Program. 
Wheeler,  G.  C.  and  J.  Wheeler.  1966.  The  amphibians  and  reptiles  of  North  Dakota.  University 

of  North  Dakota,  Grand  Forks.  103  pp. 
Wilson,  A.  G.  1985.  PLETHODON  VANDYKEI  IDAHOENSIS  predation.  Herp  Review 

16(4):111. 
Wilson,  A.  G.  and  J.  H.  Larsen  Jr.  1988.  Activit}'  and  diet  in  seepage-dwelling  Coeur  d'Alene 

salamanders  {Plethodon  vandykei  idahoensis).  Northwest  Science  62(5):21 1-217. 


52 


Wilson,  A.  G.,  E.  Simon,  and  J.  H.  Larsen,  Jr.  1989.  Range  extension  for  the  Coeur  d'Alene 

salamander,  PLETHODON  VANDYKEI IDAHOENSIS,  to  the  Canada-United  States  border. 
Canadian  Field-Nat.  103:93-94. 

Wilson,  L.  D.  1978.  COLUBER  CONSTRICTOR.  Cat.  Am.  Amph.  Rep.  218.1-218.4. 

Wishard,  L.  N.  1977.  Larvae  growth  in  RANA  PRETIOSA:  ecological  and  genetic  factors.  M.A. 
thesis.  Univ.  of  Mont. 

Yoffe,  E.  1992.  Silence  of  the  frogs.  New  York  Times  Magazine  13  Dec  1992:36-39,  64-66,  76. 


53 


APPENDIX  1. 

DATA  SHEETS  USED  FOR 

AMPHIBIAN  AND  REPTILE 

SURVEYS  AND  OBSERVATIONS 


54 


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AMPHIBIAN     SURVEY    DATA    SHEET    -  us  BSHI  wildlife  service.  4B  12  McMURRYAVE.Fr.  COLUNS.CO  bob J6-3«00 

(drd«  ch04C«  for  »h*d«d  variabl«a;  supply  v*Jue  lot  oth«n)  (vor.   2/7/92) 


DATE 


BEGIN 
TIME 


END 
TIME 


OBSERVERS 


LOCAUTY 


STATE 


COUNTY 


MAP 

NAME 


OWNER 


ELEVATION 
(circle  tcaJe) 


M 
FT 


SECTION 
DESCRIPTION 


UTM 
ZONE 


NORTHING 
(Of  LAT) 


EASTING 
(M  LOfJl 


AMPHIBIAN  AND/OR  GARTER  SNAKE  SPEOES  PRESENT 
(INDICATE  NUMBERS  IN  CATEGORIES  IF  POSSIBLE) 


CmCL£  METHOD  AND  1^CMCATE  tP 
VOUCHER  SPEOMBJ  WAS  COUJECTED 


SPECIES 


ADULTS/JUVENILES 


CALUNG? 


TADPOLES/LARVAE 


EGG  MASSES 


METHOD: 


VISUAL^AURAL  ID      DIP  NET/SEINE 
HAND  COLLECTED     TRAPPED 
VOUCHER  COLLECTED?       YES      NO 


VISUAUAUFIAL  ID      DIP  NET/SBNE 
HAND  COLLECTED     TRAPPED 
VOUCHER  COLLECTED?       YES      NO 


VISUAL/AUFl&L  ID      DIP  NET /SEINE 
HAND  COLLECTED     TRAPPED 
VOUCHER  COLLECTED?      YES      NO 


VISUAUAURAL  ID      DIP  NET/SEINE 
HAND  COLLECTED     TRA.-PED 
VOUCHER  COLLECTED?       YES      NO 


VISUAUAURAL  ID      DIP  NET/SEINE 
HAND  COLLECTED     TRAPPED 
VOUCHER  COLLECTED?      YES      NO 


FISH  PRESENT? 


YES 


?7? 


FISH 

SPECIES: 


ENTIRE  SITE 
SEARCHED? 


IF  NO.  INDICATE 
AREA 


METERS  OF  SHOREUNE 
KP  Of  HABTTAT 


PHYSICAL  AND  CHEMICAL  ENVIRONMENT  (CHEMISTRY  VARIABLES  OPTIONAL  -  USE  EXTFW.  SPACES  FOR  ADDITIONAL  MEASUREMENTS) 


WEATHER: 


CLEAR 


OVERCAST 


RAIN 


SNOW 


CALM 


UGHT 


STRONG 


AIRTEKff" 

Idrcl*  icalfl) 


WATER  TEMP 

(circle  icttle) 


COLOR; 


CLEAR 


STAINED 


TURBIDITY: 


aJEAH 


CLOUDY 


PH 


ANC 


SITE  DESCFUPTIONS  •  (SKETCH  SITE  AND  PUT  AIXMTIONAL  COMMENTS  ON  BACK  OF  SHETH 
OMfT  THIS  SECTION  IF  DATA  HAVE  BEEN  COLi£CTED  ON  A  PFIEVIOUS  VISIT 


NATUFIAL 


MAN-MADE 


DRAINAGE; 


PERMANENT 


OCCASIONAL 


NONE 


DESCRIPTION; 


PERJAlkNENT 
LAKE/POND 


TEMPORARY 
LAKE/POND 


MARSH /BOG 


STREAM 


SPRING/SEEP 


ACTIVE 
BEAVER  POND 


INACTIVE 
BEAVER  POND 


SITE 
LENGTH  (Ml 


SITE 
WIDTH  (Ml 


MAXIMUM  DEPTH: 


<  1  M 


1  -  2  M 


>  2  M 


STREAM  ORDER 


5  + 


PRIKAARY  SUBSTRATE: 


SILT/MUD 


SAND/GRAVEL 


COBBLE 


BOULDEF.'EEDROCK 


OTHER 


%  OF  POfJD  LAKE  lAARGIN  WITH  EMERGENT  VEGETATIOtJ: 


1  -  25 


25  -50 


>  50 


EMERGENT  VEGETATION  SPECIES 
(UST  l^'  ORDER  OF  ABUNDANCE! 


NORTH  SHOF£UNE  CHARACTERS; 


E:4ALL0WS 
PRESENT 


SHALLOWS 
AESEKT 


E'v?erg£i;t  veg 

PrUlSENT 


i?\GENT  ViG 
/ SScNT 


DISTANCE  (M)  TO  FOREST  EDGE 


FOREST  TREE 
SPECIES; 


ROUGH  SKETCH  OF  SITE 

GRID  SPACING  IS            METERS  BFPA/EEN  LINES 

^ 

^      ^ 

; 

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: 

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:          i 

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\          \ 

ADDITIONAL  NOTES: 


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Natural  Heritage  Rare  Animal  Species  Reporting  Form 

This  form  is  used  to  report  a  personal  field  sighting  of  a  rare  species  tracked  by  the  Montana  Natural  Heritage 
^fcrogram.   It  may  also  be  used  to  summarize  locational  information  from  a  published  or  unpublished  report.   Animal 
^^pecies  tracked  include  those  on  the  U.S.  Fish  and  Wildlife  Service  Threatened,  Endangered  or  Candidate  Lists,  the 
U.S.  Forest  Service  Sensitive  List,  the  Montana  Department  of  Fish,  Wildlife  and  Parks  Species  of  Special  Interest  or 
Concern  List,  and  the  Heritage  Program  Animal  Species  of  Special  Concern  List.   The  Heritage  Program  can  provide 
a  copy  of  the  list  upon  request.   For  most  bird  species,  only  reports  of  confirmed  breeding  are  requested. 

In  order  for  this  form  to  be  processed,  the  sections  preceded  by  two  asterisks  (**)  must  be  completed. 

Send  completed  form  to:  Montana  Natural  Heritage  Program,  1515  E  6th  Ave.,  PO  Box  201800,  Helena  MT  59620. 

Scientific  Name **Common  Name 

Location: 

Location  Map:  A  mapped  location  of  the  occurrence  should  accompany  this  form.  The  ideal  format  is  to  locate  the 
site  on  a  photocopied  section  of  a  USGS  7.5  minute  topo  map;  Forest  Service,  BLM,  or  other  maps  may  be  used.  Be 
sure  to  provide  the  name  of  the  map. 

County: Township: Range: Section: 


•'Directions  to  Site:  Describe  in  detail  how  to  get  to  the  site  from  a  readily  located  permanent  landmark  such  as  a 
road  intersection. 


t 


iology/Habitat 

*Date  and  Approximate  Time  of  the  Observation: 


•*Niimber  of  Individuals  Observed: 

D  1-5         0  5-10          D  11-50          D  51-100           D  101-1000  Q  >  1000 

If  possible,  provide  the  exact  number  of  individuals: 

Life  Stages  Present:  Check  off  the  life  stages  observed  or  provide  an  estimate  of  the  numbers  of  individuals  for 
each  Hfe  stage: 

eggs larvae immature adult  female adult  male adult,  sex  unknown 

Comments: 

Additional  Status  Information:  What  else  was  observed?   Provide  information  on  the  behavior  of  the  species 
particularly  that  which  could  indicate  or  confirm  breeding  at  the  site.   For  birds  this  could  include  singing  males, 
carrying  nest  material/food,  dependant  young  observed,  entry  of  adults  into  possible  nesting  cavity,  etc. 


Associated  Species:  List  any  asr-ociated  species  such  as  predators,  prey,  food  plants,  host  species,  or  additional  rare 
""tecies  observed  at  the  site. 


*=  Required  Fiald 


Habitat  Data:  Describe  the  general  area  where  the  occurrence  is  located.   List  community  types,  dominant 
vegetation,  and  information  on  the  physical  environment  such  as  substrate  type,  hydrology,  moisture  regime,  slope, 
elevation,  and  aspect.   Also,  if  possible,  provide  information  on  the  surrounding  land  use  and  extent  of  additional 
kitable  habitat. 


Weather  Conditions: 

n  clear         D  overcast  D  calm  O  vnndy 

Describe  temperature,  precipitation,  and  other  significant  weather  factors: 


Conservation:  Are  there  any  natural  or  human  threats  to  this  occurrence?  Please  describe. 


Ownership:  If  known,  please  provide  landowner  name,  address  and  telephone  number. 


Jnformation  Source: 

Jame,  Address,  and  Telephone  Number  (of  person  filing  report) 


'♦Does  this  information  come  from  □  a  field  visit,  D  a  2nd  party  observation,  or  □  a  published  or  unpublished  report? 

Citation:  For  information  taken  from  a  published  or  unpublished  report,  please  provide  a  copy  of  the  cover  page  and 
pertinent  portions  of  the  report.   If  a  copy  cannot  be  provided,  list  below  the  author,  date,  title,  publisher,  and  page 
numbers. 


Voucher:  Was  the  observation  vouchered  with  n  a  photograph?  Q  a  specimen? 

If  possible,  attach  a  copy  of  the  photograph.  If  specimen  voucher,  please  provide  the  collection  #  and  name  of  the 

repository: 

Identification:  How  was  the  species  identification  made?  Was  it  based  on  a  sighting,  track,  call,  scat,  road  kill,  etc.? 
Name  the  identification  manual(s)  used  or  expert(s)  consulted.   Were  there  identification  problems? 


Confirmation:  Would  you  accompany  a  biologist  to  the  site  if  needed?   Q  yes    D  no 
additional  Comments:  (use  additional  sheets  if  needed) 


Required  Field 


APPENDIX  2. 

SITES  SURVEYED  DURING  1995 

AMPHIBIAN  AND  REPTILE  SURVEYS 

ON  THE 
HELENA  NATIONAL  FOREST 


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''B  3 


APPENDIX  3. 

AMPHIBIANS  AND  REPTILES 

OBSERVED  DURING  SURVEYS  OF  THE 

HELENA  NATIONAL  FOREST 

IN  1995 


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APPENDIX  4. 

AMPHIBIANS  AND  REPTILES 

REPORTED  FROM  IN  AND  AROUND  THE 

HELENA  NATIONAL  FOREST 


66 


Natural  Heritage  Program  03/21/1996 

Montana  Animal  Atlas  (Herpelile)  Species  Report 

County  Precision    Date         Breed  Data  Type 


LONG-TOED  SALAMANDER 

Jefferson        .5  to  5  mil  4/28/1962  No      Museum  Specimen 
2  mi.  S.  of  East  Helena  on  branch  McClellen  Creek 

Jefferson        .5  to  5  mil  4/28/1962  No     Museum  Specimen 
S.  of  East  Helena  on  Al  Palmer  Ranch 

Jefferson        .5  to  5  mil  4/28/1962  Yes    Museum  Specimen 
2  mi.  s.  of  East  Helena  on  branch  McClellen  Creek 

Jefferson        .5  to  5  mil  8/25/1959  No      Museum  Specimen 
Horse  trough  off  McClellen  Creek 

Jefferson        .5  to  5  mil  7/ /1 972  No      Museum  Specimen 
Near  Clancy 

Jefferson       <  .5  mile.    5/15/1995  Yes    Observation 
Mill  Creek  headwaters 

Jefferson       <  .5  mile.    5/15/1995  Yes    Observation 
Upper  Willard  Creek 

Jefferson       <  .5  mile.    5/20/1995  Yes     Observation 
Old  beaver  pond  in  upper  Corral  Gulch. 

Jefferson       <.5mile.    5/20/1995  Yes     Observation 
Large  pond  just  below  Park  Lake 

Jefferson       <  .5  mile.    9/6/1995  Yes     Observation 
Cliff  Lake,  Red  Rock  Drainage 

Lewis  &  Clark    .5  to  5  mil  7/13/1983  No      Museum  Specimen 
Upper  Grizzly  Gulch 

Lewis  &  Clark   <  .5  mile.    5/20/1995  Yes     Observation 
Old  beaver  pond  on  tributary  of  Beaver  Creek 

Lev/is  &  Clark    <  .5  mile.    5/24/1995  Yes     Observation 
Middle  Copper  Creek  drainage  in  pond. 

Lewis  &  Clark   <  .5  mile.    5/25/1995  Yes     Obser\'ation 
Pond  N.  of  Heart  Lake 

Lewis  &  Clark   <  .5  mile.    7/11/1995  Yes     Museum  Specimen 
Ponds  en  Austin  Creek 


67 


Natural  Heritage  Program  03/21/1996 

Montana  Animal  Atlas  (Herpetile)  Species  Report 

County  Precision    Date        Breed  Data  Type 


LONG-TOED  SALAMANDER    (cominued) 

Lewis  &  Clark   <  .5  mile.    7/11/1995  Yes     Observation 
Beaver  ponds,  Meadow  Creek 

Lewis  &  Clark   <  .5  mile.    7/11/1995  Yes    Observation 
Dog  Creek  beaver  ponds 

Lewis  &  Clark   <  .5  mile.    9/28/1995  No     Observation 
Marysville 

Powell  <  .5  mile.    7/11/1995  Yes     Museum  Specimen 

Lilly  Lake 

Powell  <.5mile.    7/11/1995  Yes     Museum  Specimen 

Mine  shaft  pond 

Powell  <.5mile.    7/11/1995  Yes    Observation 

Bryan  Creek  Beaver  Pond 

Powell  <.5mile.    8/23/1995  Yes     Museum  Specimen 

Small  pond  off  BLM  road,  Garnet  Mountains 

Powell  <.5mile.    8/24/1995  Yes    Observation 

Lower  Chamberlain  Meadows  ca.  3/4  down,  Garnet  Mountains 

Powell  <.5mile.    8/25/1995  Yes     Museum  Specimen 

Old  oxbow  at  Blackfoot  River,  1  mi.  SW  of  Blackfoot  Canyon  Campground 


TAILED  FROG 

Lewis  &  Clark   5  to  10  mil   / /I966  No     Observation 
Tributary  of  Copper  Creek,  tributary  of  the  Blackfoot  River 

Lewis  &  Clark   <  .5  mile.    7/5/1994  Yes     Museum  Specimen 
Falls  Creek,  crossing  down  to  falls. 


WESTERN  TOAD 

Broadwater      >  10  miles.    /  /I966  Yes    Observation 
Near  Canyon  Ferry  Reser\'oir,  small  spring  in  open  ponderosa  pine. 

Cascade         .5  to  5  mil   /  /   0  No     Museum  Specimen 
Little  Belt  r.Icuntains,  1 .2  n-.iles  N.  (Hwy  £9)  of  Kings  Hill  Pass,  7200  "t. 


68 


Natural  Heritage  Program  03/21/1996 

Montana  Animal  Atlas  (Herpetile)  Species  Report 

County  Precision    Date         Breed  Data  Type 


WESTERN  TOAD    (continued) 

Cascade         .5  to  5  mil   / /1 966  No     Observation 
Kings  Hill,  7000  ft. 

Granite  .5  to  5  mil  7/1 1/1977  Yes     Museum  Specimen 

Vi  mi  W  of  Bearmouth,  rest  area  off  1-90. 

Jefferson        .5  to  5  mil  8/7/1951  No      Museum  Specimen 
Prickly  Pear  Creek 

Lewis  &  Clark   <  .5  mile.    7/11/1995  Yes    Museum  Specimen 
Dog  Creek  beaver  ponds 

Lewis  &  Clark    .5  to  5  mil  5/25/1995  No     Observation 
McDonald  Pass  area 

Meagher         >  10  miles.  8/4/1899  No     Museum  Specimen 
Deep  Creek  Canyon,  Big  Belt  Mountains 

Meagher         >  10  miles.  8/23/1919  No     Museum  Specimen 
Fort  Logan,  Camas  Creek  (4  mi  S). 

Meagher  .5  to  5  mil  8/20/1951  No      Museum  Specimen 

Sheep  Creek  near  Jumping  Creek  Campgrounds 

Meagher  .5  to  5  mil  8/24/1951  Yes     Museum  Specimen 

Adams  Ranch,  Sheep  Creek 

Meagher         .5  to  5  mil  8/  6/1958  No     Museum  Specimen 
Lake  Creek 

Meagher         <  .5  mile.    6/26/1995  No      Observation 
Beaver  ponds  on  Daniels  Creek 

Powell  <.5mile.    8/25/1995  Yes    Museum  Specimen 

Old  oxbow  at  Blackfoot  River,  1  mi.  SW  of  Blackfoot  Canyon  Campground 

Powell  .5  to  5  mil  7/9/1995  No      Obser\'ation 

30  mi.  NW  of  Lincoln,  Coopers  Lake 


WOODHOUSE'S  TOAD 

Lewis  &  Clark    .5  to  5  mil  II 9/1948  No     Museum  Specimen 
Mouth  of  Trout  Creek,  Mirscuri  Ri\  >;r 


69 


Natural  Heritage  Program  03/21/1996 

Montana  Animal  Atlas  (Herpelile)  Species  Report 

County  Precision   Date        Breed  Data  Type 


WESTERN  CHORUS  FROG 

Cascade  5  to  10  mil  6/ 7/1993  No      Observation 

Chestnut  Valley  Sand  hills 

Jefferson        <  .5  mile.    8/20/1995  Yes     Observation 
Jefferson  Creek,  Elkhom  Mtns.  In  Cow  Pond. 

Lewis  &  Clark   <  .5  mile.    5/21/1995  No     Observation 
Ca.  17  mi.  S.  of  Augusta,  on  Wrangle  Creek 

Lewis  &  Clark    .5  to  5  mil  5/25/1995  No     Observation 
McDonald  Pass  area 


PLALNS  SPADEFOOT 

Cascade         .5  to  5  mil  8/31/1948  No     Museum  Specimen 
3  mi.  S.  of  Cascade,  Missouri  River 

Lewis  &  Clark   <  .5  mile.    6/ /1988  No     Observation 
Dave  Center's  house. 


NORTHERN  LEOPARD  FROG 

Broadwater      <  .5  mile.    5/10/1993  No     Observation 
Deepdale  FAS.  Directly  E.  across  channel  from  Deepdale  BE  nest 

Broadwater      .5  to  5  mil  8/13/1899  Yes    Museum  Specimen 
Townsend 

Cascade         .5  to  5  mil  8/31/1948  No     Museum  Specimen 
3  mi.  S.  of  Cascade,  Missouri  River 

Cascade         .5  to  5  mil  8/12/1954  No     Museum  Specimen 
Spanish  Coulee 

Granite         .5  to  5  mil  7/27/1961  No     Museum  Specimen 
Lake  Albacaulis 

Lewis  &  Clark    <  .5  mile.    7/ /1993  No     Observation 
very  small  riparian  ponds  along  Dearborn  River 

Lewis  &  Clark    .5  to  5  mil  8/1 1/1948  No     Museum  Specimen 
Wolf  Creek 


70 


Natural  Heritage  Program  03/21/1996 

Montana  Animal  Atlas  (Herpetile)  Species  Report 

County  Precision    Date         Breed  Data  Type 


SPOTTED  FROG    (continued) 

Jefferson        <  .5  mile.    9/6/1995  No      Observation 
Cliff  Lake,  Red  Rock  Drainage 

Judith  Basin     .5  to  5  mil  8/8/1944  No      Museum  Specimen 
12  miles  S.  of  Neihart 

Lewis  &  Clark    .5  to  5  mil    /  /1 966  No      Obser\'ation 
2.3  miles  W.  of  Flasher  Pass  tributary  of  Blackfoot  River 

Lewis  &  Clark   <  .5  mile.    9/3/1994  No     Observation 
Prickley  Pear  Creek  just  W.  of  Stansfield  Lake. 

Lewis  &  Clark    <  .5  mile.    7/27/1975  No      Observation 
Pond,  ca.  400  m  from  the  main  stack  of  East  Helena  Smelter. 

Lewis  &  Clark   <  .5  mile.    7/27/1975  No     Observation 
Ca.  1.6  km  SSE  from  the  East  Helena  Smelter  in  springs 

Lewis  &  Clark    <.5mile.    7/27/1975  No      Obser\'ation 
McClellan  Creek  Quarr>'  Lake 

Lewis  &  Clark    <.5mile.    5/22/1994  No      Museum  Specimen 
7-Up  Pete  Proposed  Gold  Mine  area  on  HWY  200,  ca.  10  mi.  E.  of  Lincoln. 

Lewis  &  Clark    <  .5  mile.    5/20/1995  Yes     Museum  Specimen 
Old  beaver  pond  on  tributary  of  Beaver  Creek 

Lewis  &  Clark   <  .5  mile.    5/20/1995  Yes     Observation 
Pond  0.5  mi.  w.  of  Chessman  Reservoir. 

Lewis  &  Clark    <.5mile.    5/24/1995  No      Obser\'ation 
Copper  Creek  beaver  pond  near  campgrounds. 

Lewis  &  Clark    <  .5  mile.    5/24/1995  No      Observation 
Middle  Copper  Creek  drainage  in  pond. 

Lewis  &  Clark    <  .5  mile.    5/25/1995  Yes     Obser\'ation 
PondN.  of  Heart  Lake 

Lewis  &  Clark    <  .5  mile.    7/11/1995  Yes     Observation 
Ponds  on  Austin  Creek 

Lewis  &  Clark    <  .5  mile.    5/24/1995  No      Observation 
Copper  Creek 


73 


Natural  Heritage  Program  03/21/1996 

Montana  Animal  Atlas  (Herpetile)  Species  Report 

County  Precision    Date         Breed  Data  Type 


SPOTTED  FROG    (continued) 

Lewis  &  Clark    <  .5  mile.    8/31/1995  Yes     Observation 
Poorman  Creek,  before  road  leaves  creek 

Lewis  &  Clark    <  .5  mile.    8/31/1995  No      Observation 
Upper  Blackfoot  River  marshes 

Lewis  &  Clark    <  .5  mile.    7/11/1995  Yes     Obser\'ation 
Beaver  ponds,  Meadow  Creek 

Lewis  &  Clark    <  .5  mile.    7/11/1995  Yes     Observation 
Dog  Creek  beaver  ponds 

Lewis  &  Clark    <  .5  mile.    6/27/1995  No      Obser\'ation 
S.  of  Park  Lake 

Meagher  .5  to  5  mil  6/ 1/1978  No      Museum  Specimen 

Dry  Fork  of  Musselshell  River 

Meagher  <  .5  mile.    5/29/1994  Yes     Museum  Specimen 

Crater  Lake  and  ponds  above,  5880  ft. 

Meagher         <.5mile.    7/8/1994  No     Obser\'ation 
W.  fork  Checkerboard  Creek,  Castle  Mountains,  6200  ft. 

Meagher         <  .5  mile.    7/9/1994  No      Observation 
Onion  Park,  Little  Belt  Mountains,  also  sec.  5. 

Meagher         .5  to  5  mil  8/6/1958  No     Museum  Specimen 
Lake  Creek 

Meagher         .5  to  5  mil  5/13/1950  No     Museum  Specimen 
Near  Ringling  Hot  Well 

Meagher         <  .5  mile.    6/26/1995  No      Observation 
Irrigation  ditch  below  Wolsey  Creek  on  Sheep  Creek 

Meagher         <  .5  mile.    8/3/1995  No      Observation 
N.  Fork  Eagle  Creek 

Meagher         <  .5  mile.    8/15/1995  No     Observation 
E.  Fork  Grasshopper  Creek 

Meagher         <  .5  mile.    8/30/1995  No      Obser\'ation 
Grasshopper  Creek 


74 


Natural  Heritage  Program  03/21/1996 

Montana  Animal  Atlas  (Herpetile)  Species  Report 

County  Precision    Date         Breed  Data  Type 


SPOTTED  FROG    (continued) 

Powell  .5  to  5  mil  7/21/1891  No      Museum  Specimen 

Elliston,  near  Little  Blackfoot  River 

Powell  .5  to  5  mil  7/22/1891  No     Museum  Specimen 

Deer  Lodge,  Cottonwood  Creek 

Powell  .5  to  5  mil    /  /    0  No      Museum  Specimen 

Ovando,  N.  of  Camp  Lake 

Powell  .5  to  5  mil    /  /    0  No      Museum  Specimen 

Browns  Lake 

Powell  <.5mile.    7/11/1995  Yes     Museum  Specimen 

Lilly  Lake 

Powell  <.5mile.    7/11/1995  Yes     Observation 

Beaver  pond,  Little  Blackfoot 

Powell  <  .5  mile.    7/11/1995  Yes     Museum  Specimen 

Mine  shaft  pond 

Powell  <.5mile.    7/11/1995  No     Observation 

Mine  shaft  pond 

Powell  <.5mile.    7/11/1995  Yes     Observation 

Bryan  Creek  Beaver  Pond 

Powell  <.5mile.    7/11/1995  No      Observation 

Little  Blackfoot  River  Beaver  Dam 

Powell  <.5mile.    8/23/1995  Yes     Museum  Specimen 

Small  pond  off  BLM  road,  Garnet  Mountains 

Powell  <.5mile.    8/24/1995  Yes     Museum  Specimen 

Lower  Chamberlain  Meadows  ca.  3/4  down,  Garnet  Mountains 

Powell  <.5mile.    8/31/1995  No     Observation 

Pond  off  road,  Sucker  Creek 

Powell  <  .5  mile.    8/22/1995  No      Observation 

Cottonwood  Meadow,  Garnet  Mountains 

Pov/ell  <.5mile.    8/22/1995  No     Observation 

South  side  Old  Balcv  '■  ''■'••  ■'    Game!  Mountains 


75 


Natural  Heritage  Program  03/2 1  /1 996 

Montana  Animal  Atlas  (Herpetile)  Species  Report 

Count>'  Precision    Date         Breed  Data  Type 


SPOTTED  FROG    (continued) 

Powell  <.5mile.    8/31/1995  No     Obser^'ation 

Pond  off  road,  Sucker  Creek 

Powell  <  .5  mile.    8/25/1995  Yes     Museum  Specimen 

Old  oxbow  at  Blackfoot  River,  1  mi.  SW  of  Blackfoot  Canyon  Campground 


PAINTED  TURTLE 

Broadwater       <  .5  mile.    5/  /1993  No      Observation 
Canyon  Ferry  WMA  east  side. 

Broadwater      5  to  10  mil   /  /   0  No      Specimen  Reported 
See  map  in  Black  1970 

Broadwater       <  .5  mile.    5/9/1995  No      Observation 
Deepdale  fishing  access 

Granite  .5  to  5  mil  6/ /1995  Yes     Observation 

In  pond  by  rest  area  along  1-90  near  Bearmouth. 

Jefferson       <  .5  mile.    7/26/1995  No     Observation 
Prickly  Pear  Road 

Lewis  &  Clark   <.5mile.    / /1994  Yes    Observation 
Spring  Meadow  Lake 

Powell  .5  to  5  mil  9/7/1994  No     Observation 

Lahrity  Lake  in  Ovando  Valley 

Powell  .5  to  5  mil  9/8/1994  No     Observation 

Evans  Lake 

Powell  5  to  10  mil    /  /    0  No      Specimen  Reported 

See  map  in  Black  1970 


SPINY  SOFTSHELL 

Broadwater      5  to  10  mil   /  /   0  No      Specimen  Reported 
Canyon  Ferry  Reservoir 


76 


Natural  Heritage  Program  03/21/1996 

Montana  Animal  Atlas  (Herpetile)  Species  Report 

County  Precision    Date         Breed  Data  Type 


RUBBER  BOA 

Broadwater       5  to  10  mil    //   0  No      Specimen  Reported 
See  map  in  Davis  1963 

Broadwater       .5  to  5  mil  9/  /1957  No      Museum  Specimen 
Near  Toston 

Gallatin        .5  to  5  mil  8/1 1/1959  No     Museum  Specimen 
Middle  Fork  16  Mile  Creek 

Lewis  &  Clark    5  to  10  mil   //   0  No      Specimen  Reported 
See  map  in  Davis  1963 

Lewis  &  Clark    .5  to  5  mil  3/  /1949  No      Museum  Specimen 
Worth  Ranch,  Canyon  Creek 

Lewis  &  Clark    .5  to  5  mil   /  /1 994  No      Observation 
Grizzly  Gulch  SSW  of  Helena 


RACER 

Lewis  &  Clark    5  to  10  mil   //   0  No      Specimen  Reported 
See  map  in  Davis  1963 

Lewis  &  Clark    5  to  10  mil   //   0  No      Specimen  Reported 
See  map  in  Davis  1963 

Lewis  &  Clark    5  to  10  mil   //   0  No      Specimen  Reported 
See  map  in  Davis  1963 

Lewis  &  Clark    5  to  10  mil   //   0  No      Specimen  Reported 
See  map  in  Davis  1 963 

Lewis  &  Clark    5  to  1 0  mil   /  /   0  No      Specimen  Reported 
See  map  in  Davis  1963 

Lewis  &  Clark    <  .5  mile.    5/20/1995  No     Museum  Specimen 
1 .5  mi.  ESE  of  Sieben  Ranch 


Natural  Heritage  Program  03/21/1996 

Montana  Animal  Atlas  (Herpetile)  Species  Report 

County  Precision   Date        Breed  Data  Type 


GOPHER  SNAKE 

Broadwater      <.5mile.    5/21/1994  No     Observation 
Hwy.  12  between  Helena  and  Townsend 

Broadwater      <  .5  mile.    8/30/1995  No     Observation 
Whites  Creek 

Cascade  5  to  10  mil  7/6/1993  No      Observation 

Chestnut  Valley  Sandhills 

Lewis  &  Clark    .5  to  5  mil  10/15/1982  No      Museum  Specimen 
Junction  Sheep  and  Little  Prickly  Pear  Creeks 

Lewis  &  Clark    .5  to  5  mil  5/24/1941  No      Museum  Specimen 
Wolf  Creek 

Lewis  &  Clark    .5  to  5  mil  7/28/1951  No      Museum  Specimen 
Prickly  Pear  Creek 

WESTERN  TERRESTRIAL  GARTER 

Broadwater       .5  to  5  mil  9/ /1 962  No      Museum  Specimen 
W.  of  Winston 

Broadwater       <  .5  mile.    7/12/1995  No      Observation 
Lake  off  of  Springs  Gulch,  Big  Belt  Mountains 

Broadwater       <  .5  mile.    5/29/1995  No      Observation 
Deepdale  Fishing  Access  Site 

Broadwater       <  .5  mile.    7/12/1995  No      Observation 
FS  RD  359  along  Avalanche  Creek  benveen  McGregor  and  Spilling  Gulch 

Broadwater       <  .5  mile.   10/15/1995  No      Museum  Specimen 
Indian  Road  Recreation  Area,  just  N  of  Townsend 

Gallatin         5  to  10  mil    //    0  No      Specimen  Reported 
See  map  in  Davis  1963 

Granite  5  to  10  mil  10/31/1944  No      Museum  Specimen 

Rock  Creek 

Jefferson        .5  to  5  mil  7/30/1951  No      Museum  Specimen 
Prickly  Pear  Creek 


Natural  Heritage  Program  03/21/1996 

Montana  Animal  Atlas  (Herpetiie)  Species  Report 

County  Precision    Date         Breed  Data  Type 


WESTERN  TERRESTRIAL  GARTER    (continued) 

Jefferson        .5  to  5  mil  7/30/1951  No      Museum  Specimen 
Prickley  Pear 

Jefferson       .5  to  5  mil  4/28/1962  No     Museum  Specimen 
S.  of  East  Helena  on  Al  Palmer  Ranch 

Lewis  &  Clark    .5  to  5  mil  7/20/1891  No      Museum  Specimen 
12  miles  E  of  Helena  on  McClellen  Creek 

Lewis  &  Clark    5  to  10  mil    //    0  No      Specimen  Reported 
See  map  in  Davis  1963 

Lewis  &  Clark    <  .5  mile.    7/27/1975  No      Observation 
Pond,  ca.  400  m  from  the  main  stack  of  East  Helena  Smelter. 

Lewis  &  Clark    <  .5  mile.    7/27/1975  No     Observation 
Where  Prickly  Pear  Creek  flows  along  slag  pile  at  East  Helena  Smelter. 

Lewis  &  Clark    <  .5  mile.    7/27/1975  No      Observation 
McClellan  Creek  Quarry  Lake 

Lewis  &  Clark    .5  to  5  mil  6/30/1949  No      Museum  Specimen 
Prickly  Pear  Creek 

Lewis  &  Clark    .5  to  5  mil  9/10/1951  No      Museum  Specimen 
Prickley  Pear  Creek 

Lewis  &  Clark    .5  to  5  mil  9/10/1951  No     Museum  Specimen 
Prickley  Pear  Creek 

Lewis  &  Clark    .5  to  5  mil  8/ /1 994  Yes    Observation 
Little  Prickly  Pear  Creek  N.  of  Helena 

Lewis  &  Clark   <  .5  mile.    7/5/1995  No      Observation 
Helena 

Lewis  &  Clark   <  .5  mile.    8/20/1995  No      Observation 
Falls  Creek 

Lewis  &  Clark    .5  to  5  mil  5/25/1995  No      Observation 
McDonald  Pass  area 

Madison  .5  to  5  mil    /  /1 994  No      Obser\'ation 

Little  Blackfoot  River  W.  of  A\'on 


79 


Natural  Heritage  Program  03/21/1 996 

Montana  Animal  Atlas  (Herpetile)  Species  Report 

County  Precision    Date         Breed  Data  Type 


WESTERN  TERRESTRIAL  GARTER    (continued) 

Meagher  5  to  10  mil  S/22/1919  No      Museum  Specimen 

Fort  Logan,  Camas  Creek,  (4  mi  S) 

Meagher         <  .5  mile.    8/30/1995  No     Observation 
Grasshopper  Creek 

Powell  >  10  miles.  7/31/1967  No     Museum  Specimen 

North  Fork  of  the  Blackfoot  River 

Powell  >  10  miles.  7/3/1973  No      Museum  Specimen 

North  Fork  of  the  Blackfoot  River 

Powell  <  .5  mile.    6/  /1950  No      Specimen  Reported 

Cottonwood  Creek 

Powell  .5  to  5  mil  6/6/1950  No      Museum  Specimen 

0.5  mi.  above  mouth  of  Cottonwood  Creek 

Powell  <.5mile.    8/31/1995  No      Observation 

Road  past  Stemple  Pass 


COMMON  GARTER  SNAKE 

Broadwater      <  .5  mile.    9/25/1995  No     Observation 
Eureka  Creek 

Cascade         .5  to  5  mil  9/6/1994  No     Observation 
Schrammeck  Lake 

Lewis  &  Clark    5  to  10  mil   //   0  No      Specimen  Reported 
See  map  in  Davis  1963 

Lewis  &  Clark    5  to  10  mil    //    0  No      Specimen  Reported 
See  map  in  Davis  1963 

Lewis  &  Clark    5  to  10  mil  7/30/1951  No      Museum  Specimen 
Prickly  Pear  Creek 

Powell  .5  to  5  mil   / /1994  No     Obser\'ation 

N.  of  Ovando 

Powell  <.5mile.    8/3/1995  No     Obsen'ation 

Ontario  Creek 


80 


Natural  Heritage  Program  03/21/1996 

Montana  Animal  Atlas  (Herpetile)  Species  Report 

County  Precision    Date         Breed  DataTjpe 


WESTERN  RATTLESNAKE 

Cascade         .5  to  5  mil   / /1927  No     Museum  Specimen 
Cascade 

Gallatin        5  to  10  mil   /  /   0  No      Specimen  Reported 
See  map  in  Davis  1963 

Lewis  &  Clark    .5  to  5  mil  7/28/1949  No     Museum  Specimen 
Wirth  Ranch 

Lewis  &  Clark    <.5mile.    8/  /1 995  No     Observation 
Melony  Bruhn's  house 


81 


APPENDIX  5. 

NOTES  ON  HARLEQUIN  SURVEYS 

AND  EXAMINATION  OF  POTENTIAL 

NORTHERN  BOG  LEMMING  HABITAT 

ON  THE  HELENA  NATIONAL  FOREST 


82 


Appendix  5.  Notes  on  harlequin  surveys  and  examination  of  potential  northern  bog 
lemming  habitat  on  the  Helena  National  Forest. 

Northern  Bog  Lemmings.  All  sites  surveyed  for  amphibians  and  reptiles  were  also 
examined  for  their  potential  suitability  as  northern  bog  lemming  {Synaptomys  borealis) 
habitat.  None  examined  seemed  suitable.  Particular  attention  was  given  Indian 
Meadows,  however  no  extensive  moss  mats  were  found.  There  may  be  small  pockets  of 
suitable  habitat  within  this  large  wetland  complex  which  were  missed,  as  only  about  4 
hours  were  spent  exploring  the  area,  and  the  water  level  was  very  high.  Summaries  of 
current  knowledge  of  Montana  distribution  and  habitat  is  available  (Reichel  1996, 
Reichel  and  Beckstrom  1994). 

Harlequin  Ducks  were  surveyed  on  the  Landers  Fork  of  the  Blackfoot  River, 
Copper  Creek,  and  the  East  Fork  of  the  North  Fork  Blackfoot  River  (Table  A5.1).  None 
were  found  on  the  Helena  National  Forest,,  however  one  pair  was  located  near  North  Fork 
Falls,  less  than  a  mile  below  HNF  lands  on  the  Lolo  NF.  It  seems  likely  that  ducks  will 
be  found  to  use  the  East  Fork  of  the  North  Fork  on  the  HNF.  It  was  surprising  that  no 
ducks  were  located  on  the  Landers  Fork  given  the  apparently  good  habitat  and  lack  of 
disturbance  on  that  stream.  I  would  recommend  that  1-2  additional  surveys  during  pair 
season  (1  May  to  1  June)  be  done  before  a  final  conclusion  that  ducks  are  not  currently 
present  is  reached.  The  section  above  which  we  surveyed  was  not  accessible;  high  water 
made  crossing  the  river  impossible  and  cliffs  on  both  sides  of  the  river  made  walking  the 
river  also  impossible.  It  is  unknown  at  this  time  if  the  section  above  is  boatable  and  may 
be  surveyed  that  way.  GiN'en  the  easy  access  to  Copper  Creek  and  the  amount  of  the 
stream  surveyed,  it  seems  doubtful  that  this  stream  currently  has  harlequins  present;  I 
would  not  recommend  resurveying  it  at  this  time  without  unless  either:  1)  harlequins 
ducks  are  reported  in  the  Copper  Creek-Landers  Fork  drainage;  or  2)  major  land 
management  activities  are  plamied  in  the  drainage.  Statewide  Harlequin  survey  data  is 
currently  being  summarized  and  will  be  available  soon  (Reichel  and  Center,  in  prep.),  as 
will  a  Conservation  Assessment  and  Strategy  for  the  U.S.  Rocky  Mountains  (Cassirer  et 
al.,  in  review). 


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^Appendix  6.  Heritage  program  species  ranking  definitions. 

Taxa  are  evaluated  and  ranked  by  the  Heritage  Program  on  the  basis  of  their  global  (range-wide)  status,  and 
their  state-wide  status.  These  ranks  are  used  to  determine  protection  and  data  collection  priorities,  and  are 
revised  as  new  information  becomes  available. 

A  scale  of  1  (critically  imperiled)  to  5  (demonstrably  secure)  is  used  for  these  ranks,  and  each  species  is 
assigned  the  appropriate  combination  of  global  and  state  ranks. 

Example:  common  loon  =  G5  /  S3  (i.e.,  species  is  demonstrably  secure  globally;  in  Montana  is  found  within  a 
restricted  range). 

Global  and  state  ranks  are  assigned  according  to  a  standardized  procedure  used  by  all  Natural  Heritage 
Programs,  and  are  defined  below. 

Global/state 

Rank  Definition   (G  =  Range-wide;  S  =  Montana) 

Gl    SI  Critically  imperiled  because  of  extreme  rarity  {5  or  fewer 

occurrences,  or  very  few  remaining  individuals),  or  because  of  some 
factor  of  its  biology  making  it  especially  vulnerable  to  extinction. 

G2    S2  Imperiled  because  of  rarity  (6  to  20  occurrences) ,  or  because  of  other 

I  factors  demonstrably  making  it  very  vulnerable  to  extinction 

throughout  its  range. 

03    S3  Either  very  rare  and  local  throughout  its  range,  or   found  locally 

{even  abundantly  at  some  of  its  locations)  in  a  restricted  range,  or 
vulnerable  to  extinction  throughout  its  range  because  of  other 
factors;  in  the  range  of  21  to  100  occurrences. 

G4    84  Apparently  secure,  though  it  may  be  quite  rare  in  parts  of  its  range, 

especially  at  the  periphery. 

G5    S5  Demonstrably  secure,  though  it  may  be  quite  rare  in  parts  of  its 

range,  especially  at  the  periphery. 

GU    SU         '  Possibly  in  peril,  but  status  uncertain;  more  information  needed. 

GH    SH  Historically  known;  may  be  rediscovered. 

GX    SX  Believed  to  be  extinct;  historical  records  only,  continue  search. 


85 


'  other  codes  : 

A      Accidental  in  the  state;  including  species  (usually  birds  or  butterflies) 

recorded  very  infrequently,  hundreds  or  thousands  of  miles  outside  their  usual 
range . 

B      A  state  rank  modifier  indicating  breeding  status  for  a  migratory  species. 
Example:  S1B,SZN  --  breeding  occurrences  for  the  species  are  ranked  SI 
(critically  imperiled)  in  the  state,  nonbreeding  occurrences  are  not  ranked  in 
the  state. 

E      An  exotic  established  in  the  state;  may  be  native  in  nearby  regions. 

HYB     Element  represents  a  hybrid  of  species. 

N      A  state  rank  modifier  indicating  non-breeding  status  for  a  migratory  species. 
Example:  S1B,SZN  --  breeding  occurrences  for  the  species  are  ranked  SI 
(critically  imperiled)  in  the  state,  nonbreeding  occurrences  are  not  ranked  in 
the  state. 

P       Indicates  the  element  may  potentially  occur  in  the  state. 

Q      Taxonomic  questions  or  problems  involved,  more  information  needed;  appended  to 
the  global  rank. 

'  R      Reported  in  the  state;  but  lacking  documentation  which  would  provide  a  basis  for 
either  accepting  or  rejecting  the  report. 

T      Rank  for  a  subspecific  taxon  (subspecies,  variety,  or  population) ;  appended  to 
the  global  rank  for  the  full  species. 

Z      Ranking  not  applicable. 

#      A  modifier  to  SX  or  SH;  the  species  has  been  reintroduced  but  the  population  is 
not  yet  breeding  and  established. 

USF&WS  (USESA)  Status:  The  codes  in  this  column  denote  the  categories  defined  in  the  U.S. 
Fish  and  Wildlife  Service  Notices  of  Review  (1990,  1993,  1994),  and  indicate  the  status  of 
a  taxcn  with  respect  to  the  federal  Endangered  Species  Act  of  1973: 

E/SA    Treat  as  endangered  because  of  similarity  of  appearance. 

LE      Endangered 

LT      Threatened 

P     Proposed  E  or  T 

CI      Notice  of  Review,  Category  1  (substantial  biological  information  on  file  to 

support  the  Epprcpriaueness  of  prcpcsing  to  list  as  endangered  or  threatened) . 


66 


C2       Notice  of  Review,  Category  2  (current  information  indicates  that  proposing  to 
list  as  endangered  or  threatened  is  possibly  appropriate,  but  substantial 
biological  information  is  not  on  file  to  support  an  immediate  ruling) . 

C2*      Category  2,  and  the  taxon  is  possibly  extinct. 

3A       Taxa  for  which  the  USFWS  has  persuasive  evidence  of  extinction. 

3B      Names  that,  on  the  basis  of  current  taxonomic  understanding,  do  not  represent 
taxa  meeting  the  Endangered  Species  Act's  definition  of  "species." 

30      Taxa  that  have  proven  to  be  more  abundant  or  widespread  than  was  previously 
believed,  and/or  those  that  are  not  subject  to  any  identifiable  threat. 

NL      Not  listed/no  designation.   (See  note  below.) 

XN      Nonessential  experimental  population. 

Note:   A  species  can  have  more  than  one  federal  designation  if  the  species'  status 
varies  within  its  range.   In  these  instances  the  Montana  designation  is  listed  first. 

Examples:       bald  eagle  =  LELT.  Species  is  Listed  Endangered  in  Montana;  elsewhere  in 
its  range  it  may  be  Listed  Threatened. 

trumpeter  swan  =  C2NL.  Species  is  a  Category  2  in  Montana;  elsewhere  in  its  range 
it  may  not  have  USF&WS  designation. 

common  tern  =  NLC2 .  Species  has  no  USF&WS  designation  in  Montana;  elsewhere  in 
its  range  it  may  be  a  Category  2. 

USFS  Status:   The  status  of  species  in  Montana  as  defined  by  the  U.S.  Forest  Service 
Tianual  (2670.22)  .   These  taxa  are  listed  as  such  by  the  Regional  Forester  (Northern 
Region)  on  Montana  National  Forests. 

State  Status:   These  codes  give  the  state  legal  status  of  vertebrates  as  listed  in  the 
1989  Statutes  of  Montana  for  the  Department  of  Fish,  Wildlife  and  Parks. 

GA  =  game  animal 

GF  =  game  fish 

F3  =  fur  bearing  animal  Management  Status: 

MB  =  migratory  bird 

T'ii  =  upland  game  bird  CD  =  closed  season 

E   =  endangered  RH  =  restricted  harvest 

NG  =  nongame  wildlife 

P   =  protected  species 

U   =  unprotected  species 


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