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CHECKLIST OF AMPHIBIANS AND REPTILES
OF THE
SAGUA-BARACOA MOUNTAINS, EASTERN CUBA

Ansel Fong G' and Nils Navarro”
1BIOECO, Museo de Historia Natural “Tomas Romay”
*Museo de Historia Natural “Carlos de la Torre”

/EXMHSON aN

MAR 1 2 2093

SMITHSONIAN
HERPETOLOGICAL INFORMATION
SERVICE
NO. 130

2001

SMITHSONIAN
HERPETOLOGICAL
INFORMATION
SERVICE

The SHIS series publishes and distributes translations,
bibliographies, indices, and similar items judged useful
to individuals interested in the biology of amphibians
and reptiles, but unlikely to be published in the normal
technical journals. Single copies are distributed free
to interested individuals. Libraries, herpetological
associations, and research laboratories are invited to
exchange their publications with the Division of
Amphibians and Reptiles.

We wish to encourage individuals to share their
bibliographies, translations, etc. with other
herpetologists through the SHIS series. If you have
such items please contact George Zug for instructions on
preparation and submission. Contributors receive 50
free copies.

Please address all requests for copies and inquiries to
George Zug, Division of Amphibians and Reptiles,

National Museum of Natural History, Smithsonian

Institution, Washington DC 20560 USA. Please include

a self-addressed mailing label with requests.

INTRODUCTION

The mountains of eastern Cuba mountains are the main center of floristic and faunistic diversity
and have the highest endemism on the island. Considering only the vertebrates, these mountains
harbor 71% of Cuba’s species and 30% of its endemic species (Fong, 1994). Specifically, the
Sagua-Baracoa Mountains contain, together with Sierra Maestra, the most endemic amphibians and
reptiles and the greatest number of species. Fifty-three percent of Cuban amphibians and 45.6% of
Cuban reptiles occur in Sagua-Baracoa. Although these mountains comprise less than a fourth of
the Cuban territory, they have half of the Cuban herpetofauna.

The high endemism (93.1% in amphibians and 74.2% in reptiles), along with 40% of the
species being typical of Sagua-Baracoa, contribute to the national and global importance of this
area. For this reason, we prepared the following checklist. It includes all species of amphibians and
reptiles of Sagua-Baracoa described through June 2000. The list also summarizes the endemism
and distribution of the S-B herpetofauna. Despite the currency of this report, it is not conclusive
because new species continue to be described. We agree with the statement of Schwartz and
Henderson (1991): ““we hope that new species are discovered and described for many years to

come”.

The compilation of this checklist is based on the author’s field work and observations
throughout the eastern Cuba, as well as data from the Cuban herpetological collections of the
Instituto de Ecologia y Sistematica, Museo Nacional de Historia Natural, Museo de Historia Natural
of Holguin, and Centro Oriental de Ecosistemas y Biodiversidad (BIOECO) of Santiago de Cuba.
We also used the herpetological database created for a previous work (Fong, 1994), in which all
literature about Eastern Cuban vertebrates through 1993 was reviewed. In addition to that literature
we include new species descriptions from 1994 onward.

We define the Sagua-Baracoa Mountains (following Nufiez et al., 1989) as consisting of the
Sierra de Nipe (SN), Alturas del Segundo Frente (SF), Sierra del Cristal (SC), Sierra de Sagua de
Taénamo (ST), Gran Meseta de Guantanamo (MG), Cuchillas de Toa (CT), Alturas de Moa (AM),
Cuchillas y Mesas de Baracoa-Imias (CB), Gran Meseta Carsica de Baracoa (MB), Mesas Costeras
Aterrazadas de Guantanamo-Cajobabo (TG) and Llanura Costera Yaguaneque-Baracoa (LC). For a
detailed map of the Sagua-Baracoa region see Nufiez et al. (1989).

Abbreviations are as follow: EN = Endemism, E = Cuban endemic, R = Regional Cuban
endemic (known only from Sagua-Baracoa), L = Local Cuban endemic (known only from isolated
localities in Sagua-Baracoa), - = No endemic species, I = Introduced species.

i)

Acknowledgements

Most results of this report derive from the "Diversidad Biologica de los Macizos Montanosos
Sierra Maestra y Nipe-Sagua-Baracoa” project, which supported portions of our field work. Luis V.
Moreno and Orlando H. Garrido provided very valuable assistance to our studies in the last years.

S. Blair Hedges sent us some important literature on the Cuban herpetofauna. [liana Boza and

Dylan Robison help us with the translation to English.

AMPHIBIA

ANURA
Bufonidae

Bufo longinasus

Bufo peltocephalus

Bufo taladai

Hylidae
Osteopilus septentrionalis

Leptodactylidae
Eleutherodactylus acmonis

Eleutherodactylus atkinsi

Eleutherodactylus auriculatus

Eleutherodactylus bartonsmithi

Eleutherodactylus bresslerae

EN

CHECKLIST

DISTRIBUTION

MG

SN, SC, ST, MG,
CT, AM, MB, TG

SN, MG, CT, AM,
CB, MB, TG

SN,SE, S@ysi,
MG, CT, AM, CB,
MB, TG, LC

SC, ST, MG, CT,
AM, CB, MB

SN, SC, MG, CT,
AM, CB, MB, TG,
Le

ST, MG, CT, AM,
CB

ST, MB

DATA SOURCES

Valdés and Ruiz (1980)
Garrido and Jaume (1984)
Estrada et al.(1987)
Abreu et al. (1989)
Authors’ unpub. data
Garrido and Jaume (1984)
Estrada et al.(1987)
Authors’ unpub. data
Collection’s data

Estrada et al.(1987)
Abreu et al. (1989)
Authors’ unpub. data
Collection’s data

Garrido and Jaume (1984)
Estrada et al.(1987)

Abreu et al. (1989)
Collection’s data

Estrada et al.(1987)

Abreu et al. (1989) Hedges et al.
(1992)

Authors’ unpub. data
Collection’s data

Estrada et al.(1987)
Estrada and Hedges (1997)
Authors’ unpub. data
Collection’s data

Garrido and Jaume (1984)
Hedges et al. (1992)
Collection’s data

Garrido and Jaume (1984)
Silva Taboada (1988)
Abreu et al. (1989)
Collection’s data

Eleutherodactylus cuneatus

Eleutherodactylus dimidiatus

Eleutherodactylus guantanamera

Eleutherodactylus gundlachi
Eleutherodactylus iberia

Eleutherodactylus intermedius
Eleutherodactylus ionthus

Eleutherodactylus leberi

Eleutherodactylus limbatus

Eleutherodactylus mariposa

Eleutherodactylus orientalis

Eleutherodactylus pezopetrus
Eleutherodactylus planirostris

Eleutherodactylus principalis
Eleutherodactylus ricordi

SN, SC, ST, MG,
CT, AM, CB, MB,
LC

SN, MG, CT, AM,
MB

MG, CT, AM, CB

MG, CB
CB

CB

SN, SF, SC, ST,
MG, CT, AM, CB,
MB

MG

SN, SC, ST, MG,
CT, AM, MB

MG
MG, CB

SN, MG
SN, SC, MG, CT,
AM, CB, MB, LG

ST, CT, AM, CB
SN SEIMG: Ci,
AM, CB, MB

Barbour and Ramsden (1919)
Garrido and Jaume (1984)
Estrada et al.(1987)
Hedges et al. (1992)
Estrada and Hedges (1998)
Authors’ unpub. data
Collection’s data

Garrido and Jaume (1984)
Estrada et al.(1987)
Hedges et al. (1992)
Authors’ unpub. data
Hedges et al. (1992)

Fong (2000)

Authors’ unpub. data
Collection’s data

Garrido and Jaume (1984)
Estrada and Hedges (1996b)
Authors’ unpub. data
Garrido and Jaume (1984)
Garrido and Jaume (1984)
Estrada et al.(1987)
Authors’ unpub. data
Collection’s data

Garrido and Jaume (1984)
Hedges et al. (1992)
Estrada et al.(1987)

Silva Taboada (1988)
Hedges et al. (1992)
Authors’ unpub. data
Collection’s data

Hedges et al. (1992)
Garrido and Jaume (1984)
Estrada and Hedges (1996b)
Collection’s data
Collection’s data

Estrada et al.(1987)

Abreu et al. (1989)
Authors’ unpub. data
Collection’s data

Estrada and Hedges (1997)
Hedges et al. (1992)
Authors’ unpub. data
Collection’s data

Eleutherodactylus riparius
Eleutherodactylus ronaldi

Eleutherodactylus tetajulia

Eleutherodactylus toa

Eleutherodactylus varleyi

Rana catesbeiana
REPTILIA
SQUAMATA

Amphisbaenidae
Amphisbaena cubana

Anguidae
Diploglossus nigropunctatus

Gekkonidae
Gonatodes albogularis

Hemidactylus haitianus

Sphaerodactylus armast

Sphaerodactylus bromeliarum

SN
ST, MG, CT, AM,
CB

CB

SF, ST,.MG, CT,
AM, MB

SN, SC, MG, AM,
CB

SF, SN, SE

SN) SC; Cr

SCSMG, CL CB:
MB

SC, MB, LC

SN, SC, MB, LC

MB, TG

CB

Authors’ unpub. data
Garrido and Jaume (1984),
Estrada et al.(1987)
Hedges et al. (1992)
Authors’ unpub. data
Collection’s data

Estrada and Hedges (1996a)
Authors’ unpub. data
Estrada and Hedges (1991)
Fong (2000)

Collection’s data

Estrada et al.(1987)
Collection’s data

Authors’ unpub. data
Authors’ unpub. data

Estrada et al.(1987)

Thomas and Hedges (1998a)
Navarro et al. (in press)
Authors’ unpub. data

Garrido and Jaume (1984)

Estrada et al.(1987) Abreu et al.
(1989) Thomas and Hedges (1998b)
Authors’ unpub. data

Collection’s data

Authors’ unpub. data
Collection’s data

Authors’ unpub. data
Collection’s data

Schwartz and Garrido (1974)
Garrido and Jaume (1984)
Schwartz and Garrido (1985)
Collection’s data

Garrido and Jaume (1984)
Schwartz and Garrido (1985)

Sphaerodactylus celicara

Sphaerodactylus elegans

Sphaerodactylus nigropunctatus

Sphaerodactylus notatus

Sphaerodactylus ramsdeni

Sphaerodactylus ruibali

Tarentola americana

Iguanidae/Iguaninae
Cyclura nubila

Iguanidae/Polychrotidae
Anolis alayoni
Anolis alfaroi

Anolis allogus

CT, AM, CB, MB,
IU

ST, CB

MG, LC

SC, MG,CB, MB,
LC

MG

TG

SN, SC, MB, TG,
LC

CB, TG, MB

SN, SC, MG, CT,
AM, CB, MB, LC
MG

SN, SC, ST, MG,

CT, AM, CB, MB,
LC

Garrido and Jaume (1984)
Garrido and Schwartz (1982)
Estrada et al.(1987) Authors’
unpub. data Collection’s data
Schwartz and Garrido (1985)
Authors’ unpub. data
Collection’s data

Garrido and Jaume (1984)
Schwartz and Garrido (1985)
Authors’ unpub. data
Collection’s data

Schwartz and Garrido (1985)
Authors’ unpub. data
Collection’s data

Garrido and Jaume (1984)
Schwartz and Garrido (1985)
Thomas et al. (1992)
Collection’s data

Garrido and Jaume (1984)
Schwartz and Garrido (1985)
Authors’ unpub. data
Collection’s data

Barbour and Ramsden (1919)
Abreu et al. (1989)

Navarro et al. (in press)
Authors’ unpub. data
Collection’s data

Barbour and Ramsden (1919)
Authors’ unpub. data

Estrada and Hedges (1995)
Navarro et al. (in press)
Authors’ unpub. data
Garrido and Hedges (1992)
Collection’s data

Barbour and Ramsden (1919)
Estrada et al.(1987)

Abreu et al. (1989)

Authors’ unpub. data
Collection’s data

Anolis alutaceus

Anolis anfiloquioi

Anolis angusticeps

Anolis argenteolus

Anolis argillaceus

Anolis baracoae

Anolis centralis

Anolis cupeyalensis

Anolis cyanopleurus

Anolis fugitivus

SN, SC, ST, MG,
CT, AM, CB

SN, SC, CB, MB,

TG

MG, AM, CB, MB,
LC

SN, SC, ST, MG,
CT, AM, CB, MB,
TG, LG

SCJSTIMGET,
AM, CB, MB

CT, CB,MB

MG, MB,TG

SN, SC, ST, MG,
CT

SC, MG, CT,
CB,MB

AM, LC

Garrido (1980) Garrido and Jaume
(1984) Estrada et al.(1987)

Abreu et al. (1989) Garrido and
Estrada (1989) Navarro et al. (in
press) Authors’ unpub. data
Collection’s data

Garrido (1980) Garrido and Jaume
(1984) Navarro et al. (in press)
Authors’ unpub. data

Collection’s data

Garrido (1975c) Estrada et al.(1987)
Estrada and Hedges (1995)
Authors’ unpub. data

Collection’s data

Estrada et al.(1987) Abreu et al.
(1989) Navarro et al. (in press)
Authors’ unpub. data

Collection’s data

Garrido (1975a)

Estrada et al.(1987)

Authors’ unpub. data

Collection’s data

Schwartz and Garrido (1972)
Garrido and Jaume (1984)
Authors’ unpub. data

Collection’s data

Garrido (1975a)

Garrido and Jaume (1984)
Authors’ unpub. data

Collection’s data

Garrido (1975b) Garrido and Jaume
(1984) Estrada et al.(1987)
Schwartz and Henderson (1991)
Authors’ unpub. data

Collection’s data

Garrido (1975b) Garrido and Jaume
(1984) Estrada et al.(1987) Abreu
et al. (1989) Authors’ unpub. data
Collection’s data

Garrido (1980)

Garrido and Jaume (1984)
Authors’ unpub. data

Anolis homolechis

Anolis imias

Anolis inexpectatus

Anolis isolepis

Anolis jubar

Anolis loysianus

Anolis macilentus

Anolis noblei

Anolis ophiolepis

Anolis porcatus

Anolis rubribarbus

Anolis sagrei

SN, SC, ST, MG,
CT, AM; CB, LC

MB, TG

ST, CT, AM

SN, SC, ST, MG,
CT, AM, MB, LC

MB, TG, LC

SN, SC, ST, CT,
AM

MG

SN, ST, CT

SN, MG, AM

SN, SF, SC, ST,
MG, CT, AM, CB,
MB, TG, LC

ST, CT, AM, LC

SN, SF, SC, ST,

MG, CT, AM, CB,
MB, TG, LC

Garrido (1975b) Estrada et al.
(1987) Abreu et al. (1989)
Authors’ unpub. data

Collection’s data

Garrido and Jaume (1984)

Fong (2000)

Collection’s data

Garrido and Estrada (1989)
Authors’ unpub. data
Collection’s data

Garrido and Jaume (1984)
Estrada et al.(1987) Navarro et al.
(in press) Authors’ unpub. data
Collection’s data

Garrido (1973a) Garrido and Jaume
(1984) Silva Taboada (1988)
Authors’ unpub. data

Collection’s data

Barbour and Ramsden (1919)
Estrada et al.(1987) Navarro et al.
(in press) Authors’ unpub. data
Garrido and Hedges (1992)
Authors’ unpub. data
Collection’s data

Schwartz and Garrido (1972)
Garrido and Jaume (1984)
Authors’ unpub. data

Garrido and Jaume (1984) Navarro
et al. (in press) Authors’ unpub.
data Collection’s data

Estrada et al.(1987)

Abreu et al. (1989)

Authors’ unpub. data

Collection’s data

Garrido and Jaume (1984) Estrada
et al.(1987) Authors’ unpub. data
Collection’s data

Estrada et al.(1987)

Abreu et al. (1989)

Authors’ unpub. data

Collection’s data

Anolis smallwoodi

Anolis sp.
Anolis vescus

Chamaeleolis porcus

Iguanidae/Tropiduridae
Leiocephalus carinatus

Leiocephalus cubensis

Leiocephalus macropus

Leiocephalus onaneyi

Leiocephalus raviceps

Teiidae
Ameiva auberi

Boidae
Epicrates angulifer

SC, ST, MG, CT,
AM, TG, LC

CM

CB

SN, SC, MG, CB,
MB

CB, MB, TG, LC

SN, SC, MG, AM,
LE

SN, SF, SC, ST,
MG, CT, AM, CB,
MB, TG, LC

TG

MB, TG

SN, SC, ST, CT,
AM, MB, TG, LC

SNjSE) SE, .CL,
AM, CB

Schwartz and Garrido (1972)
Garrido and Jaume (1984) Estrada
et al.(1987) Schwartz and
Henderson (1991) Authors’ unpub.
data Collection’s data

Fong and Garrido (in press)

Garnido and Hedges (1992)
Collection’s data

Garrido (1982)

Garrido and Jaume (1984) Navarro
et al.(in press) Authors’ unpub.
data Collection’s data

Garrido and Jaume (1984)
Collection’s data

Garrido and Jaume (1984) Abreu et
al. (1989) Navarro et al.(in press)
Authors’ unpub. data

Garrido (1979) Garrido and Jaume
(1984) Estrada et al.(1987) Abreu
et al. (1989) Navarro et al.(in press)
Authors’ unpub. data

Collection’s data

Garrido (1973b)

Garrido and Jaume (1984)

Garrido and Jaume (1984)

Authors’ unpub. data

Collection’s data

Garrido and Jaume (1984) Estrada
et al.(1987) Navarro et al.(in press)
Authors’ unpub. data

Collection’s data

Estrada et al.(1987)
Abreu et al. (1989)
Navarro et al. (in press)
Authors’ unpub. data
Collection’s data

Colubridae
Alsophis cantherigerus

Antillophis andreae

Arrhyton landoi

Arrhyton supernum
Tretanorhinus variabilis
Tropidophidae

Tropidophis fuscus

Tropidophis melanurus

Tropidophis pilsbryi

Tropidophis wrighti

Typhlopidae
Typhlops biminiensis
Typhlops lumbricalis

TESTUDINES
Emydidae
Trachemys decussata

SN, SC, ST, MG,
CT, AM, CB, MB,
TG

SN, SC, MG, CT,
AM, CB, MB, TG

CB, MB, TG

MG, CB

SN, SC, AM
CT, AM, CB, MB

SN, SC, ST, MG,
CT, AM, CB, MB,
TG, LC

SN, MB

SN, CT, AM

MB
SN, ST, MG, CT,
AM, CB, MB

SN

Garrido and Jaume (1984) Estrada
et al.(1987) Silva Taboada (1988)
Abreu et al. (1989) Navarro et al.
(in press) Authors’ unpub. data
Collection’s data

Estrada et al.(1987) Abreu et al.
(1989) Navarro et al. (in press)
Authors’ unpub. data

Collection’s data

Schwartz and Garrido (1981)
Garrido and Jaume (1984)
Hedges and Garrido (1992a)
Hedges and Garrido (1992a)
Collection’s data

Navarro et al. (in press)
Collection’s data

Hedges and Garrido (1992b) Fong
(2000) Authors’ unpub. data
Collection’s data

Estrada et al.(1987) Hedges and
Garrido (1992b) Navarro et al. (in
press) Authors’ unpub. data
Collection’s data

Garrido and Jaume (1984)
Collection’s data

Estrada et al.(1987) Hedges and
Garrido (1992b) Authors’ unpub.
data Collection’s data

Authors’ unpub. data
Estrada et al.(1987)
Navarro et al. (in press)
Authors’ unpub. data
Collection’s data

Navarro et al. (in press)

LITERATURE CITED

Abreu, R. M., J. de la Cruz., A. Rams, and M. E. Garcia. 1989. Vertebrados del complejo
montafioso "La Zoilita", Holguin, Cuba. Poeyana 370:1-16.

Barbour, T. and C. T. Ramsden. 1919. The herpetology of Cuba. Mem. Mus. Comp. Zool. 47(2):71-
213.

Estrada, A. R. and S. B. Hedges. 1991. Nueva especie de Eleutherodactylus (Anura:
Leptodactylidae) de la Region Oriental de Cuba. Caribbean J. Sci., 27 (3-4):139-145.

Estrada, A. R. and S. B. Hedges. 1995. A new species of Anolis (Sauria: Iguanidae) from eastern
Cuba. Caribbean J. Sci., 31:65-72.

Estrada, A. R., and S. B. Hedges. 1996a. A new frog of the genus Eleutherodactylus from eastern
Cuba (Anura, Leptodactylidae). Herpetologica, 52:435-439.

Estrada, A. R. and S. B. Hedges. 1996b. At the lower size limit in tetrapods: a new diminutive frog
from Cuba (Leptodactylidae: Eleutherodactylus). Copeia, 1996 (4):852 -859.

Estrada, A. R. and S. B. Hedges. 1997. Nueva especie de Eleutherodactylus (Anura:
Leptodactylidae) del Macizo Sagua-Baracoa, Cuba. Caribbean J. Sci., 33 (3-4):222-226.

Estrada, A. R. and S. B. Hedges. 1998. Sistematica de las ranas riberefias de Cuba (Leptodactylydae:
Eleutherodactylus) con la descripcion de una especie nueva. Caribbean J. Sci., 34 (3-4):218-
230.

Estrada, A. R., G. Alayon, A. Pérez Asso, C. Pefia, and E. Solana. 1987. Lista preliminar de anfibios
y reptiles de las Cuchillas de Moa y Toa, Cuba. Garciana 8:3-4.

Fong, A. 1994. Biodiversidad de vertebrados de la Region Oriental de Cuba. Tesis de Licenciatura,
Universidad de Oriente, Santiago de Cuba.

Fong, A. 2000. Ampliacion de la distribucion de especies de anfibios y reptiles de Cuba oriental.
Biodiv. Cuba Oriental 4:87-89.

Fong, A. and O. H. Garrido. In press. Nueva especie de Anolis (Sauria: Iguanidae) de la regidn norte
de Cuba oriental. Rev. Biol. Trop. 48(2).

Garrido, O. H. 1973a. Distribucién y variacién de Anolis homolechis Cope (Lacertilia: Iguanidae) en
Cuba. Poeyana 120:1-68.

Garrido, O. H. 1973b. Nueva especie de Leiocephalus (Lacertilia: Iguanidae) para Cuba. Poeyana
116:1-19.

Garrido, O. H. 1975a. Distribucién y variacion de Anolis argillaceus Cope (Lacertilia: Iguanidae) en
Cuba. Poeyana 142:1-28.

Garrido, O. H. 1975b. Distribuci6n y variacién del complejo Anolis cyanopleurus (Lacertilia:
Iguanidae) en Cuba. Poeyana 143:1-60.

Garrido, O. H. 1975c. Variacion de Anolis angusticeps Hallowell (Lacertilia: Iguanidae) en el
occidente de Cuba y en Isla de Pinos. Poeyana 144:1-18.

Garrido, O. H. 1979. Nuevas subespecies de Leiocephalus macropus Cope (Lacertilia: Iguanidae)
para Cuba. Poeyana 188:1-16.

12

Garrido, O. H. 1980. Revision del complejo Anolis alutaceus (Lacertilia: Iguanidae) y descripcion
de una nueva especie de Cuba. Poeyana 201:1-41.

Garrido, O. H. 1982. Descripcion de una nueva especie cubana de Chamaeleolis (Lacertilia:
Iguanidae), con notas sobre su comportamiento. Poeyana 236:1-25.

Garrido, O. H. and A. R. Estrada. 1989. Nueva especie del complejo Anolis alutaceus (Lacertilia:
Iguanidae) para Cuba. Rev. Biol. 3(1):57-66.

Garrido, O. H. and S. B. Hedges. 1992. Three new grass anoles from Cuba (Squamata: Iguanidae).
Caribbean J. Sci. 28(1-2):21-29.

Garrido, O. H. and M. L. Jaume. 1984. Catalogo descriptivo de los anfibios y reptiles de Cuba.
Dojfiana, Acta Vertebrata 11(2):5-128.

Garrido, O. H. and A. Schwartz. 1982. A new species of Sphaerodactylus (Reptilia: Sauria:
Gekkonidae) from Eastern Cuba. Proc. Biol. Soc. Washington 95(2):392-397.

Hedges, S. B. and O. H. Garrido. 1992a. Cuban snakes of the genus Arrhyton: two new species and
a reconsideration of A. redimitum Cope. Herpetologica 48(2):168-177.

Hedges, S. B. and O. H. Garrido. 1992b. A new species of Tropidophis from Cuba (Serpentes:
Tropidophiidae). Copeia 3:820-825.

Hedges, S. B., A. R. Estrada and R. Thomas. 1992. Three new species of Eleutherodactylus from
eastern Cuba, with notes on vocalizations of other species (Anura: Leptodactylidae). Herpetol.
Monogr. 6:68-83.

Navarro, N., A. Fernandez, C. Pefia and E. Reyes. In press. Reptiles de la Sierra de Nipe. Garciana.
Nufiez, A., N. Vifia and A. Grafia. 1989. Regiones naturales y antropicas In Nuevo Atlas Nacional
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