Skip to main content

Full text of "Publications in biological oceanography"

See other formats


Station Lists 

and New Distributional Records 
of Littoral Marine Invertebrates 
of the Canadian Atlantic 

_and New England Regions 


ae al 
i 


Station Lists 

and New Distributional Records 
of Littoral Marine Invertebrates 
of the Canadian Atlantic 

and New England Regions 


National Museum of Natural Sciences 
Publications in Biological 
Oceanography, No. 5 


Published by the 
National Museums of Canada 


Staff editor 
Rigmore Adamson 


Musée national des Sciences naturelles 
Publications d’océanographie 
biologique, n° 5 


Publié par les 
Musées nationaux du Canada 


Station Lists 

and New Distributional Records 
of Littoral Marine Invertebrates 
of the Canadian Atlantic 

and New England Regions 


E.L. Bousfield 
and Diana R. Laubitz 


© Crown copyrights reserved 


Available by mail from the 
National Museums of Canada 
Marketing Services 

Ottawa, Ontario 

K1A OM8 


Catalogue No. NM95- 7/6 


National Museum of Natural Sciences 
National Museums of Canada 

Ottawa, Canada 

1972 


PO987654321 
Y798765432 
Litho in Canada 


Contents 


List of Maps, 6 

List of Tables, 7 
Résumé, 8 

Summary, 9 
Biographical Notes, 10 


Preface, 11 
Acknowledgements, 1 2 


Methodology, 13 
Tables, 13 


General Hydrobiological Characteristics of the Study Regions 
St. Lawrence Estuary and Gaspé Coast, 15 
Southwestern Gulf of St. Lawrence, 16 

Atlantic Coast of Eastern Nova Scotia, 18 

Northern New England and Western Nova Scotia, 20 


Geographical and Ecological Data for the Collections, 23 


Literature Cited, 50 


List of Maps 


Collection stations of littoral marine invertebrates in eastern Can- 
ada and New England regions 

1 Guide to regional station locality maps, 14 

1A St. Lawrence estuary and Gaspé coast, 24 

1B Southwestern Gulf of St. Lawrence, 32 

1C Atlantic coast of eastern Nova Scotia, 40 

1D Northern New England and western Nova Scotia, 44 


Distribution of subarctic and temperate indicator species in eastern 
Canada and New England regions 

2 Gammarus setosus, 16 

3. Mysis gaspensis, 18 

4 Ampithoe longimana, 20 

5  Bittium alternatum, 21 


‘List of Tables 


Guide to Literature 
1 Littoral Marine Invertebrate Surveys, Eastern Canada 
(1950-58), 11 


Geographical and Ecological Data for the Collections 

St. Lawrence Estuary and Gaspé Coast (1953), 25 

St. Lawrence Estuary (1969), 30 

Southwestern Gulf of St. Lawrence (1960), 33 

Atlantic Coast of Eastern Nova Scotia (1962), 39 

Northern New England and Western Nova Scotia (1963), 45 


oof WN 


Resume 


On a préparé des cartes locales et compilé les données accumulées 
dans les stations de recherche. Celle-ci s'est effectuée sur place et 
porte sur la répartition et |’'écologie des invertébrés marins cétiers, 
principalement des mollusques et des crustacés des régions sui- 
vantes: l’estuaire du St-Laurent et la céte gaspésienne, 1953, 
1969; le sud-ouest du golfe St-Laurent, 1960; la cote atlantique 
de |’est de la Nouvelle-Ecosse, 1962: et la céte du golfe du Maine 
qui borde la Nouvelle-Ecosse et la Nouvelle-Angleterre, 1963. Les 
données locales concernent un certain nombre d’espéces d'inver- 
tébrés pour lesquelles on croit avoir établi de nouvelles bornes de 
répartition ou qui sont d’un intérét zoogéographique ou écologi- 
que particulier. 


Summary 


Station data and locality maps are provided for field investigations 
on the distribution and ecology of littoral marine invertebrates, 
mainly molluscs and crustaceans, in the following regions: St. Law- 
rence estuary and Gaspé coast, 1953, 1969; the southwestern 
Gulf of St. Lawrence, 1960; the Atlantic coast of eastern Nova 
Scotia, 1962; and the Gulf of Maine coast of western Nova Scotia 
and New England, 1963. Locality records are provided for a num- 
ber of invertebrate species that are believed to set new distribu- 
tional limits, or are of particular interest, zoogeographically or 
ecologically. 


Biographical Notes 


E.L. Bousfield 

Born in Penticton, B.C., in 1926, Dr. Bousfield graduated from the 
University of Toronto in Zoology (B.A., 1948; M.A., 1949), and re- 
ceived his Ph.D. from Harvard University (Marine Biology, 1954). 
He joined the National Museums of Canada in 1950, as Inverte- 
brate Zoologist, and in 1964 became Chief Zoologist. He is the au- 
thor of over 50 scientific and popular publications, including Cana- 
dian Atlantic Sea Shells (1960), ‘‘Fresh-water Amphipod 
Crustaceans of Glaciated North America’’ (1958), and ‘‘Hausto- 
riidae of New England (Crustacea: Amphipoda)’’ (1965). His scien- 
tific research interests are primarily in the taxonomy, distribution 
and ecology of shallow-water amphipod and cirripede crustaceans; 
estuarine and intertidal ecology; and postglacial dispersal of fresh- 
water and marine invertebrates of Canada. 


Diana R. Laubitz 

Born in London, England, Diana Laubitz graduated from Cam- 
bridge University with a B.A., specializing in Zoology. She came to 
Canada in 1956, and has been associated with the National Mu- 
seum of Natural Sciences since 1964. Her research has been con- 
cerned primarily with the taxonomy and systematics of the Caprel- 
lidea, and she is the author of Studies on the Caprellidae 
(Crustacea, Amphipoda) of the American North Pacific and The 
Caprellidae (Crustacea, Amphipoda) of Atlantic and Arctic Canada, 
in the Museum's Biological Oceanography series. Recently she has 
started investigating some of the Gammaridea closely related to 
this group. 


Preface 


Since 1950 the senior author has conducted systematic field sur- 
veys of littoral marine invertebrate animals, particularly crusta- 
ceans and molluscs, of the Atlantic coast of Canada and the adja- 
cent coast of New England. These surveys have provided material 
for published faunistic studies. Those studies based on surveys 
prior to 1960 have included regional station data and locality 
maps (Table 1). The present contribution provides station data and 
locality maps for regional surveys from 1960 to 1969 (Tables 3- 
6; Maps 1, 1B-1D). Station data for the St. Lawrence estuary and 
Gaspé coast region have been given in part by Tattersall (1954); 
the complete station list and the locality map are included here 
(Table 2; Map 1A). 


Table 1. Littoral Marine Invertebrate Surveys, Eastern Canada (1950-58): Guide 
to Literature. 


Year Region Publication of 
of survey station data, maps 
1950 Coastal New Brunswick and Bousfield 1952 

Nova Scotia; Miramichi Bousfield 1954 

estuary Bousfield 19556, 1955c 
1951 Miramichi estuary Bousfield 19556, 1955c 
1953 St. Lawrence estuary and Tattersall 1954 (part) 


Gaspé coast 


1954 Cape Breton Island and Bousfield 19565 
Newfoundland 

1955 Western Nova Scotia Bousfield 1956c 

1956 - Western Nova Scotia Bousfield 19586 

1958 Minas Basin and Bay of Bousfield and Leim 1960 
Fundy Bousfield 1962a 


Publication of station lists is a useful method of avoiding dupli- 
cation of raw data and map figures where dependent multidisciplin- 
ary publications are anticipated (e.g. Dunbar and Grainger 1952; 
Bousfield and McAllister 1962). The 1960-69 material has al- 
ready provided published records of polychaete worms (Pettibone 
1963a, 19636), haustoriid amphipods (Bousfield 19626), ampelis- 
cid and melitid amphipods (Mills 1963, 1964, 1967), caprellid 
amphipods (McCain 1968), and oyster drills (Medcof and Thomas 
1969). The 1953 material provided the basis for a study on re- 
gional mysid shrimps (Tattersall 1954), a short study on fresh- 
water gastropod molluscs (Bousfield 1955a), and a preliminary re- 
gional hydrobiological analysis (Bousfield 1956a). Along with 
survey material listed in Table 1, the 1953 material was incorpo- 
rated in studies on selected invertebrate groups such as talitrid am- 
phipods (Bousfield 1958c), freshwater amphipods (Bousfield 
1958a) and shallow-water marine molluscs (Bousfield 1960, 


Preface 


1964). The present station data, together with previously pub- 
lished data (Table 1), are incorporated in forthcoming studies on 
postglacial marine invertebrate distribution (Bousfield and Thomas, 
in press), gammaridean amphipods of New England (Bousfield, in 
press), and caprellid amphipods of the northwestern North Atlantic 
(Laubitz 1972). These and other pertinent studies now in prepa- 
ration have lent urgency to the publication of the basic station data 
and primary distributional records in this paper. | 


Acknowledgements 

Many interested persons and research agencies contributed in 
various ways to the field operations during this extended period. 
The authors are especially grateful for facilities, equipment and as- 
sistance provided by officers of the Fisheries Research Board of 
Canada, particularly by Dr. R. E. Drinnan, Mr. Stanley Vass and 
Mr. W. Robichaud during fieldwork in Prince Edward Island, the 
Magdalen Islands and Shippigan regions, respectively; by Dr. H. E. 
Corbeil, Dr. P. Brunel and Mr. J. Bergeron, all of Station de Biologie 
marine, Grande-Riviére, Quebec, during work in the Gaspé region, 
1953, and on the Magdalen Islands, 1960; by Dr. Hans Boerger, 
Mount Desert Island Biological Station, Salisbury Cove, Maine, 
Dr. A. P. Stickney, U.S. National Marine Fisheries Service, Booth- 
bay Harbor, Maine, and Dr. M. R. Carriker, Marine Biological Labo- 
ratory, Woods Hole, Massachusetts, during the New England sur- 
vey, 1963; and by Dr. René Lavoie, Laval University, Dr. Geoffrey 
Power, University of Waterloo, Mr. Donald Galienne, Sept-lles, and 
Mr. Charles H. Douglas, during field operations along the St. Law- 
rence north shore in 1969. Sorting and identification of 1960 bio- 
logical material and compilation of the data were greatly assisted 
by Mrs. Anne Stamper, Lewes, England. Barbara Bousfield, wife of 
the senior author, took part in all surveys except that of 1969, and 
was most helpful in the collection of intertidal molluscs and in field 
photography. To all these persons and agencies and to others un- 
named, the authors extend their sincere thanks. 


Methodology 


Most of the collections and observations were conducted from the 
shore at stations accessible by road. Some freshwater streams and 
lakes and a few terrestrial habitats were sampled. Shore collections 
were made with long-handled dip nets and small sieves, or by 
hand-picking between the tidemarks, and at hip-boot depth. Small 
boats obtained locally were used in sampling the bottoms of la- 
goons and estuarine channels, particularly during the 1960 field 
operations. An Ekman bottom-sampler was used effectively on soft 
mud and sand to depths of over 30 feet (10 metres). A small 
triangular-frame dredge and a 3-foot iron-frame dredge were use- 
fully employed, the latter mainly on stony and shelly bottoms, to 
depths of over 50 feet (16 metres). Oyster tongs and oyster rakes 
were particularly effective on mud and shell bottoms in depths be- 
tween 5 and 10 feet. A set of standard brass sieves, minimum 
screen size 1 mm, was used to separate small organisms from 
muddy and sandy-mud substrata. All materials were fine-sorted 
into systematic groups, re-preserved, and deposited in the research 
collections of the National Museum of Natural Sciences, Ottawa. 

Surface temperatures were recorded with a mercury thermome- 
ter accurate to the nearest 0.1°C, and surface salinities were cal- 
culated from float hydrometer readings of specific gravity accurate 
to the nearest 0.0001 gm/cc. Depths were measured by cali- 
brated line soundings. 


Tables 
‘Maps consulted in the construction of the tables are from three 
main sources: 


National Topographic Series, Map scale 

Surveys and Mapping Branch, 1: 50,000 (1 mile) 
Department of Energy, Mines 1: 200,000 (4 miles) 
and Resources, Ottawa. 1: 400,000 (8 miles) 
Hydrographic Charts, various 


Canadian Hydrographic Service, 
Department of Energy, Mines 
and Resources, Ottawa. 


Army Map Service, 12°°30,000 
United States Geological Survey, 

Department of the Interior, 

Washington, D.C. 


In some cases, the names of the localities listed have been offi- 
cially changed since the maps were issued, and may no longer be 
in familiar use. 


Map 1 Collection stations of littoral marine invertebrates; guide to regional station 
locality maps of eastern Canada and New England 


General Hydrobiological Characteristics of the Study Regions 


A comprehensive and detailed presenta- 
tion of the hydrobiology of the Canadian 
Atlantic and northern New England 
coastal regions is a desirable result of 
completed long-term studies, but is be- 
yond the scope of this initial report and 
station list. However, selected field ob- 
servations on hydrobiological features of 
each of the four major subregions (Maps 
1, 1A-1D) may be usefully included in 
this preliminary report. 


St. Lawrence Estuary and Gaspé 
Coast (Tables 2 and 3; Map 1A) 

With respect to the St. Lawrence es- 
tuary, general hydrobiological features 
have been presented by _ Bousfield 
(1956a), Lavoie (1970) and Bourget 
(1971). The outer Gaspé region and 
Chaleur Bay region have been listed by 
Lacroix (1967), Brunel (1970) and 
Bousfield (1956a). 

These authors tend to subdivide the 
estuary proper “into three discrete re- 
gions. The upper estuary is the tidal 
freshwater portion, stretching for about 
100 miles above the fle d’Orléans; this 
region is warm in summer and supports 
a strictly freshwater fauna. Some spe- 
cies (e.g. Viviparus viviparus L.) extend 
significantly landward into the lower 
part of the broad tidal zone. 

The middle estuary comprises the re- 
gion from the east end of the Tle 
d'Orléans to the mouth of the Saguenay 
estuary on the north side and to Trois 
Pistoles on the south side. Salinity and 
temperature gradients are steep, both 
horizontally and vertically, and show 
large seasonal variation. The tidal ampli- 
tude is high, but water transparency is 
very low, particularly along the south 
shore, where the main seaward thrust of 
silt-laden fresh water is impelled by the 
Coriolis force. The shallow-water fauna 
consists largely of widely eurytopic and 
hardy arctic-boreal or boreal marine spe- 
cies of both plants and animals. The 


middle estuary sets downriver limits to 
freshwater organisms such as crayfish 
(Orconectes), cladocerans (Bosmina), 
limnaeid and physid gastropods, and 
unionid bivalves, the shells of which are 
ice-rafted onto the same beaches as ma- 
rine assemblages, even as far east as 
Bic. 

The lower estuary, from the Saguenay 
to Pointe des Monts and Cap Chat, is 
marked by low summer surface tem- 
peratures and high surface salinities. In- 
tense summer upwelling of very cold 
deep water immediately seaward of the 
Saguenay sill produces very high nutri- 
ent levels at the surface and very high 
primary productivity of benthic algae, 
particularly along the adjacent south 
shore. A strong inflow of fresh water 
from numerous large rivers along the 
north shore appreciably lowers surface 
salinities there in spring and early sum- 
mer. Tidal amplitudes are lower and wa- 
ter transparencies higher than in the 
middle estuary. The shallow-water 
fauna of the lower estuary (and Sague- 
nay fiord) contains a high proportion of 
arctic and subarctic endemics (e.g. 
Gammarellus homari, Gammaracanthus 
loricatus, Atylus carinatus), much higher 
than would be expected at such a low 
latitude (48°-49°N). 

In a study of breeding seasons and 
larval attachment in sessile marine in- 
vertebrates (e.g. Ba/anus), Bourget 
(1971) concluded that primary produc- 
tivity in the middle estuary is very low 
because low salinities and high turbid- 
ities drastically limit the numbers of spe- 
cies of algae and their ability to pho- 
tosynthesize. His temperature and salin- 
ity data show that, in the lower estuary, 
surface temperatures become progres- 
sively lower and salinities become 
higher during summer as the estuarine 
nutrient pump action (upwelling) Is ac- 
celerated by spring runoff and summer 
westerly winds; however, the mecha- 


General Hydrobiological Characteristics of the Study Regions 


Map 2 Distribution of subarctic and temperate indicator species: 


Gammarus setosus 
@ New record 
@ Published record 


nism tends to shut down in winter be- 
cause of the thick ice-cover, reduced 
freshet, and reduced wind effects, and 
the surface water tends to stratify and 
become relatively low in salinity. Bour- 
get (1971) also found that the reproduc- 
tive period of several arctic-boreal and 
boreal species occurs very late within 
the estuary. Spawning periods are com- 
parable to those of low arctic latitudes 
and are much later than in the Gulf of 
St. Lawrence and outer Maritimes 
coastline. 

Coastal waters of the northwestern 
portion of the Gulf of St. Lawrence, 
northwest of Anticosti Island, are some- 
what warmer and more brackish in sum- 
mer than adjacent coastal regions. Mid- 


summer surface temperatures are 14°- 
16°C and salinities 20-28%o, and the 
inshore fauna contains several boreal 
elements, such as Asterias_ vulgaris, 
Idotea balthica, Ampithoe rubricata, 
Pleusymtes glaber, Thais /apillus, and 
Littorina littorea, that are lacking within, 
or penetrate only partly into, the lower 
estuary. 


Southwestern Gulf of St. Lawrence 
(Table 4; Map 1B) 

The southwestern Gulf of St. Lawrence 
is essentially a region of sandy-mud and 
eelgrass lagoons, soft-red sandstone 
bedrock, and beautiful sandy beaches. 
Some estuaries are primarily mud-bot- 
tomed, particularly in the North- 


Southwestern Gulf of St. Lawrence 


umberland Strait region, where tidal 
amplitudes are somewhat higher than 
on the open Gulf coast. Summer surface 
temperatures are everywhere above 
15°C and usually above 18°C, and sa- 
linities are 25-28 %o. Thermal stratifica- 
tion is very pronounced, permitting 
warm-water (but winter-hardy) species 
to occupy a thin surface zone along 
shores and in estuaries, and cold-water 
faunas to exist in summer-cold bottom 
water close inshore. Winter ice is heavy, 
appearing in early December and 
frequently persisting until mid-May, fol- 
lowed by rapid vernal heating of the sur- 
face water layer. Ice scouring contrib- 
utes significantly to the rapid erosion of 
cliff faces and to the low density of ses- 
sile intertidal fauna and flora throughout 
the region (Stephenson and Stephenson 
1954). 

The eastern end of Prince Edward Is- 
land is essentially a summer-cold and 
winter-mild area. Unlike most Island es- 
tuaries, Georgetown Harbour is a deep, 
wide inlet whose open mouth is not cut 
off from the Gulf by barrier bars. Prevail- 
ing summer westerlies blow surface wa- 
ter out to sea and create inshore upwell- 
ing of cold salt water. In winter, the 
upwelling water is comparatively warm, 
and icing is less severe than elsewhere. 
The summer-cold bottom conditions are 
unfavourable to certain species (e.g. Ba- 
lanus improvisus) having warm-water 
pelagic larvae that must be retained 
within the estuary. 

The Magdalen shallows, or ‘‘Magda- 
len pocket’ (of Abbott 1968), contains 
the main northernmost populations of 
the warm-water, or Virginian, fauna that 
includes the ‘‘oyster’’ benthic inverte- 
brate association. These species meet 
their northern limit at Chaleur Bay (e.g. 
B17, B20, Restigouche estuary). The 
oyster association includes the crusta- 
ceans Neopanope texana sayi, Palaemo- 
netes vulgaris, Balanus improvisus, 


Corophium insidiosum, Caprella pe- 
nantis, and Leptochelia rapax; the gas- 
tropods Nassarius obsoletus, Odostomia 
bisuturalis, Mitrella lunata, Retusa cana- 
liculata, Pyramidella fusca, and Crepi- 
dula fornicata; the bivalves Mulinia /ate- 
ralis, Mysella_ planulata, Volsella de- 
missus, Petricola  pholadiformis, and 
Mercenaria mercenaria; and representa- 
tives of other invertebrate groups, such 
as Molgula manhattensis (Tunicata), 
Polydora websteri and Eteone heterpoda 
(Polychaeta), and Microciona prolifera 
(Porifera). 

Particularly noteworthy records for 
warm-water crustaceans in the south- 
western Gulf region are: Melita nitida 
(S27, S29); Ampithoe longimana - l|a- 
goons of the outer coast of P.E.|. (e.g. 
P2, P42), Magdalen Islands (M5, M6), 
George Bay (S7), and Buctouche (B5); 
Ovalipes ocellatus (B23, B24); and 
Rhithropanopeus harrisi (S23, S27, P8). 
Extending or confirming the molluscan 
ranges summarized by Bousfield (1960) 
are the following: the gastropods /ri- 
phora .nigrocincta (S25, $35, P43; 
B27); Cerithiopsis greeni (S25, $35, 
P12); Bittium alternatum - New Bruns- 
wick coast north to B5 and sporadically 
to B17, east in Nova Scotia to George 
Bay, widely around P.E.I., but not on the 
Magdalens (see also Map 5); Odostomia 
seminuda (S34, P5); Haminoea_ soli- 
taria- mainland north to B1/, widely 
around P.E.I., but not on the Magda- 
lens; the nudibranch Elysia chlorotica 
(S31, B22, B31); the bivalves Cum- 
mingia_ tellinoides - mainland north to 
B8, not on the Magdalens; Pandora 
gouldiana (M4, B20); and a large beach 
deposit of fossil oyster shells at M8, pre- 
viously noted by Medcof, Clarke and Er- 
skine (1965). The boreal species Zir- 
phaea crispata is common along peaty 
margins of the northeastern New Bruns- 
wick coast. Specimens of Saccog/ossus 
kowalewskii (Agassiz) were confirmed 


General Hydrobiological Characteristics of the Study Regions 


Map 3 Distribution of subarctic and temperate indicator species: 
Mysis gaspensis 

@ New record 

@ Published record 


from station P34 by Dr. N. Burdon- 
Jones (personal communication). 
Unexpected records of live shallow- 
water populations of cold-water or sub- 
arctic species include: Mysis gaspensis 
(S11, S16, P19, M4-M6, B11), Ba- 
lanus balanus (P37, M10, B24), Meso- 
desma arctatum (P26, M1, M10, B15), 
Arctica islandica (P25,M11, M12), Vol- 
sella modiolus (P37, M11), Margarites 
helicinus (P37), and Buccinum undatum 
(P25, P27,M12, B16). Apparently, very 
sharp vertical and seasonal thermo- 
clines permit both warm-water and cold- 
water faunas to exist virtually side by 
side in this region. The Magdalen Is- 
lands did not yield living specimens of 
Crassostrea virginica, Mercenaria mer- 


cenaria, Volsella demissa, Nassarius ob- 
soletus, cerithiid snails, Neopanope tex- 
ana sayi, Palaemontes vulgarus, or 
Balanus improvisus. The absence from 
the Magdalen Islands of these abundant 
and dominant indicators of Virginian 
summer conditions suggests that the 
name ‘‘Magdalen pocket’’ may not be 
entirely applicable or suitable. 


Atlantic Coast of Eastern Nova Scotia 
(Table 5; Map 1C) 

The outer coast of eastern Nova Scotia, 
particularly from Clam Harbour to In- 
dian Harbour, is essentially a cold-water 
region. Here are found the lowest sum- 
mer surface temperatures and the great- 
est seasonal range of air temperatures. 


Atlantic Coast of Eastern Nova Scotia 


The rigorous continental climate ac- 
counts mainly for the low species diver- 
sity, the low incidence of winter thermo- 
philes, and the rarity of warm-water 
endemics. The influence of the substra- 
tum on faunal composition is marked. 
The regional bedrock is igneous and acid- 
ic, forming a sharp escarpment (a more 
than 50-foot drop) a few miles inland 
from the outer coast. The estuaries are 
thus short and lack extensive sandy- 
mud tidal flats. Cold water penetrates 
up the channels, upwelling is induced 
by prevailing offshore weather systems, 
and the surface flow of fresh water into 
the estuary is low in nutrients and pH. 
All these factors combine to produce 
conditions unsuitable for warm-water 
estuarine endemics, particularly those 
with pelagic larvae. Only Oyster Pond 
and Ostrea Lake today offer suitable, if 
tenuous, physical conditions for comple- 
tion of the life cycles of such organisms 
(see also Medcof et al. 1965). 

The axis of an estuary may have an 
important bearing on its faunal compo- 
sition. If the axis is elongate and at right 
angles to the ocean front, surf and cold 
water penetrate well into the bay in 
summer; the fauna is composed largely 
of cold-water or boreal eurytherms and 
lacks endemics. Along somewhat more 
mature coastlines, estuaries tend to 
broaden at the mouth and are partly 
protected by barrier bars and shallow 
sills. With moderate protection from 
surf, inner waters stratify in summer, 
and a few warm-water species may be 
found there. Along mature coastlines, la- 
goons with sandy barrier bars develop, 
the axis of the estuary becomes more or 
less parallel to the coast, and inner wa- 
ters are completely protected from surf 
and cold-water invasion. Summer strati- 
fication and a warm-water plankton- 
retention mechanism characterize the 
circulation. Most of the estuaries of east- 
ern Nova Scotia can today be included 


among the first two categories de- 
scribed, whereas very few (e.g. Cole 
Harbour) fit into the third category or 
close to it. In hypsithermal times 
(c. 7OOO bp), however, lowered sea lev- 
els exposed extensive offshore sandy ar- 
chipelagos that had probably developed 
mature estuarine profiles, and thereby 
provided a sort of stepping-stone north- 
ward passageway for warm-water es- 
tuarine endemics into the Gulf of St. 
Lawrence (see also Bousfield and 
Thomas, in press). 

The warm-water or ‘‘oyster’’ fauna 
occurs sporadically along the eastern 
Nova Scotia coast, although surprisingly 
widespread are Corophium insidosum 
(absent only from A36-A42 and A54- 
A60), Bittium alternatum (A4, A6, A17, 
A22, A23, A27, A31, A32, A34, A47, 
A66, A70), and Jel/lina agilis (including 
A48, A53, A58, A59, A61, A66, A68, 
A69). Other notable records among the 
crustaceans are Haustorius canadensis 
(A17), Caprella penantis (A16, A73), 
Leptochelia rapax (A5, A22, A34, A37, 
A61, A70O, A79), and Mysis stenolepis 
(A4,A16, A22,A61, A63, A66, A69). 

An apparent lack of Ba/anus impro- 
visus, Ampithoe longimana, and warm- 
water decapods probably reflects the 
unsuitability of present-day summer 
temperatures and estuarine circulation 
for prolonged pelagic larval develop- 
ment and retention. Most warm-water 
molluscs occur in pockets west of Sheet 
Harbour or in the Chedabucto Bay and 
Cape Breton regions, with occasional 
specimens at intervening localities (ice- 
rafted shells), e.g.: the gastropods Mit- 
rella lunatia (A6, A17, A22, A32), Re- 
tusa canaliculata (A25, A31, A32, A3A4), 
Crepidula fornicata (A22, A34, A43), 
Nassarius obsoletus (A6, A17, A22, 
A27, A32, A70, A71, A79), and Hami- 
noea solitaria (A27); the bivalves Cras- 
sostrea virginica (A1, A6), Mysella plan- 
ulata (A17, A19), Volsella demissa (A1, 


General Hydrobiological Characteristics of the Study Regions 


Map 4 Distribution of subarctic and temperate indicator species: 
Ampithoe longimana 


A5, A6, empty valves at A52), Pandora 
gouldiana (A22, A33), Petricola pholadi- 
formis (A6, A22, empty valves at A66), 
and Pitar morrhuana (A19, A27, A35, 
A43). Winter-mild thermophiles occur 
eastward along the coast as follows: 
Amphiporeia virginiana (to A5O), Marino- 
gammarus finmarchicus (to A29), Bathy- 
poreia quoddyensis (A10, A12, A73), 
Chiridothea caeca (to A5O), and Orches- 
tia grillus (to A6O). Orchestia gamma- 
rella was taken at A22, and the pipefish 
Syngnathus fuscus at A71. 

Cold-water and subarctic species are 
not uncommon close inshore, especially 
in the region east of Sheet Harbour to 
Cape Canso (A43-A60). These include 
the crustaceans Mysis gaspensis (A31, 
A34, A35, A60, A64, A65), Balanus ba- 
lanus (A73), Mancocuma stellifera (AQ, 


20 


A17, A18, A21, A30, A31, A42, A73), 
Pontoporeia femorata (A27, A35), and 
Pagurus pubescens (A44); the gastro- 
pods Buccinum undatum (A22, A42), 
Skenea planorbis (A51, A66, A74, A75, 
A76), and Margarites helicinus (A22, 
A35, A42, A44, A54, A58, A63, A76); 
and the bivalves Mesodesma arctatum 
(AQ, A12, A17, A19, A64, A73), and 
Volsella modiolus (A8, A33, A35, A39Q, 
A42, A60O, A63). 


Northern New England and Western 
Nova Scotia (Table 6; Map 1D) 

The New England coast north of Cape 
Cod is a boreal or cold-temperate region 
that may be divided into two zoogeo- 
graphical subregions. On the basis of 
surface-water characteristics and faunal 
composition, we may recognize (1) the 


Northern New England and Western Nova Scotia 


Map 5 Distribution of subarctic and temperate indicator species: 


Bittium alternatum 


section north from Penobscot Bay by its 
uniformly cold (less than 12°C) summer 
surface temperatures and relatively high 
salinities, in which boreal and subarctic 
faunal elements are dominant; and (2) 
the section south of Penobscot Bay con- 
taining pockets of warm brackish waters 
in which Virginian faunal elements are 
dominant. 

The Cape Cod region has long been 
recognized as a practical northern limit 
of the main populations of the Virginian 
fauna and a southern limit of the boreal 
fauna. In actuality, however, many of 
the warm-water invertebrates extend 
northward into northern New England 
and cold-water invertebrates to Long Is- 
land Sound and southward. Notable 
northern New England records among 
the warm-water arthropods are Limulus 


polyphemus (M32, M33), Balanus im- 
provisus (M33, M35), Pagurus long- 
icarpus (M31, M32), Rhithropanopeus 
harrisi (M58), Exosphaeroma orego- 
nensis (M75), Heteromysis formosa 
(M73), Melita nitida (M35, M39), Am- 
pithoe valida (M35), Corophium ache- 
rusicum (M33), Gammarus palustris 
(M58), and Orchestia uhleri (M52, 
M53). Extending northward into west- 
ern Nova Scotia are Palaemonetes pugio 
(S17), Mysis stenolepis (S3, S11), 
Praunus’ flexuosus (SQ), Carcinides 
maenas (S11, S2, $17, S19), Libime 
emarginata (S1), Corophium insidiosum 
(S9, S17), and C. /acustre (head of St. 
Marys Bay). 

Records of winter-mild thermophiles 
include Orchestia gammarella (S4, M7), 
Ligia oceanica (S4, S6,S16, M56), and 


21 


General Hydrobiological Characteristics of the Study Regions 


Littorophiloscia vittata (S11, S13, S17, 
S20, M34) (see also Lemos de Castro 
1965). 

Noteworthy records of warm-water 
molluscs include the gastropods Poli- 
nices duplicata (M71, M73), Urosalpinx 
cinerea (S9, M32, M33), Bittium al- 
ternatum (M77), Haminoea solitaria 
(M19, M20), Nassarius obsoletus (S1, 
$17,M1,M20), Crepidula fornicata (S1, 
S9,M19, M20, shell at M6), Mitrella /u- 
natia (S1, S9, M32, M33), and Ovatella 
myosotis (S11, M33); the bivalves Cras- 
sostrea virginica (M39, M57), Merce- 
naria mercenaria (S1, S8, M19, M32, 
M33), Volsella demissa (S8, S17, M20, 
M32-M34), Petricola pholadiformis 
(S9, M20, M32, M33), Pandora goul- 
diana (S1, S8, S9), Mysella planulata 
(S9), and Tellina agilis (S1, M1, M30); 
and the squid Lo/igo pealii (S1). 

In the lower part of the tidal zone, 
along Maine and Scotian coasts border- 
ing the entrance to the Bay of Fundy, 
are found a number of cold-stenother- 
mal invertebrates that occur subtidally 
elsewhere: Cancer borealis, Balanus ba- 
lanus, Melita dentata, Pontoporeia femo- 
rata, Anonyx sarsi, Volsella modiolus, 
Arctica islandica, Astarte undata, Meso- 
desma arctatum, Buccinum undatum, 
and Margarites helicinus. Especially 
noteworthy records of cold-water spe- 
cies include Mysis gaspensis (M10, 
M11), Mancocuma stellifera (M9), Gam- 
marus setosus (S12, M5), and southern 
interdidal populations of Cancer borealis 
and Buccinum undatum (M72). 


22 


Geographical and Ecological Data 
for the Collections 


Geographical and Ecological Data 


GULF 


OF St. LAWRENCE 


G32 
ca O28 G25 GH G33 G34 
24 G39 
622 i ie cart 
Gi6 = G20 G54 G52 G42 
G55. at G44 
G58 oS 
G60" 659 ous 7 
G61 
Saguena d 
9 y C16 «C18 C20 )'G62 
L252 [ss $25 C14 
S224 $26 C15 C17 24 
$21 C2 cu C13 
a ct C3 co C12 
N14 Me c8_C10 
48° ae c7 
N12 FE ae ~ du 
S1 S 
: CHALEUR BAY r cs 


Map 1A Collection stations in the St. Lawrence estuary and Gaspé coast regions 


24 


St. Lawrence Estuary and Gaspé Coast 


Table 2. St. Lawrence Estuary and Gaspé Coast (1953) 
Key to station localities on Map 1A: A - Anticosti Island; C - Chaleur Bay; G - Gaspé (Ste-Flavie to Percé); 
S - south shore, St. Lawrence estuary; N - north shore, St. Lawrence estuary 


Sta Date Locality Lat N Long W Depth Temp Salinity Habitat 
No. val %o 
Ail Aug.9_ Port-Menier, SE of 49°48’ 64°21' LWtoHW 13.8 31.2 Tidal flats, limestone, 
wharf HW drift 
A2 Aug.12 Port-Menier, pier 49°49’ 64°22' HW _ -- Gravel beach 
A3  Aug.11 Baie Ste-Claire 49°53.5' 64°31' LWtoHW 78 30:6 Pebble and sand beach, 
tidal flats 
House ruins Terrestrial _ — Under stones 
Woodlands Terrestrial — — Under logs 
Small lake Terrestrial = — Under boards in and 
around dried lake 
A4 Aug.10 2 miles NE of Cap 49°56’ 64°05’ LWoHW Tho 305 Pebbles and limestone 
de Rabast flats, igneous boulders 
Small stream 20.0 Brackish 
to fresh 
C1 Aug.7 Maria, old wharf 48°10’ 66°00' LWtoHW 13.1 15.9 Stones at base of 
pilings, pebble beach 
C2 Aug.7 Cascapédia estuary 48°15.5' 65°54’ LWtoHW 16.5 18.4 Mud flats 
C3 Aug.6_ R. Petit Cascapédia, 48°10’ 65°50.5’ LW 14.9 1.4 River and tidal flats 
at New Richmond 
C4 Aug.7_ R. Petit Cascapédia 48°10.5’' 64°46’ 1 ft 16.1 Fresh Stones 
C7 Aug.6- Paspébiac, oldwharf 48°01’ 65°14’ HW — _ Sand beach 
C8 Aug.6 _ St-Godefroi, west 48°04’ 65°C?" “HW = — Under logs, fine-sand 
beach 
C9 = July 31. ~Port-Daniel, river 48°11’ 64°58’ MW 18.8 Brackish Mud flats, eelgrass 
mouti) 
Head of estuary 48°12’ 64°58’ MW. — Brackish Mud flats, salt marsh 
C10 July 31 Port-Daniel, east 4S° 11" 64°57' HW NSB -2H29 Fine-sand beach 
C11 Aug.6 Gascons 48°12' 64°52’ HW a -- Under logs, fine-pebble 
beach 
C12 July 31 Newport, beach 48°15’ 64°45’ HW _ — Fine-sand beach 
opposite Gull |. 
C13 Aug.6 Newport light AB? 17! 64°43' HW — —_ Coarse-sand beach 
C14 Aug.3 Chandler,headofbay 48°20’ 64°44’ LWtoHW 204 19.2 Marshy beach, mud 
at R. du Grand Pabos flats 
C15 Aug.2 Chandler, mouth of 48°20’ 64°42’ LWtoHW 15.2 25.1 Coarse sand, rocks 
bay 
C16 Aug.3 _ Lacdes Sept Iles 48°21’ 64°48’ <3ft 18.8 FW Stones 
R. des Sept-lles, AS 2ay “eArAy" FW 
Y2 mile below lake 
C17 Aug.3  Ste-Adelaide-de-Pabos 48°21’ 64°37’ HW _ _ Sand and pebble beach 
C18 Aug.6 _ Petit Pabos estuary 48°22' 64°35’ HW - _ Sand beach 
C19 July 30 Bog Pond, 4 miles 48°25' 64°32’ <5ft 22.8 _ Bog margin, woody 
NW of Grande-Riviére detritus, grass roots 
C20 July 30 Grande R., 1 mile 48°24! 64°21’ 1 ft 17.8 Fresh Stones 
above mouth 
C24 Aug.5 Anse-du-Cap-d’Espoir 48°25’ 64°19’ HW — — Sand beach 
G1 July9 1 mile W of Ste-Flavie 48°36’ 68°15.5' LWtoMW 14.5 27.5 Boulders 
G2 Aug.17 R.Mitis 48°37.5' 68°08’ 1-2 ft 17.6 Fresh Stones, rapids 
G3  Aug.18 Baie du Petit Mitis 48°41' 68°02’ LYN ton OlisrSn 827-7 Sandstone, slate and 


mud flats 


25 


Geographical and Ecological Data 


Sta Date Locality Lat N Long W Depth Temp Salinity Habitat 
No. rE %o 
G4 Aug.18 R. Tartigou 48°45’ 67°47' 1 ft 16.8 Fresh Stones 
G6 Aug.26 Pointe au Naufrage 48°46’ 67°47' LWtoHW 10.0 27.5 Coarse sand at HW, 
shale and boulders 
G9 Aug. 24 Matane, W of wharf 48°51’ 67°32' LWto HW 9.7. «26.9 Sand with stones 
G11 Aug.22 R. Petite Matane 48°49’ 6727.0 ott 16.0 Fresh Pebbles 
G12 Aug.22 Smallstream 5miles 48°51’ Gr? 2 7.55 ot it 11.2 Fresh 
E of Petite-Matane 
G13 Aug.22 Ste-Félicité, W of 48°54! (7 2A5/e LW to HW 98 28.8 Rocks and boulders 
wharf 
G15 Aug.20 Ruisseau a la Loutre 48°56’ 67°09' MW toHW — Brackish Boulders at MW, wood- 
lands at HW 
Cascades at mouth 13.1 “= 
G16 Aug.23 La Lorraine Cove 48°57.5' 67°07.5' LWto HW 8:6) 276 Rocks, boulders, sand 
G18 Aug.19 Small tributary of R. 49°00’ 66°58’ 1 ft 11.0 Fresh Stones 
des Grands Méchins 
Woodlands _ — Under logs and leaf 
mould 
G20 Aug.21 Capucins estuary 49°02.5’' 66°51.5’ LW toHW — Brackish Tidal flats 
R. at mouth 16.2 Stones 
G21 Aug.25 Capucins Pond 49°02’ 66°53" Natt — _ Sand, mud 
G22 Aug.25 2milesWof Cap Chat 49°05’ 66°45.5’ Subtidal NOLS 2825 Sand, pebbles, boulders 
to HW 
G23 Aug.23 Cap Chat, estuary 49°06’ 66°41’ LW — Brackish Sand and mud flats 
G24 Aug.23 Pointe Ste-Anne-des- 49°07.5’ 66°33’ LWtoHW 102 274 Rocks and boulders 
Monts 
G25 Aug.23 R.Ste-Anne estuary 49°07' 66°30.5’ LWto HW — Brackish Pebbles, mud 
G26 Aug.16 Tributary of R. 49°00’ e6°2 1" 1 ft 10.4 Fresh Waterfalls, under 
Ste-Anne mosses and stones 
Woodlot at stream 
G27 Aug.16 R.Ste-Anne 48°59’ 66°19’ 1-2 ft 15.5 Fresh Rapids, under stones 
G29 Aug.15 Smallstream 2 miles 49°13’ 66°07’ att 11.4 Fresh Under stones and 
W of Marsoui pebbles 
G31 Aug.15 Mont-St-Pierre 49°14! 65°47' LWtoMW 124 27.2 Slate bedrock 
G32 Aug.14 Mont-Louis 49°14’ 65°44" HW - _ Sand, stones, shells 
G33 Aug.14 Lacdel’Anse AG? 1-315" 6523 74 1 ft 12.0 Fresh Fine gravel along shore 
Pleureuse 
G34 Aug.14 Riviére-la-Madeleine 49°15’ 65°19.5' LWtoHW 14.6 27.2 Sand and pebble spit 
G39 Aug.13 Grand Etang, harbour 49°08’ 64°44.5’ LWtoHW 12.4 Brackish Slate and stone 
to 27-2 
G41 July 25 SE of Riviére-au- 48°59).5’ 64°23' EWAVCen NYA“ testae/ 227/57 Slate, pebbles, sand 
Renard 
G42 July 26 Anse au Griffon 48°56.5’ 64°18.5' LWtoHW 14.0 27.2 Shale, pebbles, sand 
G44 July 26 Jersey Cove 48°53.5’ 64°14.5’ LWtoHW 11.9 27.4 Slate, boulders 
July 22 R.del’Anse au Griffon 13.4 Fresh 
5 miles above mouth 
G45 July 26 CapdesRosiers, light 48°51’ 64N127 MWtoHW 13.2 268 Shale, pebble beach 
G46 July 26 Cap Bon Ami Park 48°47’ 64°12’ Terrestrial — _ Under stumps and logs 
G47 July 25 Cape Gaspé 48°45’ 64°10’ MWtoHW 14.7 25.7 Boulders, sand 


26 


St. Lawrence Estuary and Gaspé Coast 


Sta 
No. 


G48 
G50 
G51 


G52 


G53 


G54 
G55 
G56 


G58 


G59 
G60 
G61 
G62 
$2 
$3 


$5 
S6 


$8 
Sg 


$10 


S12 


$13 


$14 
$15 


S16 


S17 


$18 


$21 


$22 


$23 


Date 


July 25 
July 25 
July 27 


July 27 


July 24 


July 24 
July 27 
July 28 


July 28 


Aug. 8 
July 28 
July 29 
July 29 
Sept. 9 
Sept. 9 


10 
10 


Sept. 
Sept. 


10 
11 


Sept. 
Sept. 


Sept. 11 


July 16 


July 16 
July 17 


July 18 
July 19 


July 15 


July 20 


July 20 


July 13 


July 13 


July 12 


Locality 


Little-Gaspé 
Peninsula-Gaspé 


Bay inside Peninsula- 
Gaspé 


Gaspé Bay, 2 miles W 
of Peninsula-Gaspé 


Darmouth R., 3 miles 
above St-Majorique 


St-Majorique 
Gaspé Bay, N arm 


Gaspé Bay, S arm 


Gaspé Bay, sandy 
spit at neck 


Haldimand, south 
R. St-Jean, mouth 
Pointe St-Pierre 
Barachois 
St-Michel beach 
St-Vallier 


Montmagny, west 


Montmagny, river at 
St-Pierre bridge 


L’lslet wharf 


St-Jean-Port-Joli 


St-Roch-des-Aulnets 
wharf 
Wharf 


Pointe aux Orignaux 


Cap au Diable 


Pointe des Caps 


Pointe de la Riviere 
du Loup 


Cacouna |., W end 
E end 


R. Verte, 3 miles W 
of St-Modeste 


R. Verte, 2 miles W 
of St-Modeste 


St-Fabien, near 
Alcide Rock beacon 


Bic, Bicoques, wharf 
ruins 


Bare |., Wend 


Lat N Long W Depth ae Salinity Habitat 

48°47.5' 64°14 LWtoHW 15.8 27.1 Pebbles, sand 

48°51' 64°25’ LWtoHW 15.0 26.2 Sand beach 

48°51' 64°26’ LW 246 22.9 Sandy mud flats, 
estuarine 

48°52’ 64°28’ LWtoHW 20.4 Gis Sand, mud flats, 
sand and pebble beach 

48°54' 64°37' 1 ft 19.8 Fresh Pebbles, gravel 

48°53' 64°33: 5/ LW 20.5 74 Sandy estuary 

48°52’ 64 si2- MWtoHW 20.3 24.9 Sand spit 

48°49’ 64°31’ LW 19.4 21.1 Estuary, pebble beach, 
eelgrass 

48°49’ 64°24' MW toHW 18.6 26.0 Fine sand 

48°46.5’ 64°25.5' HW = — Sand bar 

48°46’ 64.225" Vit 20.0 Fresh Pebbles, gravel 

48°38’ 64m” LWtoHW 14.8 27.0 Sand beach 

Omori oven a 7/! MWtoHW 16.1 26.0 Rocks, sand 

46°52.5’ 70°54.5' MW -- — Shale, pebbles, sand 

46°54’ 70°49.5' LWtoHW = 18.7 0.3 Slate, pebbles, sand, 
bedrock 

AG? 58.5.) /O° S85. aw to AW 220.5 0.8 Slate and silt, bedrock 

46°59! AOC 33-52 elt 17.8. Fresh Silt, stones 

AT AOS. FOC 22°52 AW to FINN! Sl7.-S 1.9 Slate bedrock, silt 

A 13 70°16.5' LWtoHW 158 4.9 Slate bedrock, mud, 
sand 

AT 19" 70° 10:57 LW 15.0 8.3 Mud, bedrock pools 

col foctat |S 7102 110s EW 24.0 8.3 Boulders, mud 

47°29' 70°01.5' EWto AW 12:8 19.7 Boulders, sand, mud 

ASD. 69°56’ LWtoHW 19.8 19.2 Rocks, boulders, sand, 
mud 

47°43.5' 69°40.5' LWtoHW 25.0 208 Boulders, stones, mud 

ANTES INO}! 69°34’ LWtoHW 15.0 22.6 Slate, boulders, mud 

ANT polo Sys oyiment OKs) reo lhe LWtoMW 10.4 244 Boulders 

A iy: 69°30’ MW to HW = — Sand flats, beach 

AT? ST" 69°26’ 1 ft 20.8 Fresh Stones, gravel 

47°51' 69°25’ 1 ft 15.2 Fresh Under stones 

AS" 185") 68754" LW to HW 97. 262 Coarse sand, rocks, 
boulders 

ASDA OG 4 Geoe EVA ton) OO) 25721 Boulders 

AS 26.5) (682355) EWitoHW i408) 26:2 Rocks, mud flats 


27 


Geographical and Ecological Data 


Sta Date Locality Lat N Long W Depth Temp Salinity Habitat 
No. AE %o 
$24 July 10 Rimouski R., 13 48°23’ Ga So" 1 ft 19.1. Fresh 
miles above Rimouski 
$25 July 11° Branch of Rimouski 438°27.5' 68°24.5' «1 ft 17.0 Fresh 
R., 3 miles SE of 
Rimouski 
S26 July8 Smallstream 2 miles 48°27.5' 68°27’ iit 20.0 Fresh Stones, rapids 
SE of Rimouski 
$27 July8  Pointe-au-Pére,W side 48°31’ 68°28’ LW to HW 7.8 28.0 Rocks, boulders 
July 11 Eside LWtoHW 13.2 24.1 Rocks, boulders 
$28 July8 2 miles E of 28 Sib. MOS 247" LW 13tO} 27-0 Sand beach 
Pointe-au-Pére 
July 9 LW to HW — — Rocks, boulders 
$29 July 10 4miles E of 48°32’ 68°26' LWtoHW 10.9 — Boulders, mud, Fucus 
Pointe-au-Pére 
$30 July 12 Stream at Ste-Luce 48°33’ 68°24’ 1 ft 27.5 Fresh Mud 
Tourist Bureau 
$31 July 12 Stream 3 miles 48°31’ 68°22! 1 ft 10.5 Fresh 
W of St-Donat 
$32 July9 Stream 2 miles 48°29’ 68°19’ 1 ft 12.5 Fresh Stones 
NW of St-Marcellin 
S33 July 10 Ste-Luce, west 48° 33' 68°23.5' MW 10:9 25:9 Rocks, boulders 
S34 July 7 Rimouski, estuary 48°26.5’ 68°32.5' LWtoHW 16.5 13.9 Muddy sand, kelp, 
fucoids 
N3  Sept.8 Cap Tourmente 47°05’ 70°48! LWtoHW 19.2 2a3 Rocks, mud flats 
N4  Sept.8 Baie St-Paul, 47°25! TO 2925 LW to VN) 10:3) 20" Muddy sand flats 
outer bay, S side Fucus 
N5  Sept.6 Small tributary 47° 28' 70° 34’ 1 ft 10.7. Fresh Stones, wood 
of R. Baie-St-Paul 
N6  Sept.6 Baie St-Paul, W of 47°25.5' 70°29.5' HW — — Sand beach 
wharf 
N7 Sept.6 St-Joseph-de-la-Rive 47°27' HOw 22. LWtoHW 11.9 17.3 Boulders, muddy sand 
N8 Sept. 7 Pointe au Pic 47°37.5' 70°08.5' LWto HW 7.5) =25.0 Boulders, pebbles, 
sand flats 
N10 Sept.5 Port au Saumon 47°45! 69°57' LWtoHW 11.3 19.2 Mud, sand, pebbles 
wharf 
N11 Sept.5 St-Simeon, beach 47°50.5' 69°52.5' LW to HW 9.1 22.1 Rocks 
E of wharf HW. Sandy beach 
N12 Sept.4 R.dela Baie des AP By" 69°49' dott 15.2. Brackish Boulders, mud 
Rochers to fresh 
N13 Sept.4 Petit Lac Louis 48°02’ 69°49’ >1 ft 18.8 Fresh Sand, wood chips 
N14 Sept.4 Baie Ste-Catherine AYO )7/4 69°43’ LWtoHW 104 20.3 Coarse sand, rock 
N16 Sept.3 Baie du Moulin 48°09’ 69°39.5' LWto HW 8.3 24.9 Sand, sand flats 
a Baude 
N17 Sept.2 Pointe a John AS 3.5. 169-33: EVV ETON SOP 297 Sandy, gravelly mud 
N18 Sept.2 Les Escoumains, AS? 24! 69°24’ LW to HW 96 —_ Stones, sandy mud 
estuary 
N19 Sept.1  Tlets Penchés AS°24'5" 69°19’ ~“ LWtoHW 12.6 26:1 Clay mud over sand, flats 
N21 Sept.1 Pointe de Mille- A8°34.5' 69°08" MW, HW 104 28.3 Sand beach 


28 


Vaches 


St. Lawrence Estuary and Gaspé Coast 


Sta Date Locality Lat N LongW Depth Temp Salinity Habitat 

No. “Cc %o 

N22 Aug.31 Portneuf 48°38.5’ 69°05.5’ LWtoHW _ - Sand flats 

N23 Aug.31 Forestville wharf, 48°44.5' 69°03' LWtoHW 84 27.6 Sand beach and 
at Rocky Point breakwater 

N24 Aug.31 Small stream W of 48°45.5’ 69°04’ 1ft 12.6 Fresh Falls and rapids 
Baie Laval 

N26 Aug. 30 Tlets Jérémie 48°53’ 68°47.5' LWtoHW _ ~ Sand beach, mud 

N27 Aug. 30 Bersimis 48°56’ 68°39’ HW _ - Sand beach 

N29 Aug.28 Outardes estuary, 49°03.5' 68°33.5' MW — Brackish Mud 
Rageneau wharf 

N31 Aug.28 Chutesaux Outardes 49°08’ 66°29" Tt — Fresh Sand, mud 

N33 Aug.28 Pointe aux Outardes 49°02.5' 68°26’ LWtoHW 11.2 26.0 Sand, muddy sand flats 

N34 Aug. 29 Pointe LeBel 49°10’ 68°12’ - LWtoHW. 183° "456 Sand beach, flats 

N35 Aug. 29 Oldmill, 49°11’ 68°14.5’' LW toHW — Brackish Rocks, mud flats 
Manicouagan 

N37 Aug.27 Baie Comeau, 49°15’ 68°08.5' LW toHW 9.7..27.0 Boulders, sand 


cove N of wharf 


29 


Geographical and Ecological Data 


Table 3. St. Lawrence Estuary (1969) 
Key to station locality on Map 1A: L - lower estuary, Gulf of St. Lawrence 


Sta Date Locality 

No. 

L1 July 30 Pointe a John, NE 
of wharf 

L2 July 30 Les Escoumains, NW 
of Harbour I. 

L3 July 30 Forestville beach, 
inside wharf 

L4 July 31 Forestville, shore 
and breakwater 

L5 July 31 Forestville, outer 
flats at mouth of river 

L6 Aug. 1 Baie de la Boule, 
near Hall Point 

ETE Aug. 1 _~ Pointe de Moisie, 
mouth of R. Moisie 

L8  Aug.2_ Port-Cartier, at 
Dreyfus terminal 

LQ Aug.2 ___~ Port-Cartier-Ouest 

L10 Aug.2  R.Brochu 

L11 Aug.3 Pointe aux Basques 

L12 Aug.4__ Ile Grosse Boule, at 
north end 

L13 Aug.4 ile Grande Basque, 
NE bay 

L14 Aug.5  Lacdes Rapides 

L15 Aug.6 Amory Cove, near 
mouth of R. Matamec 

L16 Aug.7 Baie des Homards 
(R. Pentecdte N.) 

L17 Aug.7__ Ilets Caribou 

L18 Aug.7 Pointe des Monts 

L19 Aug.8 Off Papinachois 


30 


Lat N Long W Depth Temp. Salinity 
a @ %o 
480 1o.5, GO So: LW, HW TSE 24-2 
AS o2i 69°24’ LW — Brackish 
48° 44.5’ 69°03’ HW. — Brackish 
48°44.5’' 69°03’ LW, HW = = 
48°44’ 69°03.5’ LW 14.5 Nearly 
fresh to 
brackish 
50° 12.5’ 66°14.5' LWtoHW 14:8 274 
5OST1.5, 66704 LW, HW. 20.5 Brackish 
50°02’ 66°46’ LW. = — 
50°01’ 66°53’ LW, HW 15.6 19.5 
50°06.5’ 66°42’ HW = = 
BO ade 66°22’ LW, HW 14.5 = 
50°09.5' 66°17.5' LW, HW 18) e270) 
5OC 1055" 66222’ LW to MW 14.5 = 
5O eds: 66°25’ Shore to 20.2 Fresh 
2 ft 8.0 Spring 
50°18’ 65°57' LWtoHW 15.0 24.5 
49°49’ 67°08’ HW 16.0 High, 
brackish 
49°30’ 7a MW, HW Warm Brackish 
49°19’ 67223) LWtoHW 15.5 27.1 
49°00’ 68°38’ Naturalist 
dredge High, 
2-145 ft 13.5 brackish 
15-16 ft — — 
le=2Oitt = — 


Habitat 


Coarse sand, silt, 
pebbles, fucoids, 
Laminaria, pools in 
igneous bedrock 


Mud, sand 


Coarse to medium 
black sand 


Sand, fucoids, 
igneous bedrock pools 


Coarse to medium 
sand, river detritus 


Medium to coarse 
wave-exposed sand 


Very coarse sand, HW 
debris 


Oil-polluted muddy 
tidal flats over bedrock 


Igneous bedrock pools, 
boulders, sand, Fucus, 
Cladophora 


Steep wave-exposed 
sand beach 


Coarse-to-medium- 
sand beach, HW debris 


Sand, boulders, fucoids, 
Corallina on bedrock, 
HW debris 


Coarse sand, fucoids, 
bedrock 


Clear acidic water, 
fine-sand bottom 


Tidal lagoon, fucoids, 
sand beach, igneous 
bedrock pools 


Sand, fucoids, Entero- 
morpha, pools in 
igneous bedrock 


River bed, coarse sand, 
boulders, Fucus, HW 
debris 


Igneous bedrock, boul- 
ders, gravel, sand, 
Fucus 


Sand 
Muddy sand, kelp 
Clay, clay sand 


St. Lawrence Estuary 


Sta Date Locality 

No. 

L20 Aug.9 Lac Cinq Cents 

L21 Aug.11 Cap Colombier, E end 
of Baie du Plongeur 

L22 Aug.11 Cap Colombier, E bay 

L23  Aug.12 Franquelin 

L24 Aug.12 ° Mistassini estuary 

L25 Aug.13 Cap 4al’Orignal 


Lat N 


49°17' 


48°49.5' 


48°49.5' 


49°17’ 


49°17" 


48°22' 


Long W 


68°07’ 


68°53’ 


68°51.5’ 


67°54’ 


67°57" 


68°47.5' 


Depth 


Shore to 
2 tt 


LW 


Naturalist 
dredge 
TS tt 

20 ft 

25 ft 


LW to 
subtidal 


HW 


LW to HW 


Temp _ Salinity 


a 
22.0 


10.6 


12.2 


%o 


Fresh 


Brackish 


Habitat 


Stones, logs, dead 
leaves 


Mud and boulder 
flats, silty sand, fucoids 


Sand, filamentous algae 
Sand 
Sand, stones 


Sand flats, boulders, 
Fucus, kelp, wood chips 


Sand, gravel, HW-drift 
debris 


Slate bedrock, stones, 
silt, kelp, fucoids 


| 


Geographical and Ecological Data 


OF ST. LAW RENC E 


a 


S31 S25 
$32 S24y 


Map 1B Collection stations in the southwestern Gulf of St. Lawrence 


32 


Southwestern Gulf of St. Lawrence 


Table 4. Southwestern Gulf of St. Lawrence (1960) 


Key to station localities on Map 1B: S - Nova Scotia; P - Prince Edward Island; M - Magdalen Islands; 
B - New Brunswick 


Sta 
No. 


$1 
$2 


$3 
$4 


$5 


S6 


$7 


S8 


s9 
$10 


$11 


$12 


$13 


$14 


$15 


$16 


$17 


$18 


$19 


$20 


$21 


$22 


Date 


June 16 


June 16 


June 17 


June 20 


June 20 


June 21 


June 21 


June 22 


June 22 


June 22 


June 23 


June 23 


June 23 


June 24 


June 24 


June 26 


June 26 


June 26 


June 27 


June 28 


June 28 


June 28 


Locality 


Port Howe 
Pugwash, below 
bridge 

Wallace 

E shore Antigonish 


Harbour, at Chisholm 
Farm 


Dunn Beach, outer 
shore 
Monks Head, outer 


shore 


Pomquet Harbour, 
western pond 


Bayfield, wharf 


Bayfield, beach 


Pomquet Harbour, 
channel mouth 


Mahoney Beach, 
inner shore 


Antigonish Harbour, 
2 miles below 
Antigonish 


Chisholm Cove 


South R. estuary, 
opposite Chisholm 
Cove 


Cape Susan Cove, 
at shore road 


Nyanza, 
Bras d‘Or Lake 


St. Patricks Channel, 
South Bay 


Red Point, near 
Jamesville 


Merigomish Harbour, 
Y% mile E of Merigo- 
mish 


Big |., at Merigomish 
Harbour entrance 


Big |., inside 
entrance hook 


Merigomish Harbour, 
N side near head 


Lat N 


45°51! 
45°51! 


45°49! 
45°38 
45°41! 
45°39! 


45° 38.5’ 


45° 38.5’ 


45°39’ 
45°38’ 


45°42’ 


45°38’ 


45°38’ 


45°38’ 


45°57.5' 
46°05.5' 
45°59.5' 
45°56’ 


45°38.5' 


45°39! 
45°38’ 


45° 40.5’ 


Long W 


63°45’ 
63° 40' 


63° 28' 


61°54’ 


61°53’ 


61°49.5' 


61°50.5’ 


61°45.5' 


61°45.5' 
61°47.5' 


61°54.5' 


61°57.5' 


61°55" 


61°54.5' 


61°32’ 


60°53" 


60°59’ 


60°52’ 


62°24.5' 


62°28’ 


62°27" 


62° 2" 


Depth 


LW 


LW 


MW 


LW 


LW to HW 


LW to HW 


LW 


LW 


LW 


LW to HW 


LW 


LW 


7 it 


Ekman grab 


LW shore 


5-6 ft 


Dredge 


LW 


LW 


MW 


LW to HW 


LW 


LW 


LW to HW 


LW 


Temp 
sa 


19.0 


17.0 


13.5 


18.8 


20.3 


18.8 


18.6 


18.4 


22.5 


Salinity 


%o 


Brackish 


27.6 


29.5 


Low, 
brackish 


29.1 


274 


Habitat 


Tidal flats 


Coarse sand, Spartina 
flats 


Mud and eelgrass flats 


Mud and eelgrass flats 


Coarse sand, gravel, 
pebbles 


Muddy sand, stones, 
pebbles, Fucus, Chorda 


Fine muddy sand, 
Chorda, eelgrass, 
filamentous algae 


Stones, hard-packed 
sand 


Hard-packed sand flats 


Sand, pebbles, eelgrass 


Coarse sand, gravel, 
eelgrass 


Spartina flats, pebbles, 
mud 


Black muck, eelgrass 


Black muck, shells, 
eelgrass 


Mud and eelgrass flats 


Gravel, dead eelgrass, 
Enteromorpha 


Sand and eelgrass 
beach 


Sand and gravel spit, 
mud, shells 


Mud, muddy sand, 
stones, eelgrass, 
pilings 

Sand, stones, fucoids, 
Chorda 

Spartina flats, muddy 
sand 


Loose sand, eelgrass 


33 


Geographical and Ecological Data 


Sta Date Locality Lat N Long W Depth Temp Salinity Habitat 

No. 2C %o 

$23 June 29 Merigomish Harbour, 45°39’ 62°24.5' 10-25 ft - _ Black muck and 
channel, Masa’s Beach eelgrass 

$24 June 29 Abercrombie North, A5° 35.5" )62° 42% iW — — Muddy sand and stones 
East River 

$25 June 29 Seabreeze Point, 45°39’ 62°42.5' LW we iltshen  Arsits! Sandy mud, stones, 
opposite Pictou eelgrass, Chorda 

S26 June 29 Pictou Landing Cove 45°40’ 62°38' LWtoMW 21.5 — Mud, stones, Spartina 

$27 July5 Wallace Harbour, 45°49.5’ 63°29' LWtoHW 240 27.5 Mud, gravel, stones, 
at bridge eelgrass 

$28 July 5 Smith Point, near 45°51’ 63°24’ LW — — Sand flats, shells, 
Oak |. eelgrass 

$29 July6  PortPhilip,atbridge 45°51’ 63°44" LW 19.6 22.8 Mud, stones, eelgrass 

$30 July6 Pugwash Harbour, 45°51’ 63°40’ LW 20.3 _ Mud, stones, eelgrass 
at bridge 

$31 July 7 River John, % mile 45°45’ 63°04’ LW,HW 22.3 Variable Mud, stones, eelgrass 
below bridge 

$32 July 7 Murray Beach 45°46’ 63°07’ LW,HW — -— Sand flats 

$33 July 7 Barachois Harbour, 45°44’ 63217201 W -- _ Mud flats, eelgrass 
at bridge 

$34 July 7  Bayhead, beach at 45°45’ 63°22") CRW, AY 23.8) 42008 Muddy sand, eelgrass 
mouth of Millard 
Creek 

$35 July8 Wallace Harbour, mid- 45°49’ 63528) 15-33 ft _ — Mud, sand, shells, dead 
channel, off wharf Ekman grab eelgrass 

$36 July8 Northport 45°56’ 63252 LW Papilsy | PAS), Sand, sandy mud, 

eelgrass 

$37 Aug.22 Ottawa House Beach, 45°22’ 64°20' HW - — Steep, coarse, gravelly, 
near Parrsboro sand and shingle beach 

Pal July9 = Hillsborough Bay,at 46°14’ 63°07' LW 18.6 29.2 Mud, clay, silt, eelgrass, 
Charlottetown RR shells 
bridge 

P2 July 10 Keppoch Beach AGo 2. 63072 TLV OZR OM Fine sand, stones, 

Fucus, Chorda 

P3 July 11 Yorke estuary, at 46°16’ 63°00" LV 20.4 6) Clay, mud, stones, 
causeway Enteromorpha 

P4 July 11 Rocky Point,atferry 46°13’ 63°09"... LW 125. .29:5 Sandy mud, stones, 
wharf eelgrass 

P5 July 12 Blooming Point, A6°23.5’ 62°59" LWtoMW 22.6 29.5 Sand, sandy mud, 
Tracadie Bay eelgrass, Ruppia 

P6 July 13  Tracadie Harbour, 46°25’ 63°03" LW 21.0 28:6 Sand, dead eelgrass 
at mouth 

P7 July 13 Charlottetown Har- 46°13.5’ 63°09.5" LW Z1:0 _ Sandy mud, eelgrass 
bour, Yorke estuary 
mouth 

P8 July 14 Yorke estuary, off 46°15’ 63°09’ 1-3 ft — — Mud, dead eelgrass, 
Charlottetown Oyster tong shells 

hauls 

P9 July 14 Yorke estuary, mid- 46°14! 63°10’ 30-40 ft _ Black muck, dead 

channel Ekman grab eelgrass 


34 


Southwestern Gulf of St. Lawrence 


Sta Date Locality Lat N Long W Depth emp Salinity Habitat 

No. i %o 

P10 July 15 Stanley Bridge, at 46°28’ 63°27.5' HW _ _ Sand, dead eelgrass, 
landing stage under logs 

P11 July 16 New London Bay, 46°30’ 63°28’ 10-15 ft 22.0 — Mud, muddy sand 
mid-channel Dredge 

P12 July 16 NewLondon Bay, 46°29'6"" 63°26.5' 6-7 i — — Sandstone, eelgrass, 
E side Dredge fucoids, Chondrus 

P13 July 16 NewLondon Bay, 46°28’ 63°28’ 12-14 ft _— _ Fine black mud 
Y2 mile above bridge Ekman grab 

P14 July 17 Rustico Harbour, 46°27.5' 63°17:5' Get 19.8 ~ Sandy mud, shells 
channel off Rustico Ekman grab 

P15 July 17 Rustico Bay, off 46°27’ es° 16’ 5-8 ft — ~ Sandy mud, dead 
Little Harbour wharf Ekman grab eelgrass 

P16 July 17 North Rustico, channel 46°27.5’ 63°18’ 3-4ft — -- Mud, chord grass, 
below wharf Oyster tong eelgrass 

hauls 

P17 July 18 Linkletter Shore, 46°24’ 63°51' MWto HW — = Mud, sand, sandstone 
Bedeque Bay bedrock 

P18 July 18 ETignish Run,outer 46°57’ 63°59' HW —- oa Sand, shells 
beach 

P19 July 18 Tignish Run, near 46°57' e4°O01" . 13 25.5 N20 Sandy gravel, eelgrass 
head, at bridge 

P20 July21_ Bideford estuary,near 46°37’ 63°55’ = 10-14 ft 20.8 — Fine mud, shells, dead 
Biological Station Ekman grab eelgrass 

P21 July 22 Indian Spit, Lennox!. 46°36’ 63°51’ 30-36) _ — Sandy mud, detritus 

Ekman grab 

P22 July22 Malpeque Bay, mouth 46°35’ 63°48" ~.25 it —_ — Fine mud, dead 
of Bideford Bay Ekman grab eelgrass 

P23 July 22 Curtain I., off 46°32’ 63°47.5' 8ft _ _ Sandstone, algae 
N W point Dredge 

P24 July 22 Malpeque Bay, 46°31' 63°46’ 15 ft -— — Sand, oyster shells 
Little Rock oyster bed Dredge 

P25 July 24 Lower Montague, 46°10’ B2°S2° HV 20.0 -- Sandstone, sand 
below lighthouse 

P26 July 24 Panmurel., N end 46°08.5' 62°28' LW. 20:2) 28.7 Sandstone, sand, 
of isthmus filamentous algae 

P27 July25 Murray Harbour N, 46°03’ 62°28) LW to HW — — Sand, clay mud, gravel, 
inside breakwater debris 

P28 July 25 Murray Harbour 46°03’ 62°30’ LWtoHW _ _ Sand, dead eelgrass 
mouth, inner side 
of spit 

P29 July 25 Lower Murray Har- 46°01’ 62°30’. EW 204 29.0 Muddy sand, sand, 
bour, above wharf eelgrass 

P30 July25 Woodl., ferry terminal 45°57’ 62°45' MW toHW -~ a Salicornia and Spartina 

flats 

P31 July 26 Pinette Harbour, 46°04’ 62°54’ LW 19.5 — Muddy sand, eelgrass 
at hwy bridge 

P32 July 26 Orwell Cove, atwharf 46°09’ 62°53' MW 20.0 28.6 Sand, shells, Spartina, 

Fucus 
P33 July 26 Morell wharf, 46° 26' 62°42' LW 215 — Sand, eelgrass 


St. Peters Bay 


oH) 


Geographical and Ecological Data 


Sta Date Locality Lat N Long W Depth Temp Salinity Habitat 

No. nC %o 

P34 July 26 Annandale wharf 46° 15.5’ 62°25" . EW 20.9 _ Sand, eelgrass, Fucus 

P35 July 27 Georgetown Harbour, 46°10.5’ 62°32.5' 30-35 ft — _ Sand, mud 
at channel buoy Ekman grab 

P36 July27 Brudenell estuary, 46°12’ 62°S5.5" 7-10 ff _ = Muddy sand, dead 
opposite Provincial Ekman grab eelgrass 
Park 

P37 July 27 Shaw Point, 46°11’ 62°33" GS ft _ _ Sandstone, algae 
Georgetown Harbour Dredge 

P38 July 27 Montague estuary, AG Ol.) 462 -SS35" Sift — — Sand, dead eelgrass 
inside Shaw Point Oyster tongs 

P39 July 27 Victoria, at wharf 46°13" 63°29.5' MWtoHW 20.0 _ Sand flats, sandstone, 

rocks 

P40 July 28 Conway Narrows, 46°44’ 64°00° MW — _— Sand, eelgrass, shells 
Cascumpeque Bay 

P41 July 28 Conway Narrows, 46°44’ 63°59’ Ya-2 ft _ — Sand, eelgrass, shells 
shoal opposite large Ekman grab 
house 

P42 July 28 Conway Narrows, 46°43’ 63°58’ fas ht = = Sand, eelgrass 
at Flat |. Ekman grab 

P43 July 29 Bideford estuary, at 46°37' 63°55’ 10-12 ft 21.0 — Soft ooze 
Biological Station Ekman grab 

M1 Aug. 15 House Harbour, 47° 26' 61°50' MWto HW a _ Fine sand 
Dune-du-Nord 

M2a Aug.15 House Harbour, 47°24.5' 61°52’ 10 ft 20.0 _ Mud, dead eelgrass 
channel off Ekman grab 
Le Grand Ruisseau 

M2b Aug.15 House Harbour,atLe 47°25’ 61°52’ Surface 1 ft = — Mud, eelgrass 
Grand Ruisseau Oyster tongs 
wharf 

M3 Aug. 15 Pond, N side of 47°21' 61°53.5’ LW — _— Soft mud, eelgrass, 
Pleasant Bay fucoids 

M4 Aug.16 House Harbour,Eside 47°24.5’ 61°51’ 10-15 ft _ — Sandy mud, shells 
channel Dredge 

M5 Aug.16 House Harbour, N 47°26’ 61°47.5’ 6-10 ft _ — Sandy mud, dead 
channel Ekman grab eelgrass 

M6 July 16 Alright Channel, 47° 28' 61°46.5' LW — = Sand, stones, eelgrass 
at causeway 

M7 July 16 Oyster Pond,NEend, 47°33.5' 61°32’ HW _ _ Fine sand, shells, 
inside barrier beach eelgrass 

M8 July 16 Grand Etang Harbour, 47°34.5’ 61°29 MW _ — Sandstone, eelgrass 
at Old Harry wharf 

M9 July 16 Grosse Tle, nearLeslie 47°37.5' 61°31.5' HW _ — Sand at outlet of stream 

M10 July 16 Wolf l.,N end a7" 32° 61°43’ HW — — Sand dunes, shells 

M11 July 16 South Beach, near 47°24!’ 61°45’ HW — -- Fine sand, shells 
wharf 

M12 July 16 Etang-du-Nord, at ss fe ag 61°57'5" HW — -- Fine-sand beach, 
wharf Chondrus debris 

M13 July 16 Basques Harbour, 47°20. 5' “61° 56"" HY — _ Sand, shells 


36 


N entrance 


Southwestern Gulf of St. Lawrence 


Sta 
No. 


B2 


B3 


B4 


B5 


B6 


B7 


B8 


B9 


B10 


B11 


B12 


B13 


B14 


B15 


B16 


B17 


B18 


B19 


B20 


B21 


B22 


B23 


Date 


Aug. 


Aug. 


Aug. 


Aug. 


Aug. 


Aug. 


Aug. 


Aug. 


Aug. 


Aug. 


Aug. 


Aug. 


Aug. 


Aug. 


Aug. 


Aug. 


Aug. 


Aug. 


Aug. 


Aug. 


Aug. 


Aug. 


Locality 


Cocagne |., near 
Cormierville, inner 
shore 


Richibucto Harbour, 
channel off Fagan 
Point 


Richibucto Harbour, 


channel below 
NW arm 


Richibucto Harbour, 
inside North Beach 
Richibucto Harbour, 
off Jardine Beach 


Richibucto, estuary 
at Rexton Bridge 


Tabusintac Lagoon, 
Stymas Cove 


Tabusintac I., 
inside channel 


Wishart Point, 
channel off wharf, 
Tabusintac Lagoon 


Brantville wharf 


Tracadie Lagoon, 
channel below 
Tracadie 


Tracadie Lagoon, 
N channel 


Tracadie Lagoon, 
old wharf 


Tracadie outer beach, 
at causeway 


Little Pokemouche 
Lagoon, near mouth 


St-Simon Inlet, North 
Branch channel 


St-Simon-Nord, 
channel above church 


St-Simon-Sud, 
opposite Biological 
Station 


Miscou Harbour, 
near Portage Bay 


Pokemouche Lagoon, 
S side 


Neguac wharf 


Parlee Beach, Shediac 
Bay 


Lat N 


46°24’ 


46°41.5' 


46°42’ 


46°43’ 


46°40’ 


46°39’ 


47°18! 


47°17.5' 


47°20' 


47°22' 


47°31' 


47°33’ 


47°32! 


47°30’ 


47°41.5' 


47°45’ 


47°45’ 


47°44! 


47°52" 


47°38.5' 


47°14.5' 


46°14’ 


Long W 


64°36’ 


64°52’ 


64°51! 


64°49’ 


64°52’ 


64°52.5' 


64°57 5’ 


64°57’ 


64°57' 


64°56.5' 


64°54’ 


64° 53’ 


64°54’ 


64°52’ 


64°44’ 


64°49’ 


64°51.5' 


64°47’ 


64°33’ 


64°49’ 


65°05" 


64°30’ 


Depth Temp 


MW 25.9 


25 ft — 
Ekman grab 


30 ft _ 
Ekman grab 


1-4 ft — 
Ekman grab 


12-20 ft — 
Ekman grab 


55-60 ft = 
Ekman grab 


10-12 ft 
Oyster rake 


18.9 


6-10 ft _ 
Oyster rake 


12-30 ft _ 
Ekman grab 


LW 23.0 


20 ft = 
Ekman grab 


1-6 ft — 
Oyster rake 


LW 


HW _ 


LW 22.0 


10-25 ft 19.2 


Ekman grab 


4-6 ft _ 
Oyster tongs 


9-14 ft a 
Ekman grab 


LW, HW 21.0 


LW _ 
LW 18.9 


HW - 


Salinity 


%o 


26.4 


27.5 


28.3 


Variable 


26.6 


274 


Habitat 


Mud, coarse sand, 
eelgrass 


Sand, shells 


Sand, stones, shells 


Sand flats, eelgrass 
Sandy mud, mud, dead 
shells 


Woodchips, mud 


Sand, peat, shells, 
eelgrass 


Sand, eelgrass 


Clay mud, sand, shells 


Soft sandstone, mud, 
eelgrass, fucoids 


Sand 


Sand, mud, shells, 
eelgrass 


Muddy sand, shells, 
eelgrass 


Sand, shells 


Sand, eelgrass, mussel 
beds 


Sandy mud, shells 


Eelgrass, oyster beds 


Sandy mud, stones, 
shells, sponges 


Sandy mud, sandstone, 
shells, eelgrass 


Peat, sand 
Mud, muddy sand, 


stones, eelgrass 


Sand 


37 


Geographical and Ecological Data 


Sta Date Locality Lat N Long W Depth Temp Salinity Habitat 

No. 2C %o 

B24 Aug.7 Bourgeois Office 46°19’ 64°31' LWtoHW — _ Sand flats, sandstone 

blocks, eelgrass 

B25 Aug.7 Grande-Digue, at 46°.17.5' 64°33". UW. 27.9 - 282 Mud, muddy sand, 
wharf eelgrass 

B26 Aug.14 Shediac Harbour, 46°14’ 64°32.5' 10 Tt _ _ Mud, eelgrass 
channel off wharf Oyster tongs 

B27 Aug.14 Shediac Harbour, off 46°15.5’ 64°33’ 6-10ft _ — Shells, sandy mud, 
Indian |. Oyster tongs eelgrass 

B28 Aug.14 Shediac Harbour, 46° 15’ 64°32" AZ Tt ZA28) 262 Sand, shells 
channel opposite Ekman grab 
Pointe du Chéne 

B29 Aug.22 Buctouche Harbour, 46°30’ 64°41’ LW 24.0 — Muddy sand, eelgrass 


mouth of Little 
Buctouche River 


B30 Aug.23 Richibucto Bay, at 46°46’ 64°55’ HW PANES -- Sandy mud, eelgrass 
St-Oliver 
B31 Aug. 23 Richibucto Bay, 46°47.5' 64°54.5' MW 215 — Sand, eelgrass 


Callander Beach 


B32 Aug.23 South Kouchibouguac, 46°51’ 64°57' =LW 21/6 = Eelgrass, sandstone 
estuary at mouth 


38 


Atlantic Coast of Eastern Nova Scotia 


Table 5. Atlantic Coast of Eastern Nova Scotia (1962) 
Key to station locality on Map 1C: A - outer coast of eastern Nova Scotia, and Cape Breton Island 


Sta 
No. 


Al 
A2 


A3 


A4 


AS 


A6 


A7 


A8 


AQ 


A10 


Atl 
A112 


A13 


Al4 


A15 


A16 


A17 


A18 


A19 


A20 


A21 


A22 


A23 
A24 


Date 


July 4 
July 4 


July 5 


July 6 


July 6 


July 6 


July 6 


July 9 


July 10 


July 11 


July 11 


July 12 


July 12 


July 13 


July 13 


July 13 


July 17 


July 18 


July 18 


July 18 


July 19 


July 19 


July 19 
July 20 


Locality 


Upper Oyster Pond 


Lat N 


44°42’ 


Pleasant Point, E beach 44°41’ 


Clam Harbour beach 


Ostrea Lake, 
Musquodoboit 
Harbour 


Ostrea Lake, at mouth 
of Frost Brook 


Oyster Pond, above 
bridge 


Lake Charlotte 


Beach opposite 
Conrod |. 


Meisner Head, outer 
beach 


Martinique Beach 


Little Harbour 


Clam Harbour beach 


Clam Harbour, at 
head 


West Chezzetcook 


Chezzetcook Inlet, 
at Dyke Rd. bridge 


Chezzetcook Inlet, 
opposite Conrod I. 


Lingan Harbour, Cape 
Breton I., at mouth 
of estuary 


Point Michaud Beach, 
NE end 


Point Michaud lagoon, 
at outflow 


Grand R. estuary, 
1% miles above mouth 


L’Ardoise Beach, bay 


Bourgeois Inlet, E arm 


False Bay, near head 


Lennox Passage, 
at Grandique Point 


44°43.5' 


44° 43' 


44°43’ 


44°41.5' 


44°47.5' 


44°41.5' 


44°40.5' 


44°41.5' 


44°42.5' 


44°43’ 


44°44! 


44°42' 


44°41.5' 


44°41.5' 


46°14’ 


45°34! 


45°34’ 


45°37’ 


45°36’ 


45°38’ 


45°38’ 
45°35.5’ 


Long W 


63°04’ 


63°04’ 


62°52’ 


63°04’ 


63°04.5' 


63°03 .5! 


62°57.5" 


63°13" 


63° 12.5" 


63°09’ 


62°51" 


62°52’ 


62°50" 


63° 15.5" 


63°15" 


63°14.5' 


60°02’ 


60°40’ 


60°41 


60°39’ 


60°45.5' 


60°57’ 


60°56.5' 
61° OT’ 


Depth 


LW to HW 
HW. 


HW 
LW 


LW to MW 


LW to HW c.15.0 


Terrestrial 


MW to HW 


LW 


LW to HW 


MW 
LW to HW 


MW to HW 


LW to HW 


LW 


LW 


LW 


LW to HW 


LW to HW 


LW to MW 


LW to HW 


LW to HW 


LW 
LW 


15.9 


18.9 


t32 


15.4 


13.2 


Salinity 


%o 


Brackish 


Brackish 


Low, 
brackish 


Brackish 


17.5 


29.0 


Low, 
brackish 


25.3 


27.9 


28.4 


Habitat 


Tidal pond at LW level 
Sand, rocks, seepage 
Surf-exposed fine sand 


Mud and eelgrass flats, 
Fucus, boulders 


Acid fresh water, 
boulders, fucoids 


Boulders, shells, 
filamentous algae, 
fucoids, HW debris 


Mixed woods 


Sand, gravel, salt- 
marsh flats, HW debris 


Boulders, gravel, sand, 
Fucus 


Boulders, fine sand, 
HW rock pools 


Sandy mud, stones 


Surf-exposed fine 
sand, igneous bedrock, 
HW pools 


Muddy sand and 
eelgrass estuary, acid 
fresh water 


Salt marsh, HW debris 


Salt-marsh flats, 
eelgrass, mud, stones 


Sandy mud, eelgrass 


Surf, sand, stones, 
detritus, eelgrass, 
Chorda 


Metamorphic rock, flat 
sand, fucoids 


Sand, fine gravel 


Eelgrass, mud 


Sand and mud flats, 
eelgrass, filamentous 
algae on stones 


Sandstone, mud, 
eelgrass, fucoids, stones 
at HW level 


Mud and eelgrass flats 


Sandstone, sandy 
mud, eelgrass, Chorda 


39 


Geographical and Ecological Data 


63° Northumberland BRETON 


ISLAND 


Strait 


Cape 
AT IL ASN Tole OCEAN Breton 


Island 


Map 1C Collection stations on the Atlantic coast of eastern Nova Scotia 


Sta Date Locality Lat N Long W Depth Temp Salinity Habitat 

No. 2G %o 

A25 July 20 Bourgeois Inlet, at 45°38’ 60°57.5' 28 ft _ — Mud, sandy mud, 
forks Ekman grab eelgrass 

A26 July 20 Bourgeois Inlet, 45°37" GO°S7’ 20ft _ _ Stones, algae 
inside mouth Ekman grab 

A27 July 20 Bourgeois Inlet, 45°38’ 60°58.5' 12-20 ft 7 - Soft black mud 
W arm channel Ekman grab 

A28 July 20 Petit-de-Grat,bridge 45°30.5’' 60°59' MW _ — Cold-water channel, 

mud flats 

A29 July 20 Sampson Cove, Isle 45°29.5'’ 60°55.5' MW toHW — — Surf-exposed break- 
Madame water and rock pools 

A30 July 20 Pondville Beach,Bay 45°32’ 60°58.5' LWtoHW 14.1 29.7 Bedrock, stones, sand, 
of Rocks fucoids, Lithothamnion 

A31 July 21 Haddock Harbour A5"'33’ 61°07", LW 14.2 298 Sandstone, Fucus, soft 


mud, eelgrass 


A32 July 21 Port Royal Harbour, Ag? 32" 61°05’ LW 14.5 _ Tidal flats, mud, 
at bridge eelgrass, Mytilus beds, 
shells 
A33 July 21. Inhabitants Harbour, 45°35’ 61°16' MWto HW _ _ Gravel and sand at 
at Port Richmond stream mouth 
A34 July 21 Inhabitants R., at ro es I dl 61°14’ LW 15.0 oF Mud, eelgrass, Mytilus 
Ferry Road beds, sandstone blocks, 
wharf pilings 
A35 July 22 Guysborough Harbour, 44°27’ 61°32" TW 13:9 25:8 Muddy sand, eelgrass, 
at Boyiston Park stones 


40 


Atlantic Coast of Eastern Nova Scotia 


Sta 
No. 


A36 


A37 


A38 


A39 


A40 


A41 


A42 


A43 


Ad. 


A45 


A46 


A47 


A48 


A49 


A50 


A54 


A55 


Date 


July 22 


July 22 


July 22 


July 22 


July 22 


July 22 


July 22 


July 23 


July 23 


July 23 


July 23 


July 23 


July 24 


July 24 


July 24 


July 24 


July 24 


July 25 


July 25 


July 25 


Locality 


Guysborough Harbour, 
at Lesterdale bridge 


Guysborough Harbour, 
at Havendale 


Guysborough R., at 
Guysborough Intervale 


Dorts Cove, mouth 
of Salmon R. 


Quéénsport, mouth 
of Rock Island R. 


St. Francis Harbour, 
at mouth 


Oyster Ponds, outer 
shore 


Marshall Cove, 
Whitehaven Harbour 


Denning I., 
Whitehaven Harbour 


Port Felix 
Glasgow Head 


Glasgow Harbour, 
NE arm 


Charlos Cove, W of 
bar 


Charlos Creek, at 
mouth 


Torbay Beach 


Weber Cove, at 
causeway 


New Harbour estuary 


Webb Cove, 
Golboro Harbour 


Coddles Harbour, 
inner bay 


Seal Harbour head, 
near West Brook 


Lat N 


44°27.5' 


A5°27' 


45°28’ 
45°21? 


45° 20’ 


45°26.5' 


45°27' 


45°14! 


45°13! 


45°15! 
45°19! 
45°19! 


45°14.5' 


45°14.5' 


45°12" 


Ao 1 AS: 


45°11' 


45°10’ 


45°10! 


45°09.5' 


Long W 


61°32.5’ 


61° 34.5’ 


61°36.5' 


61°28" 


61°16’ 


61° Ts" 


61°16’ 


64? 115 


61°10.5' 


61°14" 


60°58’ 


60° 58’ 


51:°20' 


GPoe a 


a) 225: 


61°21" 


64°27 


61° 38.5’ 


61°.32:5' 


61°35! 


Depth 


LW 


LW to HW 


MW to HW 


LW 


LW to HW 


LW 


LW 


LW 


LW to HW 


MW to HW 


HW 


LW 


LW 


LW 


LW to HW 


LW to HW 


LW to MW 


LW 


Temp Salinity 
2G %o 
— Low, 
brackish 
16.5 _ 
15.1 — 
(Inner 
pond is 
fresh) 
12.0 30.4 
tt “= 
13:1 = 
17.1 29.5 
135" 29.7 
125 _— 
1Be ~ 3:3 
129 28,7 
15.1 — 
(inner) 
e150 = 


Habitat 


Bridge abutments 


Gravel, mud flats, 
eelgrass, Spartina, HW 
debris 


Stones, coarse silt, 
filamentous algae 


Surf-exposed pebble 
and sand bar 


Steep gravel and 
sand beach, HW rock 
pools, fucoids 


Firm mud, Mytilus beds, 
eelgrass 


Stones, pebbles, 
coarse-sand patches 


Sandy mud and 
eelgrass flats 


Granite bedrock, 
Ascophyllum and 
Fucus, LW pools, 
HW debris 


Coarse-sand and 
granite-stone beach 


Surf-exposed boulders, 
stones, coarse sand 


Mud, eelgrass, shells 


Sandy mud, granite 
boulders, fucoids, 
eelgrass 


Mytilus beds, stones 


Exposed sand beach, 
sandstone and granite 
boulders, tide pools 


Sand, eelgrass, fucoids, 
rock, Ascophyllum 


Sand, shells, eelgrass 


Stones, eelgrass, mud, 
detritus, fucoids 
Lithothamnion, fucoids, 
Irish moss 


Eelgrass, fine mud, 
boulders 


Mud flats, eelgrass, 
stones, detritus, fresh- 
water inflow 


41 


Geographical and Ecological Data 


Sta Date Locality Lat N Long W Depth emp Salinity Habitat 
No. “GC %o 
A56 July 25 Indian Harbour, at 45°06.5' 61°51.5' LWtoMW 21.5, Mixed Coarse, unconsolidated 
lake outflow over sand and stones, surf 
16.5 zone 
A57 July 26 Baraswa Cove, 45°05’ 61°50". )LW oo ao Mud and eelgrass flats 
inside breakwater 
A58 July 26 Wine Harbour, at 45°04.5' 61°51". RW 1522 — Sandy mud, eelgrass, 
shore road filamentous algae, 
shells 
A59 July 26 Port Bickerton,Eside 45°06’ 61°44" LW 14.8 30.4 Sand and eelgrass flats, 
stones, Fucus 
A60 July 30 Shiers Pond 44°55’ 62°18" “iWHoHW 182 _ Mud, shells, peat, 
eelgrass, Spartina, 
bridge piers, HW debris 
A61 July 30 East Quoddy Pond, 44°54’ 62.1825) JEW. 18.9 29.8 Stones, shells, kelp, 
at inlet bridge eelgrass, mud 
A62 July 31 Spanish Ship Bay,at 45°01’ 62°02’ LWto HW _ o- Stones, mud, detritus, 
Hooper Brook eelgrass, fucoids 
A63 July 31 Pye Point, Liscomb 45°00/5's 62°01". . LW 13.9 « 2631 Wave-protected sand- 
Harbour stone gravel, sandy 
mud, eelgrass, fucoids, 
boulders 
A64 July 31 MarieJoseph Beach 44°58’ 62°03' MWtoHW 12.5 Steep, surf-exposed, 
fine-gravel and coarse- 
sand beach 
A65 July 31 BakerCove,atMarie 44°58.5' 62°04.5' MW 19.5 Brackish Eelgrass, fucoids, 
Joseph detritus 
A66 Aug.1 Jewer Cove, near 44°56' 62°11’ LWtoHW 18.0 24.7 Boulders, Ascophyl/lum, 
Mitchell Bay eelgrass, shells, mud, 
cold freshwater stream 
outflow 
A67 Aug.1 Smith Cove, at hwy 44°58’ 62°13" “EW — _ Mud and eelgrass flats 
bridge 
A68 Aug.1 Moosehead, White #4°'56.5" 62° 16" - LW 16.4, 22.4 Sand, stones, eelgrass, 
Island Bay over at stream mouth © 
14.5 
A69 Aug.2  Soberlsland,atHurd 44°51’ 62°28.5' LW 15.9 20.4 Eelgrass, detritus, 
Cove bridge mud, Chondrus, fucoids 
A7O Aug.2 Malagash Cove, at 44°51' 62°32.5" LWtoHW 18:5 25:0 Mud, eelgrass, shells, 
bridge stones, boulders, 
Chondrus, detritus 
A71  Aug.2 Mushaboom Harbour, 44°52’ 62°34’ LW 18.0 _ Eelgrass, detritus, at 
at head wharf 
A72 Aug.2 Spry Bay, head of 44°50’ 62235" EW 18.2 — Protected sand, rocks, 
Tomlee Bay eelgrass, fucoids 
A73 Aug.3 Psyche Cove, south 44°48’ 62°33:5" LWtoe HW. 14:8 28,2 Fine sand, pebbles, 
end bedrock, filamentous 
algae, chord grass 
A74 Aug.3 Mason Cove, E side 44°48' 62°42’ LW c.20.0 — Eelgrass, detritus, shells, 
of Tangier Harbour Chondrus 
A75 Aug.3  PopesHarbourW,at 44°48’ 62°40' LW 22+ Eelgrass, stone chips, 


42 


head 


mud 


Atlantic Coast of Eastern Nova Scotia 


Sta Date Locality Lat N LongW Depth emp Salinity Habitat 

No. °C %o 

A76 Aug.3 _ Popes Harbour, 44°49’ 62°38’ LW 19.0 _ Eelgrass, detritus, 
near mouth of First to mud, sand, at edge 
Lake R. 21.5 of channel 

A77 Aug.4 _~ Ship Harbour, at 44°48’ 62°51.5' LW 19.5 Brackish Mytilus beds, stones, 
Rocky Brook detritus 

A78 Aug.4 LowerShipHarbour, 44°48’ 62°60’ UW 16.5 — Stones, Ascophyl/lum, 
above Whale Island . eelgrass, detritus 

A79 Aug.4 Oyster Pond, head 44°47' Gao" Lw 2u0-~ Bz Eelgrass, gravel, mud, 


of Jeddore Harbour stones, fucoids 


Geographical and Ecological Data 


M32 M26_\\(r_ 


M35 


M3233 M31 M30 
M39. : ., M28 


XX aM? 
S~M36 
< Qo 
® 


Map 1D Collection stations in northern New England and western Nova Scotia 


44 


Northern New England and Western Nova Scotia 


Table 6. Northern New England and Western Nova Scotia (1963) 


Key to station localities on Map 1D: S - western Nova Scotia; M - northern New England 


Sta Date Locality Lat N Long W Depth emp Salinity Habitat 

No. st %o 

$1 July9  St.MarysBay,above 44°34’ 65°54’ LW 170° 31.5 Sandstone, muddy sand, 
Red Bluff fucoids, Chondrus 

S2  July8 Centreville, Trout 44° 33’ 66°02' MWtoHW 104 32.3 Boulders, bedrock, 
Cove, Digby Neck kelp, Corallina 

$3 July9 _— E Sandy Cove, St. 44°29! 66°05’ LW 150 eSile7, Sandy mud, fine gravel, 
Marys shore, Digby fucoids, Zostera 
Neck 

S4 July9 Sandy Cove, Fundy 44°30’ 66°06’ LW, HW 10.0 _ Coarse sand, bedrock 
shore, Digby Neck pools 

$5 July9 Little River Wharf, 44°27' 66°08’ HW -- = Pools, drift debris 
St. Marys Bay 

S6 July9 Whale Cove, Digby 44°26’ 66.117 HW — _ Bedrock fissures and 
Neck pools 

S7 July9 Midway Lake, Digby 44°32’ 66°03’ Shoreline 22.0 Fresh Stones, gravel, Nuphar 
Neck 

S8  July9 _~ Barton old wharf, 44°32’ 65°53’ LWtoMW 16.8 31.7 Coarse sand, mud, 
St. Marys Bay Zostera, Cladophora 

S9 July 10 Brighton Bay head, 44°33’ 65°52" “EW toMy PT FSe “SNF Mud, sandy mud, 
near old wharf stones, fucoids 

S10 July 10 Meteghan Centre 44°12' 66°09’ LW 14.5 32.4 Sand beach, stones, 

gravel, fucoids 

$11 July 11 Gilbert Point, W of 44°29' 65°58’ LWtoHW 15.6 318 Pebbles, sandy mud 
lighthouse 

$12 July 11 Upper Saulnierville, 44°17! 66°08’ LW toHW — Brackish Stones, algae in stream 
at stream mouth outflow 

$13 July 12 Grosses Cocques, at 44°22! 66°06": LW 14.3 31.4 Fine-sand flats, Zostera 
old harbour entrance clumps 

$14 July 12 Meteghan R., salt 44°13’ 66°08’ HW _ = Salt-marsh pools, mud 
marsh banks, drift 

$15 July 13 Port Maitland, at 43°59’ 66°09’ LW 14.0 — Sand beach, slate- 
wharf pebble foreshore 

S16 July 14 Cape St. Mary, at 44°05’ 67°13" HW _ — Slate bedrock and 
wharf igneous boulders, 

HW-drift debris 

$17 July14 Eel Pond,HwyNo.3 43°50’ 65°56" - LW. 22.0 Brackish Gravel, mud, stones, 
culvert fucoids, Zostera 

$18 July 14 Cape Fourchu, at 43°48’ 66°09’ HW — _ Bedrock spray pools, 
Yarmouth Light HW drift 

$19 July13 Salmon R., South 44°03’ 66°10’ LWtoMW 13.8 32.1 Sand, pebbles, algae 
Beach 

$20 July13 Mavillette, at road 44°06’ 66°12? HW — Brackish Salt-marsh pools, 
bridge debris 

M1 July 16 Mount Desert Nar- 44°25’ 68°22’ LW 21.6 30.4 Mud, boulders, eelgrass, 
rows, W of causeway, fucoids 
Maine 

M2 July2 Pond Cove beach, 44°37’ 67°30' HW = _ Sand, pebbles 
near Roque Bluffs 

M3 July2 Roque Bluffs, salt 44°37' 67°29" | LAW 17.5 Brackish Mud, stones, gravel 


marsh above bridge 


45 


Geographical and Ecological Data 


Sta Date Locality Lat N Long W Depth Temp Salinity Habitat 
No. IG %o 
M4 July2 _ Little KennebecBay, 44°40’ 67°26" LW. 21+ Brackish Mud, gravel, H2S 
2 miles W of Larabee 
M5 July2 ~ Fort O’Brien Point, 44°41' 67°24’ LWto MW- — Brackish Stream mouth, mud, 
near Machiasport stones, fucoids 
M6 July3 Sandy River Beach 44°34’ 67°34’ LW 120 326 Coarse igneous sand, 
boulders, fucoids 
July 4 Sandy River Beach 44°34’ 67°34’ HW — — Sand 
M7 July3 Henry Point, E end 44°32’ 67°34’ LW _ — Bedrock, fucoids, debris 
Sawyer Cove, near 
Jonesport 
M8 July4 Little Machias Bay, 44°40’ BF 13. 6 EA 17.0 Brackish Gravel, mud, boulders 
mouth of Cutler Brook 
M9 July4 South Trescott 44° 46' 67°04’ =LW 955 323 Muddy sand, boulders, 
(Baillie’s Mistake) Ascophyllum 
M10 July4 Straight Bay, Cow 44°52' 67°08’ LW 124 31.8 Reducing mud flats, 
Neck, Cobscook Bay Ulva, Cladophora 
M11 July5  Cobscook Bay, SW 44°49’ 67°10' LWtoHW 13.4 29.6 HW debris and 
arm, 2 miles below bedrock, LW mud, 
Whiting Zostera, Ascophyllum 
M12 July5  Dennys estuary, at 44°56’ 67°14’ oO LW 174A 6.50 Gravel, mud, fucoids, 
hwy bridge, NW Zostera, debris 
Cobscook Bay 
M13 July5 Reversing Falls, 44°53' 67°08' MW 10.5 _ Bedrock, fucoids 
Cobscook Bay 
M14 July 21 Mosquito Head,near 43°56’ 69°13’... HW —_ _ Twin sand beaches 
Martinsville 
M15 July 21 Spruce Head, at 44°01’ 69°08’ HW — _ Drift debris on igneous 
causeway bedrock, gentle slope 
M16 July 21 South Thomaston,at 44°03’ 69°07’ HW _— — Fine igneous sand, 
town beach stones 
M17 July 17 Sand Beach, near 44°20’ 68°11’ HW — — Steep surf-exposed 
Great Head, Mount fine-sheil sand beach 
Desert I. 
M18 July 18 TheLedges,northof 44°19’ 68°11’ HW. ~ — Igneous bedrock spray 
Otter Cliff Point pools 
M19 July 17 Mill Pond, near 44°21' 68°25") “LW to HW -°22.6: S13 Mud, fucoids, stones, 
Prettymarsh Zostera, filamentous 
algae 
M20 July 17  Prettymarsh Pond, 44°21' 68°24’ LWtoHW 246 31.1 Mud, gravel, stones, 
Mount Desert I. Zostera, Enteromorpha 
M21 July 18 Marlboro Beach, 44° 28' 68°17’ LW Soft black muck, 
Raccoon Cove pebbles, Myti/us beds 
and Echiurus 
M22 July 18 Lamoine Beach 44°27' 68217" LW 16.6 31.4 Sand, muddy sand, 
pebbles, stones 
M23 July 19 Off Mount Desert I. 44°26’ 68°18' 15-60 ft _ _ Black reducing mud 
Biological Station, Dredge 
Salisbury Cove, 
near buoy N6 
M24 July 19 Near Googin's Ledge, 44°27’ 68°18" 15-50 ft _ -— Mud, stones, sand 
off Lamoine Dredge 


46 


Northern New England and Western Nova Scotia 


Sta Date Locality Lat N LongW Depth Temp Salinity Habitat 
No. *¢ %o 
M25 July 19 Bayside, mouth of 44°28' 68°26’ LWtoMW 244 26.2 Gravel, boulders, 
Union R. fucoids 
M26 July 21 North Cove, Tenant's 43°58’ 69°12’ LWtoMW 13.6 31.7 Bedrock, silty mud, 
Harbour Ulva, Chondrus 
M27 July 21 1mile NE PortClyde 43°56’ 69°15’ LWtoMW_ 12.0 — Sand, boulders 
M28 July 21 Pemaquid Beach 43°52’ 69°32' LW 17.44.3818 Bedrock, pebbles, 
coarse sand, Chondrus 
M29 July 21 Pemaquid Point 43°50’ 69°31' LWtoHW 16+ Sandstone schist 
bedrock, heavy surf 
M30 July 22 Round Pond, atwharf 43°56’ 69°27’ LWtoMW 16.2 — Silty mud, bedrock, 
Mytilus beds 
M31 July 22 Hockamock Point,S 43°58’ 69°25' LW 15.1. 31.5 Mud, gravel, shells, 
of Medomak, at wharf Zostera, Chondrus 
M32 July 22 Sylvester Pond, 43°59’ 69°25’ LWtoMW 22.3 31.4 Fine mud, stones, 
Greenland Cove filamentous algae 


M33 July 23 Damariscotta R., at 44°02’ 69°32' LWtoHW 22.0 Brackish Boulders, mud, 


town bridge Chondrus 

M34 July 23 Damariscotta Salt 44°01’ 69°32'  Supratidal — Brackish Thick mud, Ruppia, 
Pond, 1 mile above Zostera debris, Spartina 
Newcastle 

M35 July 23 Mouth of Oyster Creek, 44°04’ 69°31' LW — Brackish Shells, mud, Ruppia, 
at Damariscotta Ulva, at bridge 
estuary abutments 

M36 July 23. Hendricks Point Beach, 43°49’ 69°41' LW 17.3) “34;0F Coarse sand, bedrock, 
Boothbay Harbour fucoids, Chondrus 

M37 July 24 Ocean Point, 43°49’ 69°35’ LWtoHW 15.3 Sie Surf-exposed sand- 
E Boothbay stones, schists, 

intrusives 

M38 July 24  Linekin Bay, at 43°52’ 69°35’ LWtoMW_ 18.2 — Gravel, chips, mud 
Paradise Point 

M39 July 24 SheepscotR.,athwy 44°03’ 69°37’ LW 25:3 C1910 Muddy sand, stones, 
bridge, above sill Enteromorpha, Zostera 


M40 July 24 Reid State Park beach 43°47’ 69°43’ LWtoHW 14.8 31.8+ Sandstone bedrock, 
coarse sand, surf 


M41 July 25 Dune Beach, Hermit!. 43°43' 69°51" =LW 16.0 30.9 Schistose bedrock, 
Chondrus, sand, fucoids 
M42 July 25 Popham Beach, at 43°45’ 69°47' MW toHW _ — Coarse to fine sand, 
Fort Popham igneous rock pools 
M43 July 26 Prouts Neck, Saco 43°32’ 70°20' LWand 195 31.4 Graded sands, stones, 
Bay at yacht club subtidal Zostera 
M44 July 26 Cape Elizabeth 43°34’ 70°12" HW 16.2 —_ Surf-exposed pools in 


vertically bedded 
schistose rocks 


M45 July 27 Biddeford Pool, at 43°27' 7O°2A"° LW 16:8" S2.2 Coarse sand, bedrock, 
South Beach fucoids, debris 

M46 July 27 Biddeford Pool, atGut 43°28’ 70°21' LWtoMW 19+ - Graded sands, organic 
entrance sands, stones, Zostera 


M47 July 28 Scarborough estuary, 43°34’ 70°22' LWtoHW 25+ Brackish Coarse gravel, algae, 
at abandoned RR bridge abutments 
culvert 


47 


Geographical and Ecological Data 


Sta Date Locality Lat N Long W Depth Temp Salinity Habitat 
No. (6: %o 
M48 July 28 Pine Point, at 43°33" 70°21’ LWtoMW 22+ _ Graded sands, muddy 
Fisherman's Dock sand, mussel shells 
M49 July 29 Bridge Point, Cousins |. 43°46’ 70°08’ MWtoHW 21.8 — Coarse sand, boulders, 
fucoids 
M50 July 29 Staples Point(Moore 43°48’ 70°06’ LWtoMW 22.2 _ Mud, bedrock, shells, 
Point), at Keeshawa to Zostera, fucoids 
Bay 26.9 
M51 July 31 Cove E of Walker's AS 2" 70°27' LWtoMW 14.8 _ Igneous and volcanic 
Point, Kennebunkport bedrock, Chondrus, 
fucoids 
M52 July 31 Mousam estuary 43° 217 70°31"  LWLHW 17.3 — LW graded sands, HW 
mouth to salt marsh 
24+ 
M53 Aug.1 Wells Inlet,S branch 43°16’ 70°34’ =LW 24+ 31+ Mud flats, cliffs, 
Spartina, fucoids 
M54 Aug.1 Ogunquit SaltMarsh 43°16’ 70°35’ MW toHW — —_ Clean sand, Spartina, 
fucoids 
M55 Aug.1 Cape Neddick Beach, 43°12’ 70°36). = LW 18.8 Brackish Graded sand, gravel, 
at creek mouth mud, salt grasses 
M56 Aug.1 Cape Neddick, at 43°10’ 70°35’  HWWand 16.1 — Granite bedrock, spray 
Nubble Light supratidal pools, under drift 
M57 Aug.2 _ Piscataque estuary,at 43°07’ 70°50' MW to HW — Brackish Mud flats, Spartina 
Hwys 4 and 16 junct., 
N.H. 
M58 Aug.2 _ Piscataque estuary,at 43°08’ 7O°S2". LW 22.8 Brackish Mud, boulders, shell, 
mouth Bellamy R. Zostera, fucoids 
M59 Aug.3 Seabrook beach, at 42°53’ 70°49’ =LW 17.1 a Surf-exposed coarse 
Beckman’s Point to sand 
19.5 
M60 Aug.4  Plum!.Sound,Nside 42°42’ 70°47’ = LW Wa Shs Coarse sand 
of entrance, Mass. 
M61 Aug.4  Plum|. estuary, 42°48’ 70°48’ MW — — Coarse ripple sand, 
at entrance Parker R. mud, Spartina 
refuge 
M62 Aug.5_ Little Harbour,mouth 43°04’ 70°44’ LW 1 52 ee OulEe Muddy sand, boulders, 
of Piscataque R., N.H. fucoids, kelp 
M63 Aug.5 Odiornes Point, oppo- 43°03’ 70°43’ MW toHW — — Hard bedrock, stones, 
site Isles of Shoals pools, debris 
M64 Aug.5 Castle Neckestuary, 42°40’ 70°43) LW 18.1 - Fine white sand, 
E side of mouth, Mass. boulders 
M65 Aug.5 _ Essequit salt marsh, 42°39' 10 As — — Mud, Spartina, 
inside Wingarsheak Ascophyllum var. 
Beach 
M66 Aug.6- Rockport, town beach 42°40’ 70° S37? LWtoMW 15.4 31.7 Variable sand, boulders, 
bedrock, fucoids 
M67 Aug.6  Essequit Channel, 42737! 70°40' MWtoHW 18.6 _ Mud banks, Mytilus 
inside Gloucester beds, Ascophyllum var., 
entrance Spartina 
M68 Aug. 7 Magnolia Beach, A235. 70°43' LWtoMW 14.6 — Fine sand, boulders, 
Cape Ann stones, bedrock, 
fucoids 


48 


Northern New England and Western Nova Scotia 


Sta 
No. 


M69 


M70 
M71 


M72 


M73 


M74 


M75 


M76 


M77 


M78 


Aug. 


Aug. 


Aug. 


Aug. 


Aug. 


Aug. 


Aug. 


10 


4 


12 


Locality 


Annisquam Pond, 
Cape Ann 


Folly Bay, Cape Ann 


Marshfield salt pond, 
W of Brant Rock at 
hwy gates 


Brant Rock, trailer 
camp shore 


Duxbury Bay, at 
causeway to beach 


Warren Cove, at 
Beach Park 


Eel Creek, upper 
estuary, below head 
of tide 


Cape Cod Bay, S side 
of canal mouth 


East Sandwich Beach, 
at inlet mouth 


North WVeymouth, 
S of Boston 


Lat N 

42°39' 
42°42! 
42°05’ 
42°05! 
42°03’ 
41°56’ 


41°56! 


41°47! 


41°45! 


42°15’ 


Long W 


70°39’ 


70°37’ 


70°38’ 


70°37’ 


70°38" 


70°37" 


70°37" 


10°30’ 


70°26" 


FOLD T* 


Depth 


MW to HW 


HW 


LW 


LW 


LW 


LW 


MW to HW 


LW to HW 


LW 


LW 


16 + 


Zar 


15.0 


25 


15.6 


19.4 
to 
23+ 


20.4 


Salinity 


%o 


Brackish 


Fo | 


32.4 


Brackish 


32.0 


Habitat 


Mud flat, eelgrass 
debris 


Igneous bedrock pools 


Mud banks, Spartina, 
muddy gravel, fucoids 


Bedrock, boulders, 
gravel, sand, Chorda 


Mud, coarse sand, 
eelgrass, stones 


Fine sand, organic and 
muddy sand 


Mud, gravel, Spartina, 
10-15 cfs. 


Coarse sand, stones, 
Chorda, Zostera, 
fucoids, HW sand 


Sand, pebbles, gravel 
riffle, debris 


Sandy silt, stones, 
Chonarus, Ulva 


49 


Literature Cited 


Abbott, R.T. 

(1968). Sea shells of North America; a guide to 
field identification. Golden Press, New York. 
280 pp. 


Bourget, E. 

(1971). Aspects saisonniers de la fixation de 
l‘epifaune benthique de |'étage infralittoral de 
l‘estuaire du St-Laurent. M.Sc. thesis, Laval Univ. 
110 pp. 


Bousfield, E.L. 
(1952). Zoological investigations in the Maritime 
Provinces. Nat. Mus. Can. Bull. 126: 188-94. 


(1954). The distribution and spawning seasons of 
barnacles on the Atlantic coast of Canada. Nat. 
Mus. Can. Bull. 132: 112-54. 


(1955a). Viviparus viviparus L. in eastern Canada. 
Can. Field-Nat. 69: 27-28. 


(19556). Some physical features of the Miramichi 
estuary. J. Fish. Res. Board Can. 12(3): 342-61. 


(1955c). Ecological control of the occurrence of 
barnacles in the Miramichi estuary. Nat. Mus. Can. 
Bull. 137: 1-69. 


(1956a). Studies on the shore fauna of the St. 
Lawrence estuary and Gaspé coast. Nat. Mus. Can. 
Bull. 136: 95-101. 


(19566). Studies on the shore Crustacea collected 
in eastern Nova Scotia and Newfoundland, 1954. 
Nat. Mus. Can. Bull. 142: 127-52. 


(1956c). Malacostracan crustaceans from the 
shores of western Nova Scotia. Proc. Nova Scotian 
Inst. Sci. 24(1): 25-38. 


(1958a). Fresh-water amphipod crustaceans of 
glaciated North America. Can. Field-Nat. 72: 55- 
Ta ee 


(19585). Littoral marine arthropods and mollusks 
collected in western Nova Scotia, 1956. Proc. 
Nova Scotian Inst. Sci. 24(3): 303-25. 


(1958c). Distributional ecology of terrestrial Talit- 
ridae (Crustacea: Amphipoda) of Canada. Proc. 
Tenth Int. Congr. Entomol. 1: 883-98. 


(1960). Canadian Atlantic sea shells. National Mu- 
seums of Canada, Ottawa. 72 pp. 


(1962a). Studies on littoral marine arthropods 
from the Bay of Fundy region. Nat. Mus. Can. Bull. 
183: 42-62. 


(19626). New haustoriid amphipods from the Ca- 
nadian Atlantic region. Nat. Mus. Can. Bull. 183: 
63-75. 


(1964). Coquillages des cdtes canadiennes de 
|‘Atlantique. National Museums of Canada, Ot- 
tawa. 89 pp. 


(In press). Shallow-water gammaridean Amphi- 
poda of New England. Cornell Univ. Press. 340 pp. 


50 


Bousfield, E.L., and A.H. Leim 
(1960). The fauna of Minas Basin and Minas 
Channel. Nat. Mus. Can. Bull. 166: 1-30. 


Bousfield, E.L., and D.E. McAllister 

(1962). Station list of the National Museum Ma- 
rine Biological Expedition to southeastern Alaska 
and Prince William Sound. Nat. Mus. Can. Bull. 
183: 76-103. 


Bousfield, E.L., and M.L.H. Thomas 

(In press). Post-glacial dispersal of littoral marine 
invertebrates of the Canadian Atlantic region. 
J. Fish. Res. Board Can. 


Brunel, P. 

(1970). Catalogue d’invertébrés benthiques du 
Golfe Saint-Laurent recueillis de 1951 4 1966 par 
la Station de Biologie marine de Grande-Riviére. 
Trav. Péch. Québec 32: 1-54. 


Dunbar, M.J., and E.H. Grainger 

(1952). Station list of the ‘‘Calanus’’ expeditions, 
1947-1950. J. Fish. Res. Board Can. 9(2): 65- 
82. 


Lacroix, Guy 

(1967). Recherches sur le zooplancton de la Baie- 
des-Chaleurs. Rapp. Annu. Sta. Biol. Mar. Grande- 
Riviére 1966: 37-53. 


Laubitz, D.R. 

(1972). The Caprellidae (Crustacea: Amphipoda) 
of Atlantic and Arctic Canada. Nat. Mus. Can. 
Pubs. Biol. Oceanogr. 4: 1-82. 


Lavoie, R. 

(1970). Contribution a la biologie et a |’écologie 
de Macoma balthica L. de |’estuaire du Saint-Lau- 
rent. Ph.D. dissertation, Laval Univ. 249 pp. 


Lemos de Castro, A. 

(1965). On the systematics of the genus Litto- 
rophiloscia Hatch (lsopoda, Oniscidae). Arg. Mus. 
Nac. Brazil. 53: 85-98. 


McCain, J.C. 
(1968). The Caprellidae (Crustacea: Amphipoda) 
of the western North Atlantic. U.S. Nat. Mus. Bull. 
278: 1-147. 


Medcof, J.C., A.H. Clarke and J.S. Erskine 
(1965). Ancient Canadian east-coast oyster and 
quahaug shells. J. Fish. Res. Board Can. 22(2): 
631-34. 


Medcof, J.C., and M.L.H. Thomas 

(1969). Canadian Atlantic oyster drills (Urosa/- 
pinx)-distribution and industrial importance. 
J.Fish. Res. Board Can. 26(5): 1121-31. 


Mills, E.L. 

(1963). A new species of Ampelisca (Crustacea: 
Amphipoda) from eastern North America, with 
notes on other species of the genus. Can. J. Zool. 
41: 971-89. 


(1964). Noteworthy Amphipoda (Crustacea) in the 
collection of the Yale Peabody Museum. Posti/la 
79: 1-41. 


Literature Cited 


(1967). The biology of an ampeliscid amphipod 
crustacean sibling pair. J. Fish. Res. Board Can. 
24(2): 305-55. 


Pettibone, M.H. 

(1963a). Revision of some genera of polychaete 
worms of the Family Spionidae, including the de- 
scription of a new species of Sco/e/epis. Proc. Biol. 
Soc. Wash. 76: 89-104. 


(19636). Marine polychaete worms of the New 
England region. |. Families Aphroditidae through 
Trochochaetidae. U.S. Nat. Mus. Bull. 227(1): 
1-356. 


Powell, N.A. 

(1968). Studies on Bryozoa (Polyzoa) of the Bay of 
Fundy region, Il: Bryozoa from fifty fathoms. Can. 
Biol. Mar. 9(3): 247-59. 


Stephenson, T.A., and Anne Stephenson 
(1954). Life between tide-marks in North America, 
II1A: Nova Scotia and Prince Edward Island. J. 
Eco/. 42(1): 14-45. 


Tattersall, O.S. 

(1954). Shallow-water Mysidacea from the St. 
Lawrence estuary, eastern Canada. Can. Fie/d-Nat. 
68: 143-54. 


51 


ol a eee ete ar yy wou |} = rl - 
ri vi Aj - vi My Dans + 
@ ae | ' 
i 34 ' ? 
4 A ( 
a - wy iv 
; ian / 1 
‘TP 
: in 
| ‘ ne ' 
or ; “ r os - ¥) 7 
j i ' 
ee ee Se oT ee ee ae ba Ae 
- P 1 ve ‘ i» wy : f ar = tow ats ay! Wi 
Wend ae, ‘ 4 pak +a ae 
Ang , , i. a OUR 
; +e 
ey ae i, » 7 
‘ ‘ Pipe: T ; sult 1 port 
4 P) : Ay! pues tt ' ; . 
' , NS } 
F 7 4 ’ | 7 
4 
b go é 
Neel py » 
; j b 
' a | oT a j pe 
8 
— é 
— Py, 


»4/ pes 
‘ j j 
a R - 
ye ? =" x 5 J, 
ee | ‘ 
? <) 
mu io. a “ , - 
7 37 
ay! Pail i< 
: re oe ey 7 ’ 
| ee Le ra 
a ) , 
al oe p ‘ ‘ o % 
2 ies : 
; ( Wip Cae ae = : 
Si Re a ‘5 a \ - = 
pie ~ | © sie id 
\) a . 28 
ad d q , ww f 
‘ y 4 ¥ 
i - 
oh @ . 
a 
y i U " 
2 * f % 
(eras eS 
1 
\ 
_ v/ a = 
PAWS ' j 
t 
4 
) ew ny 
YA \Ap ig iy 4 
».9 
rw). » } 
i ed 7] Lal i /F g , x 
oe hal ¥ J tia Gig ne ad 7 f 
oF 44 
PL (eviews ee lv ‘ * 
‘ ° q 
" ¢ aii if » 2, * , pat 
oe 8 " 
Pei Sy: ii ie \ : 5 tes, OM) 
u -— 
af o veut yur a = (s oi ix? Hs ill F\'s a : ' éu f ke TS : ar | y 
‘ri PAE ae, Sted a Bae.) 
i : i F 


} 


CALIF ACAD OF SCIENCES LIBRARY 


3 1853 10004 6478