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CONTRIBUTIONS TOWARD A FLORA OF NEVADA 


NO, 37 ) | 


SCROPHULARIACEAE OF JEVADA « 
en te 
by” 

GABRIEL) EDWIN 
June 27, 1955 


A series prepared through the cooperation of 
the National Arboretum and the Section of Plant Introduction 
Horticultural Crops Research Branch 
Agricultural Research Service. 
U. S. Department of Agriculture 
Plant Industry Station 


Beltsville, Md. 


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Address all inquiries concerning this series 
to W. Andrew Archer, Plant Industry Station, 
Beltsville, Maryland 


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SCROPHULARIACEAE OF NEVADA 
PART I - MIMULUS 
By Gabriel Edwin * 


Contribution Toward a Flora of Nevada, No. 37 


MEMULUS°L.; Spt Pl. 2.°1753. 

Annuals or perennials; in Nevada herbs; glabrous or pubescent; 
leaves mostly opposite, occasionally in a basal rosette or scattered, 
sessile or petioled, entire, toothed or lobed; calyx with sepals at 
least partly united, teeth equal, subsqual or unequal. Corolla equally 
or unequally bilabiate, yellow to blue, red, reddish purple, sometimes 
white; tube included or exserted: upper lip 2-lobed, lower 3-lobed, 
mostly exceeding the upper, often with 2 hairy patches extending into 
the throat; stamens ), didynamous, filaments adnate to the corolla, 
anthers equally 2-celled, confluent at the top, sometimes exserted; 
Stigma lobes distinct, style simple; capsule oblong or ovoid, 2-valved, 
loculicidally dehiscent; seeds mostly numerous, ovoid, ellipsoid or 
subrotund, usually reticulate, rarely smooth. 


Typs species: M. ringens L., Sp. Pl. 63h. 1753. 


SYNOPSIS OF SUBGENERA AND SECTIONS 
Subgenus I. Synplacus Grant. Placentae completely united into a cen- 


tral coiumn or splitting less than half way to the base; pedicels 


% Assistant Curator, National Arboretum Herbarium, Beltsville, Maryland 


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usually longer than the calyx; style glabrous or puberulent, rarely 

pubescent; capsule mostly dehiscent to the base along both sutures; 

corolla dropping soon after anthesis (except in M. breweri). 

1. Calyx teeth equal or subequal, not folding over the calyx orifice 
at maturity; palate, when present, not prominent, not constricting, 
nor even partly closing the corolla throat. 
2. Stamens exserted; calyx teeth -6 mm. long. . ERYTHRANTHE (A) 
2. Stamens mostly included, when exserted the calyx teeth not ex- 

ceeding 3 mm. in length ... « «+ « » « «.« « « PARANDANTHUS (B) 

1. Calyx teeth very unequal, folding over the orifice at maturity; 
palate always present, well-developed, frequently at least partly 
etouiue tip corollas throat «5 + + 6 one) ¥u tee v,& SIMLOLUS, (C) 

Subgenus II. Schizoplacus Grant. Placentae mostly splitting to the 

base and adherent to the valves; pedicels usually shorter than the 

calyx; style glandular-pubescent; capsule mostly dehiscent to the base 
along the inner suture and part way or not all along the outer; ea 
la persisting on the developing capsule. 


Corolla tubular-funnelform to nearly campanulate, style glandular- 


pubescent, at least along the upper half .. . . EUNANUS (D) 


(A) ERYTHRANTHE Greene 
1. Corolla strongly bilabiate, tube only very slightly exserteds; stigma 
DS CE er 6) Hous oh en tHe int uiien a @ 4 le Me CARDINALIS 
1. Corolla only slightly bilabiate, tube much exserted; stigma lips 


eciliate e e e e e * e e ° ° e . * . ° ° ° . e 2 e M. EASTWOODIAE 


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(B) PARADANTHUS Grant 
1. Corolla lobes distinctly unequal. 
2. Mature calyx distinctly inflated; leaves ovate, dentate. 
3. M. FLORIBUNDUS 
2. Mature calyx little or not at all inflated, sometimes distended 
by the mature capsule; leaves linear or linear-lanceolate, en- 
Ene sw el ss os le «ar so 0 We MS MONTTOUDES 
1. Corolla lobes equal or subequal. 
3. Anthers bearded (rarely glabrous in M. primuloides); perennial. 

). Flowers mostly solitary, never racemose; stems often scapose. 

5. M. PRIMULOIDES 
4. Flowers racemose; stems leafy. 
5. Stems glandular-pubescent; *orolla red or pinks; calyx 
Resch) s-OUNM, LOWE ss «<6 6 «+ « « O» M. DEWISTE 
5. Stems with soft, long pubescence, slimy to the touch; 
corolla yellow; calyx teeth 1.5-3 mm. long. 
7. M. MOSCHATUS 
3. Anthers glabrous; annuals. 

6. Clayx teeth arising from the tube at different levels, tube 
deeply cleft with sinuses of various lengths, lowest teeth 
about equaling the length of the tube; one pair of stamens 
often reduced in length, but remaining functional. 

8. M. PILOSUS 

6. Calyx teeth all arising from the tube at the same level, 
not exceeding 1/3 the size of the tube, sinuses equal in 
length or almost so; both pairs of stamens fully developed 


in length. 


4 


7. Plants glandular-pubescent or giandular-villous; corolla 
persisting on the developing capsule; stems slender, 2-15 
CMe Uee oes Se ee ee ew 8s 9. M. BREWERI 

7. Plants glabrous or minutely puberulents; corolla failing 
soon after anthesis, plants usually not exceeding 8 cm. 
8. Stigma lobes unequal; calyx teeth eciliate. 

10. M. SUKSDORFII 
8. Stigma lobes equal; calyx teeth ciliate. 


11. M. RUBELLUS 


(C) SIMIOLUS Greene 
1. Corolla throat open; flowers axillary, or in a few-flowered raceme, 
Bear eria. Cae. SUMMIb.. » o. s,s. + « «© « « » « les M. GLABRATUS 
1. Corolla throat partly, or nearly closed by the prominent palate; 
flowers commonly numerous, in a definite raceme, sometimes few- 
flowered. 
2. Upper calyx tooth rarely more than twice the length of the 
others. 
3. Upper leaves conspicuously connate-perfoliate, glaucous or 
BaUCEGENL ss 5 6 2 6 6 he et ee ee oe ol. M. GLAUCESCENS 
5. Upper leaves neither connate-perfoliate nor glaucous, seldom 
pubescent. 
4. Style much shorter than the smaller pair of stamens; leaves 
usually broader than long; corolla very little exceeding 
Wiemeairm ee gs 6 + ee «ae sw Ly. M. BRACHYSTYLIS 


4. Style equaling or longer than the stamens; leaves usually 


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longer than broad; corolla usually well-exserted. 
5. Flowers mostly in definite racemes; pedicels 
usually shorter than the corollas; rootstocks 
rarely fleshy or yellow. . . 15. M. GUTTATUS 
5. Flowers mostly 1-3, rarely 5 on a stem; pedi- 
cels usually longer than the corollas; stems | 
arising from a mass of fleshy, yellowish root- 
BLOCKS. es e- 8 «© vows o) 16, MB. TILINGT 
2. Upper calyx tooth almost 3 times the length of the others. 


17. M. NASUTUS 


(D) EUNANUS Gray 
1. Mature calyx distinctly inflated. 
2. Corolla manifestly bilabiate. 
3. Calyx throat oblique, teeth unequal. 
h. Calyx teeth mostly blunt. .... . ». 18. M. TORREYI 
. Calyx teeth triangular or lanceolate, acute. 
19. M. ANGUSTIFOLIUS 
3. Calyx throat slightly, if at all oblique, teeth nearly equal. 
20. M. NANUS 
2. Corolla not obviously bilabiate, the lobes equal or nearly so. 
5. Anthers glabrous; stigma lobes unequal. 21. M. SPISSUS 
5. Anthers hispid; stigma lobes equal. .. 22. M. BIGELOVII 
1. Mature calyx little, if at all inflated, sometimes distended by 
the mature capsule. 


7. Stamens mostly included, only the upper pair sometimes slightly 


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8, Calyx teeth acute, about 1/l as long as the tube; capsule 
ovate-lanceolate, usually well-exserted. 
9. Calyx 6-10 mm. long; plant not noticeably mephitic. 
23. M. DENSUS 
9. Calyx -6 mm. long; plant when fresh, strongly mephitic. 
24. M. MEPHITICUS 
8. Calyx teeth lanceolate-subulate, at least 1/3 as long as 
the tube; capsule broadly oblong or ovate, scarcely exserted. 
10. Stamens included; calyx 3-5 mm. long, teeth unequal; 
corolla tube 2 to 3 times the length of the calyx. 
25. M. COCCINEUS 
10. Upper pair of stamens little exserted; calyx 0.8-1.1 
em. long, teeth subequal; corolla tube 14 to 2 times 
the length of the calyx... ... 26. M. WASHOENSIS 


7. Both pairs of stamens distinctly exserted . . 27. M. STAMINEUS 


1. MIMULUS CARDINALIS Dougl. in Benth., Sercph. Ind. 28. 1835. 

Erythranthe cardinalis Spach.; Mimulus cardinalis var. rigens 
Greene; M. cardinalis var. griseus Greene; M. cardinalis var. exsul 
Greene. 

Perennial; erect to procumbent, villous or villous-viscid; stems 
from a running rootstock up to about 30 cm.$; leaves obovate, oblong 
or lance-obovate, 2-12 cm. long, 1-5 cm. wide, sessile, auriculate, 
serrate with Salient teeth; pedicels usually exceeding the subtending 


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Dry or damp places. Douglas, Mineral and Ormsby Counties. 


2, MIMULUS EASTWOODIAE Rydb., Bull. Torr. Bot. Club 0: 83. 1913. 
M. cardinalis Hastw. 

Perennial by stolons or rootstocks, procumbent, usually strong- 
ly woolly-canescent; stems terete, short; leaves varying from obovate 
to oblong, with acute tips, 2-6 cm. long, 0.5-3 cm. wide, sessile, 
coarsely dentate along the upper half, lower leaves usually reddish 
below, with cuneate bases; pedicels exceeding subtending leaves; 
flowers few. 

Wet shaded places. Reported in Nevada by Wheeler's Expedition, 


1872. 


3. MIMULUS FLORIBUNDUS Dougl., Bot. Reg. 13, pl. 1125. 1828. 
M. peduncularis Dougl.; M. serotinus Suksd.; M. floribundus 
var. minor Hook.; M. pubescens Benth.; M. deltoideus Gdgr. 

Annual; viscid-villous, more or less slimy plants; stem branch- 
ed from the base, procumbent or weakly erect, up to about 50 em. high; 
leaves mostly winged petiolate (occasionally sessile), scattered, 
thin ovate, obovate or ovate-lanceolate, 1.5~5 cm. long, 0.5-3 cm. 
wide, acute, saliently dentate, ciliate, base broadly subcordate, 
sometimes cuneate, 3-5-nerved from the base; pedicels filiform, mostly 
spreading at maturity, frequently shorter than the subtending leaves. 

Mostly dry places. 

Churchill, Douglas, Elko, Lincoln, Mineral, Nye and Washoe 


Counties. 


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4. MIMULUS MONTIOIDES Gray, Proc. Am. Acad. 7: 380. 1867. 
M. rubellus var. latiflorus Wats.; M. rubellus Gray (in part). 
Annual; erect, simple or branched, viscid-puberulent to gland- 
ular-pubescent plant; leaves linear to linear-lanceolate, entire, ob- 
tuse, 1.5-2.5 cm. long, 2-8 mm. broad, upper sessile, lower usually 
short-petiolate; pedicels 0.5-3 em. long. 
Dry open places. 


Douglas, Nye, Ormsby and Washoe Connties. 


5. MIMULUS PRIMULOIDES Benth., Scroph. Ind. 29. 1835. 
M. pilosellus Greene; M. nevadensis Gdgr.3 M. prinuloides 
var. pilosellus (Greene) Smiley. 

Perennial; glabrous to viscid-villous or puberuient, producing 
stolons, runners or bulbils, scapose or with true stem leaves; flowers 
often solitary or subsolitary; leaves broadly ovate to oveate-lanceo- 
late, occasionally cuneate, always sessile. (Having the habit of the 
genus Primula). 

Mostly in moist to wet (occasionally dry), open places, 6000- 
11,060 ft. 

Douglas, Elko, Humboldt, Mineral, Nye, rvmsby and Washoe 


Counties. 


6. MIMULUS LEWISII Pursh, Fl. Am. Sept.” 220 27,: pl. e200"" 182), 
M. lewisii var. exsertus Coult. & Fisher; M. roseus Dougl.; 


M. roseus var. glabrior Hook.; M. lewisii var. tetonensis A. Nels.; 


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M. lewisii var. alba Henry; M. lewisii f. tetonensis (Nels. ) Macb. 
& Pays. 

Perennial from a running rootstock; stems erect, 3-8 dm. tall, 
usually unbranched, mostly viscid-pubescent, sometimes viscid-villous; 
leaves thin, oblong-lanceolate to ovate-oblong, 2-8 cm. long, 1-2.5 
em. wide, 3-5-nerved from *he base, sessile, entire to irregularly 
dentate with short blunt teeth; pedicels mostly exceeding the sub- 
tending leaves. Superficially much like the species of section 
Erythranthe. 

Mostly damp and wet, exposed places, 6000-9100 ft. 


Clark, Douglas, Elko, Ormsby, Storey and Washoe Counties. 


7. MIMULUS MOSCHATUS Dougl., Bot. Reg. 13, pl. 1118. 1828. 

Perennial; rootstocks; stem weak, creeping or decumbent, up to 
33 em. long; entire plant viscid-villous and sometimes slimy to the 
touch; leaves broadly or narrowly ovate, 1-) cm. long, 0.5-2.5 em. 
wide, acute or obtuse, mostly denticulate, sometimes entire, short- 
petiolate, pinnately veined; flowers few; pedicels shorter than the 
subtending leaves. 

This plant is perhaps much more variable than is at present 
indicated. Specimens with equal calyx teeth, different pubescence 
and with other variations in character have been noted that are 
either close to or synonymous with this entity. However, in all 
cases the material was incomplete and no definite conclusions could 
be drawn. 

Damp, wet or rarely dry, open or shaded places. 


Douglas, Elko and Washoe Counties. 


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8. MIMULUS PILOSUS (Benth.) Wats., Bot. King's Exp. 225. 1871. 
Mimetanthe pilosa (Benth.) Greene; Mimulus exilis Dur. & Hilg. 
Annual; stem villous, erect, branched or unbranched; leaves 
shorter than the pedicels. 
This species is easily recognized due to the very unusual calyx 
lobes and the often reduced stamens. 
Wet, damp or dry places, about 5000-6500 ft. 


Churchill, Elko, Humboldt, Lander and Washoe Counties. 


9. MIMULUS BREWERI (Greene) Coville, Contr. U. S. Nat. Herb. ): 171. 
1893. 

Annual; simple or branched, up to 17 cm. high, glandular-pubes- 
cent or puberulent; leaves linear to oblong, small, up to 1.8 cm. long, 
h mm. wide, weakly toothed or entire, mostly obtuse, short-petiolate 
or usually sessile and always nerved from the base; pedicels up to l 
cm. long, mostly exceeding the calyx. 

Mostly damp, exposed places, 8000-11,000 ft. 


Elko, Nye and Washoe Counties. 


10. MIMULUS SUKSDORFII Gray, Syn. Fl. N. Am. ed. 2, 2, pt. 1, suppl. 
450. 1886, 

Annual; low bushy, much-branched plants, up to about 6 cm. tall, 
with sparse, viscid puberulence and a red tinge; leaves mostly linear 
or almost so, sometimes oblanceolate-linear, not more than 2 mm. wide, 
12-1) mm. long; obtuse, with small, sinuate teeth or subentire, mostly 


sessile, the lowest occasionally short-petiolate. 


ee ia88) ome a! aoe 108, F 
- Bernal - rhenedentian hy ni m0 beta 


eeflss Iaptuar (tev @ 


“ot oo%8 0008 5 ee soca — 10 ob et 


FYE sl .dvoli ite . 0.0. paused. gall V0 (once roe an 
t : & 


=< 


~wodvatalube faa "9 


£, — 


TH... .M9 Sf Ok Oty Spm 


ag | 
t 


7 a : eT 
steloiteq=trode ,enudtlty: cred 62 suis Id. basttone eile veoh ot 


& % 


o 


Loge oe alestheag p,agud ‘one gor bevien sree oe eiteass vitensi 
inyias ent ; sy:toeeoKs. Yliwon gaol: 3 

: ; rae “ — a ; 

($3 000, 12eh008. 4 goveta besoqis <8 Et POM 


ag. ttntod: aor baw oUt on 


f° 


.2eae, ORL 
fied: io O ducds oF ae: re: sitnat ¢ bvonion-ed coum enelasse wor 


ysenkl yiseom sevect pogait bee rs bre souatisrrocig’ bloat PE bg : 


| 
ee 


ial a ae 
LeRom au dooce. ag: ittesd: odes. tiem thiw penddo % 


fio 
Yh ieee “ee 


eb Me mt Sorand: etoer ter cseontSuadatosoaaid contsonon 


Lage 


Very close to M. rubellus Gray but the stigma lobes are clearly 
different. However, M. breweri (Greene) Coville, M. suksdorfii Gray 
and M. rubellus Gray present a confusing problem. A rather intensive 
study (which is beyond the scope of this paper) seems necessary to 
establish, if possible, the true limits and relationship of these 
threes taxa, assuming that three taxa do exist. 

Dry or wet places, ),000-11,300 ft. 

Douglas, Elko, Esmeralda, Humboldt, Mineral, Nye, Washoe and 


White Pine Counties. 


11. MIMULUS RUBELLUS Gray ex Torr., Bot. Mex. Bound. 116. 1859. 
M. gratioloides Rydb. 

Annual; stems reaching 20 cm. , simple or branched; entire plant 
often beset with glandular pubescence but sometimes glabrous; leaves 
varying from linear to oblong, often tinged with red, up to 2 cm. 
long, and 5 mm. broad, mostly smaller and narrower, sessile, or the 
lowest sometimes short-petiolate, the margins mostly with irregular, 
short teeth, 1-3-nerved from the base. 

Dry; sometimes shaded places, 3000 ft. upwards. 


Elko, Esmeralda, Lincoln, Nye, Washoe and White Pine Counties. 


12. MIMULUS GLABRATUS HBK., Nov. Gen. et Spe 29 370. 1617. 
M. andicolus HBK.; M. jamesii var. texensis Gray; M. tener Phil. 
Perennial, glabrous or almost so; stems stoloniferous or creep- 


ing, rooting at the nodes, branched, up to ) cm. tall; leaves broadly 


Lara 


Pip aag 


an ae re i en 


smd £ fei top opeiue ae at Etvoo (onse30) goxesd x 


ovisnsta saute aS " amrony, paddies a ‘Yuoseag wa Es 


uimeie wie nado aagte od snd : . ip. ET ie gine N38 . 


Or. 2 aR 9995 ‘emege: (roms, ssa to oqooe pit eqoyed, ak ee ¥, ng ; 
ere ‘9 qiiteaotoates bigies. ‘aati nas ott eldtescq 4 seataabes : 

Lin bes ob Bunt seni? dais abate cexat ‘Be | 

ft! Ge, fx-0001 pavaly dew 40 ‘ao 

bee eofesw ,oyll sianaet ot fe sn ot eb lasoned ours ashanod™ 

| aw BS ‘sastinse9 ont oak e 


~ort ,tgaed .feM «FOR. aiid hit can 8 a ~— s 


O88 
jeslg aritne p5adsgatd to ‘olgete age ) ds galdosoy ame $8 Lh 


ey 


gevacl ;adoudsin som atdorce tud conevecity iatebrnly oh toved nade 5 


WO 3 Os a wat atte begat teste. “8 sGae tde oe sean? 3 Eos! gat 


* 


att so ,olisnes aoner esas Ett ‘tied Lp «it fom teamed 1 é bite va 


—nfegerct dtimoyioeom akg ies ETC ete lobteq- torte cembboms zs wads) 


aead eit gost parent idee: % 


< Rew 60 OYE uae bohsris cenltonoe. et) 


5 PERE eters Wd 


wot ext edt sw Brin suinaW ,eet watasabl. aatevoned outa m 


WESl OTE a8 .oB do aad eH i arama Es 


oP 


7 s aN 
rot «Mg went’ ebecesco! aw Toe enst, om RE, gut autzodbn tp il lin 


aueaire “tb auorotine tote awots 108 agoats 0 snordaly Ts: 


Lied <6 


vy 
7 
fg 


yibsocd eavest {i fad amo Gd ae peadonged c2sbon oft 


ee i) 


a 


=e 


ovate, subcordate, irregularly dentate, 2-5 cm. long, 1-3 em. wide, 
upper sessile, lower petiolate, cuneate; petioles longer or shorter 


than the blades. 


KEY TO VARIETIES 
Leaves mostly broadly ovate, irregular and strongly dentate; pedicels 
and calyx mostly glabrous . ......-. +... 12a. M. GLABRATUS 
var. GLABRATUS 
Leaves mostly suborbicular, oval with a rounded apex, entire undulate, 
sparsely dentate or denticulate; pedicels and calyx mostly pubescent. 
12 b. M. GLABRATUS 


var. FREMONTII 


12 a. MIMULUS GLABRATUS var. GLABRATUS 


Aquatic or semi-aquatic. Elko and Lyon Counties. 


12 b. MIMULUS GLABRATUS var. FREMONTII (Benth.) Grant, Ann. Mo. Bot. 
Gard. 11: 190. 192). 


Washoe Co. 


13. MIMULUS GLAUCESCENS Greene, Bull. Calif. Acad. Sei. 1: 113. 1885. 
Annual; erect, glabrous or glaucous; stems terete, simple or 
branched, occasionally h-angled above, up to 0 cm.; lower leaves 
petiolate, upper few conspicuously connate-perfoliate and usually 
glaucous, margins subentire with short scattered teeth; pedicels about 


as long as the mature calyx. 


ae i 
ee 


a 
ie 
Soe 
Pa 
a5 
eS 
5 


, Setvode 10 tegnot selolead redaenio tage eq 


ears 0 OT mi 1, 
nab wlenerde baie. te: csiugerrt ouEro: xibaor 


TPASTATY 1 os GPs. ee ee eee ee anorde ly sutton 


CUT ARHAI .tav- > a - ly 
oo ee : | 


3 alt 


eseinbay eridne .xeqe Debiven s° -asEw eee: alata videon & 

-tqsoescug yYlizor xylso bas & Teotbeg: povalwetinge * 0 ‘otathieh vss 
BUTATEAIO «Mid SL 
TETMOMENT Ate 


QUTATGAIO <xev QUTATEAID Cree 


ay i ine 


eo 


eetengen com. bror cml fie " oitaups-tmee O° okveupa 


eHSel GEL 4 {t had 2 
8 
“AGL (EEE 2h kee .bsot obited Lind sie eutgeaouato eu: 


+ 


ro elquie ,eveTed amade gesoonals 10 “swordely i 


savesl wawot- = ito on OF, cy Rahiaan: 


ods slenkbeq edited boredseoe gtosde i ste 


Dry or damp places. Humboldt and Lander Counties. 


lb. MIMULUS BRACHYSTYLIS Edwin, L. West. Bot. 73 138-139. 195k. 

Annual; stem glabrous, upright, simple or little-branched from 
the base, l-angled, up to 22 em.; leaves ovate, suborbicular or 
rotund with reniform bases, up to 1.6 em. long, mostly as broad as’ 
or broader than long, subentire or weakly and irregularly dentate, 
lower long-petiolate, upper subsessile, uppexmost sessile, more or 
less villous; pedicels longer above, shorter below than ‘the sub- 
tending leaves and calyxes. 


Elko and Nye Counties, 5600-7000 ft. 


15. MIMULUS GUTTATUS DC., Cat. Hort. Monsp. 127. 1813. 

M. Jangsdorfii J. Donn; M. luteus J. Donn, not L.; M. lyratus 
Benth.; M. rivularis Benth.; M. scouleri Hook.; M. luteus var. graci- 
iis Gray; M. glabratus var. ascendens Gray; M. guttatus war. grandis 
Greene; M. langsdorfii var. platyphyllus Greene; M. langsdorfii var. 
argutus Greene; M. langsdorfii var. grandis Greene; M. langsdorfii 
ver. californicum Jepson; M. langsdorfii var. guttatus Jepson; M. — 
hirsutus Howells; M. grandiflerus Howell; M. grandis Heller; M. equinus 
Greene; M. pamiculatus Greene; M. prionophyllus Greene; M. clementinus 
Greene. 

Annual, cr perennial by stolons, creeping rootstocks or stems 
rooting at lower nodes; glabrous or pubescent; stems terete or };-angled, 


fistulous and erect or soft and reclining, up to 55 dm., mostly simple 


Leottaned mebmaed bre thladeo 3808. 


4 ‘ et! 


SRE ee E88 AY dod det” cL path ba BETTTETI Ae SEMIN: 
sox? barlonexd-plaeht so atgats diigo puvidelg mode wlawmak | 
kde pica ci ad cosara ‘néveot (ome 98 0d ga bolans-d onad. “a 


‘ee bacid 2a vltaon egnot «nt Ore ‘od wy  aeaed vero eben. atin) 
-atadaed els. fuporrk bree einen seco) extinedva, neat deus a 0 
~~ PTO eitenas seomenagps albessedue ‘requ otelottequgeoE owl 

mihgs ails ¢ nacd woled aeozads wevods. stogivod sieathed jewel Eby. oaet 


eexglee bas ‘apvanl: 3 ee : 
“ot 0 1 ono pe aeleabd ex bas ent” 


oka 


efOL VSL nen tol FBI » okt arr any TOM 


2.2 fon sarod % asedal M youl 1% bE rsot 


» ‘er 
Ry eh te gal % 
mccbmcdtomphase an 


wtoavs .sav guedat oe i vaioeHl: selaoes of ed bee! rete 
wibitete . TET audedia cry HE eget aiehnoons cov emterdely .M § 
“ev Eitzabeural seneovi ent iydayiata av Picks : 


aceeaUeee 


Pa ov ‘ ~ aad ; 7 f BS 
ittwobeysel 6 gated eice'ts See idtsabe 


TT 


_.K qnvsqet gubsioug . tee Livsobe eae t M Hanae | 


euntvpe .M grotiel sioner « wp ftowl giro} toners. 
x ee ousd uso SS Wf gepmond and 


mo & - = Lay 
On a a 


a 
Ge €e. ¢ 
water. rari 


arate “EC sdadoor acikomeen sxoteda acl fatamereg tO. sane 
efgna~) xo aceted newdo rdanonedira £6 nnordafy eon * 


elemte yeeom ,.mb dd oF os apitabtoet bos Hos co) toene re 


Zi. 


but sometimes branched; leaves also very variable, ovate, ovate-oblong 
or oblong-lanceolate, 0.8-15 cm. long, 0.6-8 cm. wide, coarsely den- 

tate or lyrate and sometimes with small projections at the base of the 
blade, occasionally dentations approach regularity, then are finer and 
smalier, lower long-petiolate, upper sessile; internodes mostly longer 
than the leaves; pedicels mostly short and thick, not twice the length 


of the calyx, occasionally thin and elongate. 


KEY TO VARIETIES 
Leaves often longer than 2 cm., stems taller than 2.5 cm.; corolla 
mostly much exceeding the calyx. 
Leaves 8-15 cm. long, 6-8 cm. wide; calyx in fruit 1-2.5 em. 
oRe ee Oso= ls SCR WIde «se 8 4 Ale we ee «= ee MM. BOTT 
var. GUTTATUS 
Leaves 1-3.5 cm. long, 1-1.5 cm. wide; calyx in fruit 0.5-1.2 
em.eoneyr Ds-0.0 tmaiwide . 1 6 «© « « « i « « 15Bs. Me. GUITATUS 
var. DEPAUPERATUS 
Leaves mostly shorter than 2 cm.; stems not exceeding 2.5 cm.; corolla 
ijteleswenger than therendyx whi... . . » . + 15 ¢. M. GUITATUS 


var. HALLII 


15 a. MIMULUS GUTTATUS var. GUTTATUS 
Widely distributed in wet, exposed places throughout the State, 


variable; the varieties confluent with the typical. 


seb “feansoo ie ee ‘Bed a peg 
Ad: te esed edt ds eno Lesetort see she oonttonoe bia 8 Hg 


vet - 


ng meat ove cedd pea sosonae bsg aa 


aatrrateav ik 7 


eiforson zt 2.5 ssid relied erage ¢ Md § aid sognol ato. ae 


xyes ers greebaeers: dour. vif 
: . - = 

mo 2.3-£ ftext of wefeo-tebiw oo 8-3. yenol amo 20-8 eovsed . 
CUTATTOD fee ef Leb ate » wpe 2 oR smo 25L+8.0 goof i 


osTaTTy) «ay 


GS. f-d,0 tiovt at ayfso pebiw .mo eefal sgaiol amo Re -£ sovast Pen: 
TATT MP RE a ne ee we Ot am 6 
oh TARA a JR Ey 


s{foxoo ¢.9d 2.8) en nid esoxe tom emeda as ae S sets sorrel citaom & 


SUTATIUO .M 2 ef: MINE. 2 osu gls2 ont aeett nonsot ott 


LIJAM iter 


34s. 


15 b. M. GUTTATUS var. DEPAUPERATUS (Gray) Grant, Ann. Mo. Bot. Gard. 
Lia (270. LAL 92h 


Clark and Elko Counties. 


15 c. M. GUITTATUS var. HALLII Greene) Grant, 1. c. 11: 172. 192). 


Elko Co. 


16. MIMULUS TILINGI Regel, Gartenfl. 18: 321, pl. 631. 1869. 


KEY TO VARIETIES 
Plants 5-20 cm. long; leaves elliptical to broadly ovate, lower short- 
petioled; stems usually bearing more than 3 flowers. 
16 a. M. TILINGI 
var. TILINGI 
Plants 20-0 cm. long; leaves oblong to suborbicular, lower long-petio- 
dates stem bearing 1-3 flowers. VAMLINTIO®N. . .... 16d. M. TILINGI 


var. CORALLINUS 


16 a. MIMULUS TILINGI var. TILINGI 
From dry to aquatic (mostly wet or muddy) places. 
Churchill, Douglas, Elko, Esmeralda, Mineral, Nye, Ormsby and 


Washoe Counties. 


16 b. MIMULUS TILINGI var. CORALLINUS (Greene) Grant, Ann. Mo. Bot. 
Gard. ll: 155. 192k. 


Wet places..Washoe Co. 


ae 


Sei: SYP eh ag-sk. us ( saved) uae ad -eonareso 


BB Ed dq AME BE Mnedied Legell TOMLITT 20 
“QT TSLAAY OF Ya 
CORLITT co ae Of 


TORLILT sxe 


ere ye gre t tewol qmnlvaks ye od egetts eeveo! 


‘ye, OM pn r 
Rae iT J << Gt 
te a Me 
Conia sO 4 "eo 
B22 ed 


bre vdeo ,eyi ,iavanih poe ee 


40d .oM .nod ,daexd fenwet8) 
f 


eo 


17. MIMULUS NASUTUS Greene, Bull. Calif. Acad. Sci. 1: 112. 1885. 
M. luteus in part of authors; M. langsdorfii in part of 

authors; M. subreniformis Greene; M. glareosus Greene; M. langs- 
dorfii var. nasutus Jepson; M. erosus Greene; M. minusculus Greene; 
M. cordatus Greene; M. cuspidatus Greene; M. puberulus Gdgr.; M. 
bakeri Gdgr.; M. parishii Gdgr., not Greene. 

Annual; puberulent to almost glabrous, rarely pubescent plants; 
stem erect or ascending, almost always branched, up to 6 am., l- 
angled, often winged; leaves round-ovate to oblong or rarely some- 
what ovate-lanceolate, 5-12 cm. long, almost as broad, rarely equal- 
ing the length, 3-5-nerved from the base, often red-tinged below, 
lower with clasping petioles, upper sessile; flowers in a raceme or 
raceme-like inflorescence; pedicels 2 to 3 or more times longer than 


the calyx, erect but recurving in fruit. 


KEY TO VARIETIES 
Floral leaves tinged with red below; stems usually stout, erect; co- 
Pete Oe iaeeS CMs LORE «cis 8 6s se we ew 17 a. M. NASUTUS 
var. NASUTUS 
Floral leaves densely, white villous below; stems weak, reclining; 
Sorel = ts5-1.5 Gm. tome. . «5 + ss 1 «+» « 17 b. M. NASDTUS 


var. MICRANTHUS 


17a. MIMULUS NASUTUS var. NASUTUS 
Dry tc aquatic, mostly exposed, damp to wet places. 


Clark, Elko, Humboldt, Lander, Lyon and Washoe Counties. 


\B6BE Skt ef £08 Mien eo it es 
to deg nt £ Hrobasa! H setoHides % reg. wm agerut | 


' apacdel .M. penesid ey ‘= paneese, esmso be 


Ae 


Need 


(petest) auediroe uel tis oth" panger®, arpoxe ait noege suggest ay : 


iy 


oe 


Mo evrbd ay Lioredry at Ye yenpen peseb Geir: ee artebree 
| sree th. tort: get de bee. pee gunsbo 
cadaala tnaot sede ‘yfowst awotdals saonit ee - dookeredeg qlesnmk 
ai , smh 4 od ad: sberlomaud eqn, éaomle cnetbnesea 10, = 
<emee yieteat tC giolde od. sdevo-baure't, eovsel woegaty nedte ss 


aul | 


~laspe yister .bao ard eB teomis ae ot Stee: vetefosmnal-odsv9 | 


TO CHGGAT s.r ae wad ,of te 


tact eognol semis erom wo € oF e: er toathen yeonooseroltat out Iononet 


eal TeETAy “et. “tea ae wae 
: ere DE 
93 siooTo .dirote elisvar anode & rwoled bet Atiw hegntt pre: fen LT a 
aed 0 1a TE 1 ww ha ee ww ae we ee BOL wo E.S-T, 0 
Gus at .18T 
‘auintfoet jisew emede yvoled svoliiv edinw <foena eevnel 
: A b ; ‘ aot 
ROAR LETS t 8 id med vL e * * = er “ . * ° a eee mrad. — ERB 
GUBTUAHOIM ster 
, a ee 
tran Be aera 
: soosl4 ane od on .bonogre yiteon oth ft 


ms a 


q 
nes Pole) ania elt bree ane prerrs ish Rescate ae ot 


Ee 


17 b. MIMULUS NASUTUS var. MICRANTHUS (Heller) Grant, Ann. Mo. Bot. 
Gard. ll: 182. 192k. 


Wet places. Elko Co. 


18. MIMULUS TORREYI Gray, Proc. Am. Acad. 11: 97. 1876. 
Eunanus torreyi Greene; E. fremontii Gray, not Greene. 

Annual; erect, glandular-pubescent plants; stems 0.5-3.5 cm. 
high, simple or branched; leaves scattered, varying from spatulate to 
ovate, 1.5-3.5 cm. long, 0.3-1.3 em. broad, short-petiolate, entire, 
or with a few teeth, obtuse, often red-tinged below; internodes usu- 
ally exceeding the leaves; pedicels 2-3 mm. long. 


Rare. Lincoln Co. 


19. MIMULUS ANGUSTIFOLIUS (Greene) Grant, Ann. Mo. Bot. Gard. 11: 298. 
192). 
Eunanus angustifolius Greene 
Annual; glandular-pubescent plants; stem red, up to 10 cm., 
mostly branched; leaves numerous, often clustered, linear or oblance- 
Olate, 1-1.7 cm. long, 1-3 mm. wide, entire, sessile; corolla lips 
very unequal. 


Known only from Nevada. Elko, Mineral and Washoe Counties. 


20. MIMULUS NANUS Hook. & Arn., Bot. Beechey's Voyage, 378. 18). 
M. nanus pluriflorus Hook. & Arn., Eunanus tolmei Benth., 
E. nanus Holz. 


Annual; viscid-puberulent to glandular-pubescent plants; stems 


onset ae +5 beanoment : tensetd ser $ suns 

md @.€-2.0 amste ¢ adns i¢ aeoueduq-za Lubna ly efoere elseanh 

3 adelodage mont arty BY .beredtsoa novael ebodonastd: tO ofgmie = 
eaten otelotie nd rode paoid «iD £. fat, 0 cgno ama: 2 BEAR, ics 
~yau weboutedck 74 woke Se egnidabet ted Lo sentdo “iess. wet 8 stg 


spool .mm €-8° nigueee isovsal od? gittbeosxs . 


499 niooaid oral f, 


328 aa bys) .fo8 .oM cand .dnetd (easesd) QULIOITevOMA. ee 
es ASAE 

enso rd ayiiotteugas eunsasi essere 

2m Of of qu bot mode pediatq jnooueduqenatobrals view 

~sonetide ra tesa tt ootsten ie medto _EsOrLO Mt eaveel sbedonead 5 f 

sqif-sfioro jolieves yotisne .ebiw om Eat ygarot ar) PafaL a 


Py 
w 


|) 


up to 15 cm., usually branched, low and spreading, tinged with red, 
or straw-colored or green; lower leaves petiolate, upper sessile, all 
1-3-nerved from the base, obovate, narrowly ovate or oblong, up to 3 
em. long and 6 mm. wide, subentire, acute or obtuse; pedicels 2-3 cm. 
long; flowers many. 

Dry, exposed places. 

Douglas, Elko, Humboldt, Lander, Lyon, Nye, Ormsby, Pershing and 


Washoe Counties. 


21. MIMULUS SPISSUS Grant, Ann. Mo. Bot. Gard. 11: 277. 192). 
Annuals; stem 15 cm. tall, glandular-villous, much-branched; 
flowers and leaves densely crowded near the apex; leaves broadly ob- 

ovate, short-petiolate, acute or cuspidate, entire, 1-15 cm. long, 


)-~6 mm. broad, 3-5-nerved from the base; pedicels 1-2 mm. long. 


KEY TO VARIETIES 
Leaves ovate; stigma lobes unequal, ciliate . .. 21a. M. SPISSUS 
var. SPISSUS 
Leaves lance-ovate or ellipticals; stigma lobes equal, eciliate. 
21 b. M. SPISSUS 


var. LINCOLENSIS 


21 a. MIMULUS SPISSUS var. SPISSUS 


Mostly dry places. Clark, Lincoln and Nye Counties. 


~ 


ber deior begatd. csoitbneage ‘ade. wor be 
Is flesea soca anaes tog scrial some 4 
& oF ay _3ne Ido 10 ednwe 3 ceo sis <sterada sand veut 


a 


as im Rea 
no £.S. 2fesfhed peautde 0" ave. soidnedee ebtw comin 8 me 
4 8 * : Bach 0 


endo eoectinatibas cits to 2 meta = ta 


wo 
A 
a) 
cs, 
3 


SRiTETeAT OF ord 


; + oteilto Cab tescotidaeia enti 


epeerte .ccy 


Breve TOOL . rar 


aertavad exit bre afoonda oto 


* 


wha: 


21 b. MIMULUS SPISSUS var. LINCOLNENSIS Edwin, L. West. Bot. 7: 222. 
1955. 


Lincoln Co. 


22. MIMULUS BIGELOVII Gray in Proc. Am. Acad. 11: 96. 1876. 
Eunanus bigelovii Gray 
Annuals; stem mostly branched, up to 25 cm. high, glandular- 
pubescent; leaves mostly elliptical, sometimes narrowly obovate, ta- 
pering to an acute apex, scattered, mostly entire or with a few teeth 
near the tip, short-petiolate, subsessile or occasionally sessile; 
pedicels 1.5-3 (-6) mm. long; calyx sometimes viscid-villous, usually 


having pronounced pubescence. 


KEY TO VARIETIES 
Leaves only narrowly obovate, tapering to narrowly sessile bases, l- 
nerved from the base, not cuspidate-tipped. ... 22 a. M. BIGELOVII 
var. BIGELOVII 
Leaves broadly obovate or ovate, sessile by a broad base, 3-5-nerved 
from the base, strongly cuspidate-tipped. ... . 22 b. M. BIGELOVII 


var. OVATUS 


22 b. MIMULUS BIGELOVII var. BIGELOVII 
Dry sites, 2500-11,200 ft. 
Clark, Douglas, Esmeralda, Lincoln, Mineral, Ormsby and Washoe 


Counties. 


Sc& :¥_ cod -dueW vd ates eremiaugoin 


© 


Ton etd aa Panera! vi he cea Ter ss 


retiseas 1 Sasadessae 0 eliepsaulia ere 7 oe ie 


faivepy ,crolliv-bisutw-aenttomon' eelEo- qari tans (B-) EBL 2 é 


ae eonsosadirg © penmronas as 


oe. faces ceili Seg (eend, add mort & 
Vo GROTE tee 
eTTotei—— ,eted bsowd 2 yo eitacer ,.etexo vo edevodo yibsorrd - 


VOIGNEE Mid SS. . 4 w cbeqgtt-etehtgeco ylgnotta. gesed|e 


J 


eonss* bas ydamy0) trons, casan spetaranat 


~20- 


22 b. MIMULUS BIGELOVII var. OVATUS Gray, Syn. Fl. No. Am. ed 2, 2, pt. 
1, suppl. 45. 1886. 
M. bigelovii var. cuspidatus Grant; M. cusickii (Greene) Piper; 
Eunanus cusickii Greene. 
Dry places. 


Clark, Esmeralda, Lander, Lincoln, Nye and Washoe Counties. 


23. MIMULUS DENSUS Grant, Ann. Mo. Bot. Gard. 11: 298. 192). 

Annual; glandular-pubescent, viscid plants, sometimes viscid- 
villous; stem mostly much-branched, up to 15 cm.; leaves numerous, 
oblong, linear or elliptical, 1.5-2.5 cm. long, 2.5 mm. broad, sessile; 
flowers axillary, numerous; pedicels 2.5 mm. long. 

Very difficult to separate from the older M. mephiticus Greene, 
possibly a synonym. Eastwood & Howell 9138 (Calif. Academy of Sci- 
ences and State College of Washington) is an intergrading specimen. 

Mostly dry places, up to about 10,000 ft. 

Churchill, Elko, Lander, Mineral, Nye, Ormsby and Washoe 


Counties. 


2). MIMULUS MEPHITICUS Greene, Bull. Calif. Acad. Sci. 1: 9. 188). 
Eunanus mephiticus Greene. 
Annual; stems erect, simple or branched, glandular-pubescent 
ane¢ viscid; leaves varying from nearly linear to oblanceolate, 1-2.2 
em. long, 2-5 mm. broad, entire, sessile or short-petiolate, obtuse or 
acute; pedicels 2-l mm. long; the mephitic odor often disappears on 


drying. 


Ht (sage oi} PRISE eyo M gids sO Bybee 


des s£e be ent fo om an dain ae: 


® 


bioaiy sacckioraa “ eeagtd big ane . droosedias tobi € 
ry : . 


sorter eevee gem ef at oa be seit cbbeiine ‘eideom: cate 


oe uot st € or = of ieobightte ee asenit 


\pnot! unit 2 ‘elesibed ¢ paurortomset (eens 
puoi shies at ag Oe mre ere’. adeesqee og as rae eN tb vse 
+ xt oe 7 


Gaarecmecnhdarin = AUT 


ae x 
& Ate osu 


soO Ley wwebwok \tRisd}. ELE? Ifow a  boo'ehast - uaa a ee 


~ 


amkbese gntbergietad as eb-tdooar tieatt ‘26 egeifos aseae baie 


at O00 Of duoda ee! ep “a0 Leg pte at oh 
} i 
safes! bre. efsrrto 


faa 


ue 


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SSH ateloosns fig od aepmtl st lmao org ee seb 


at i> 
iy 


iy ganda asnls bfsghexoda acy olimaee certdes is sate 


mo anmpeaqseib rn Hie gba 2 Bilas ont. wy 


ii | er 


~21- 


Dry places. Esmeralda and Washoe Counties. 


25. MIMULUS COCCINEUS Congdon, Erythea 7: 187. 1899. 

Annual; stem erect, up to 6 cm., often branched; entire plant 
glandular-puberulent (occasionally almost glabrous); leaves linear to 
oblanceolate, small, up to 1 cm. long, ) mm. broad, entire, usually 
obtuse, very short-petiolate or subsessile; pedicels rarely more than 
1 mm. long; flowers clustered at the tips of the stems; calyx cylin- 
drical, 3-5 mm. long. 


Rare. Douglas Co. 


26. MIMULUS WASHOENSIS Edwin, L. West. Bot. 7: 221. 1955. 

Annual; stem glandular-pubescent, upright, unbranched, up to 
10 cm. high; leaves sessile, entire or almost so, lanceolate, lance- 
linear, or elliptic (occasionally obovate) up to 1.8 cm. long, 3-6 
(-~8) mm. wide, ciliate; pedicels 2-) mm. long; calyx 8-11 mm. long, 
ovate to somewhat campanulate. 


Washoe Co. 


27. MIMULUS STAMINEUS Grant, Ann. Mo. Bot. Gard. 11: 302. 192h. 
Annual; stem branched, rarely simple, up to 10 cm. tall, dense- 
ly glandular-pubescent; leaves sessile, numerous, °blanceolate to 
elliptical or occasionally linear, 1-2.5 cm. long, 2-5 mm. broad, en- 
tire, obtuse; pedicels 1.2 mm. long; calyx tubular, 5-8 mm. long. 
Reported as occurring in Ormsby and Esmeralda Counties by Grant, 


i. ¢. The only specimen seen by the author is C. F. Baker 1375, Snow 


yiiace certian 


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