A stereo-atlas of ostracod shells

A Stereo-Atlas of Ostracod Shells

edited by R. H. Bate, J. W. Neale, Lesley M. Sheppard

and David J. Siveter

Volume 9, Part 1; 16th July, 1982

Published by the British Micropalaeontological Society

Editors

Dr R.H. Bate, Department of Palaeontology, British Museum (Natural History), Cromwell Road,
London SW7 5BD.

Prof. J.W. Neale, Department of Geology, The University, Hull HU6 7RH.

Dr Lesley M. Sheppard, Department of Palaeontology, British Museum (Natural History), Cromwell
Road, London SW7 5BD.

Dr David J. Siveter, Department of Geology, The University, Leicester LEI 7RH.

Editorial Board

Dr G. Bonaduce, Stazione Zoologica, 80121 Napoli, Italy.

Dr J.-P. Colin, Esso Production Research - European, 213 Cours Victor Hugo, 33321 Begles,
France.

Dr P. De Deckker, Research School of Pacific Studies, Australian National University, PO Box 4,
Canberra ACT 2600, Australia.

Dr D. van Harten, Universiteit van Amsterdam, Geologisch Instituut, Nieuwe Prinsengracht 130,
Amsterdam, The Netherlands.

Dr I. Purper, Departamento de Paleontologia e Estratigrafia, UFRGS, 90 000 Porto Alegre RS, Brazil.

Dr R.E.L. Schallreuter, Universitat Hamburg, Geologisch-Palaontologisches Institut, Bundesstrasse
55, D 2000 Hamburg 13, West Germany.

Officers of the British Micropalaeontological Society

Chairman Dr B. Owens, Institute of Geological Sciences, Ring Road, Halton, Leeds.

Secretary Dr A.R. Lord, Department of Geology, University College London, Gower Street,
London WC1.

Treasurer Dr R.H. Bate, Department of Palaeontology, British Museum (Natural History), Cromwell
Road, London SW7 5BD.

Editor Dr L.M. Sheppard, Department of Palaeontology, British Museum (Natural History),
Cromwell Road, London SW7 5BD.

Circular Editor Dr E.G. Spinner, Department of Geology, The University, Mappin Street, Sheffield SI.

Conodont Group:

Chairman Dr R.L. Austin, Department of Geology, The University, Southampton.

Secretary Dr A.C. Higgins, Department of Geology, The University, Sheffield SI.

Foraminifera Group:

Chairman Dr M.B. Hart, Department of Geology, Plymouth Polytechnic, Drake Circus, Plymouth
PL4 8AA.

Secretary Dr M.D. Fewtrell, Erico Research Laboratories Ltd., Lane House, 233-235 Roehampton
Lane, London SW15 4LB.

Microplankton Group:

Chairman Mr J.R.G. Fenton, Robertson Research International Ltd., Ty’n-y-Coed, Llanrhos,
Llandudno, Gwynnedd LL30 ISA.

Secretary Mr R. Woollam, Institute of Geological Sciences, Ring Road, Halton, Leeds.

Ostracod Group:

Chairman Dr R.C. Whatley, Department of Geology, University College of Wales, Aberystwyth,
Dyfed SY23 3DB.

Secretary Dr D.J. Horne, Department of Geology, City of London Polytechnic, Walburgh House,
Bigland Street, London El.

Palynology Group:

Chairman Mr M.J. Fisher, British National Oil Corporation, 150 St. Vincent Street, Glasgow G2 5LJ.

Secretary Dr D.J. Smith, BP Research Centre, Chertsey Road, Sunbury-on-Thames, Middlesex.

Instructions to Authors

Contributions illustrated by scanning electron micrographs of Ostracoda in stereo-pairs are invited.

Format should follow the style set by the majority of papers in this issue. Descriptive matter apart

from illustrations should be cut to a minimum; preferably each plate should be accompanied by one

page of text only. Blanks to aid in mounting figures for plates may be obtained from any one of the

Editors or Editorial Board. Completed papers should be sent to Dr L.M. Sheppard.

The front cover shows a female left valve, external and internal views, of

Bilobatia serralobata Schallreuter.

Printed by United Printing Services, Blackpool, England.

Stereo-Atlas of Ostracod Shells 9(1) 1- 8 (1982)

595.336.13 (113.312) (492.71 : 161.008.54) : 551.35 + 552.55

Braderupia asymmetrica (1 of 8)

ON BRADERUPIA ASYMMETRICA (NECKAJA)

by Roger E. L. Schallreuter
(University of Hamburg, German Federal Republic)

Genus Braderupia gen. nov.

Type-species: Pseudostrepula asymmetrica Neckaja, 1958.

Derivation of name: Braderup, the locality of the figured specimens. Gender feminine.

Diagnosis: Small to medium-sized palaeocope. Unisulcate; S2 moderately long. Asymmetrical: right valve has a
distinct spine-like posteroventral lobe; in left valve it is absent or occurs (especially in females) only
as a weak inflation. Velum in males and larger tecnomorphs is a rounded ridge, in females it forms a
flange-like dolon, and is absent in young instars. Velar antrum admarginal, occurs antero- and
centroventrally. Histium developed only in anteroventral and posteroventral regions, forms a more
or less distinct, rounded ridge, and is only slightly dimorphic (weaker posteroventrally in males);
histial antrum absent. Histiovelar furrow fissum-like anteriorly, posteriorly has a v-shaped profile
and irregular row of puncta, continued posteriorly as an indistinct semisulcus. Laterohistial furrow
anteriorly forms a distinct fissum parallel to histiovelar fissum-like furrow, posteriorly forms a more
or less distinct u-shaped furrow or is absent. Marginal sculpture is a ridge or row of spines.

Explanation of Plate 9, 2

Figs. 1, 2, 9 LV (GPIMH 2472, 880 yam long): fig. 1, ext. lat . ; fig. 2, ext. vent. obi.
Scale A (100 /xm; x 110), figs. 1, 2.

■ft

- n

J.U

T + +

- + + -1
- + + H

-1-1-1

+ + -t

Stereo-Atlas of Ostracod Shells 9, 3

Braderupia asymmetrica (3 of 8)

Remarks: B. asymmetrica was formerly questionally placed within the ctenonotellid genus Pseudostrepula
(Schallreuter, Palaeontographica (A) 153 (4/6), 194, 1976; V. A. Ivanova, Trudy Paleont. Inst.,
172, 158, 1979), but because of the confirmed presence of a histium the species is here assigned to
the Tetradellidae.il. asymmetrica is a sigmoopsine because it has a dimorphic histium which, more-
over, is also present anteroventrally and anterocentrally as, for example, in Sigmoopsis (see below).

The only other unisulcate sigmoopsine genera are Severobolbina Schallreuter (Geol. For.
Stockh. Fork. 96 (3 = 558), 278, 1974) and Valdarella Qvale ( Norsk Geol. Tidsskr., 60 (2), 102,
(1980). In contrast to Braderupia, in Severobolbina the histium is also present centroventrally in
females (where it is confluent with the velar dolon) but is missing in males and in the anterior regions
of both dimorphs. Thus, Braderupia represents a different phylogenetic line leading to unisulcate
members of the Sigmoopsinae and descends presumably from Sigmoopsis- like forms in which the
histium is also present anteriorly as, for example, in S. rostrata (Krause) (Schallreuter, Geologie
15 (7), pi. 4 (p.873), fig. 4, pi. 5 (p.875), fig. 5). A third lineage is represented by Valdarella, which
may have descended from Kiesowia (Carinobolbina)- like forms.

Valdarella resembles Braderupia by having histial and velar ridges parallel to each other and
to the anteroventral margin in the females, but differs by the absence of a histium and velum in
tecnomorphs and in the development of these sculptures in the anterior centroventral region of
the females.

Ullerella ventr optical a Henningsmoen (Norsk. Geol. Tiddsskr. 32 (1), 47, 1953) resembles
Braderupia asymmetrica in certain respects but is not placed within the new genus because of the
presence of a ridge in front of the velum in the anterior part of the valve (Henningsmoen, loc. cit.,
text-fig. 1 lower).

Explanation of Plate 9, 4

Fig. l.d'LV, ext. lat. (GPIMH 2473, 820 /xm long); fig. 2, juv. car., ext. vent. (GPIMH 2474, 510/xm long).
Scale A (100 /xm; x 117), fig. 1; scale B (100 /xm; x 192), fig. 2.

•rr-

■rr

.XL.

i Stereo-Atlas of Ostracod Shells 9, 2

i

Braderupia asymmetrica (2 of 8)

u

r

-l_ J_U-
I

- -1 4

Stereo-Atlas of Ostracod Shells 9, 4

Braderupia asymmetrica (4 of 8)

t-r

i i

l-

T

I i i
-XXJ.

T-l — t-

Stereo-Atlas of Ostracod Shells 9,

Braderupia asymmetrica (5 of 8)

Braderupia asymmetrica (Neckaja, 1958)

1958

1959

1960

1970

1971

1971

1973

1973

1976

1976

1976

1979

1979

1980

Pseudostrepula asymmetrica sp.n. A. I. Neckaja, Trudy vses. neft. nauchno-issled. geol.-razv. Inst. (VNIGRI) 115 (= Mikrofauna
SSSR 9), 352, 353, pi. 1, figs. 8, 9.

Pseudostrepula asymmetrica Neckaja; L. I. Sarv, Eesti NSV Tead.Akad. Geol. inst. uurimused 4, 96-98, 193, tab. 2 (187),
pi. 16, figs. 1 1-16, text-fig. 10G.

Pseudostrepula asymmetrica Neckaja; L. I. Sarv, Ibid. 5, tab. 1.

Pseudostrepula asymmetrica Neckaja; A. Roomusoks, Stratigrafija viruskoj i char' juskoj serij (ordovik) Severnoj Estonii I,
216, 236, 267, 268, 289, 291, tabs. 12, 13, 15 (220, 246, 296).

Pseudostrepula asymmetrica; R. E. L. Schallreuter, Neus Jb. Geol. Palaont., 1971 (4), 250.

Pseudostrepula asymmetrica Neckaja; R. E. L. Schallreuter, Ibid., 1971 (11), 691.

Pseudostrepula (resp. Psuedostepula) asymmetrica Neckaja; A. E Neckaja, Trudy VNIGRI 324, 65, 66.

Pseudostrepula asymmetrica Neck.; L. K. Gailfte, Problemy regionalnoj geologii Pribaltiki i Belorussii, 67, tab. 2 (68).
Pseudostrepula ? asymmetrica Neckaja; R. E. L. Schallreuter, Palaeontographica (A) 153 (4/6), 194, 198.
Pseudostrepula asymmetrica Neckaja; N. Sidaraviciene, Sovet. geol. 1976 (8), 54, tab. 1 (50).

Pseudostrepula asymmetrica (Neckaja); V. Jaanusson, The Ordovician System (Proc. Palaeont. Assoc, symp. Birmingham
Sept. 1974; Ed. M. G. Bassett), text-figs. 10, 11 (faunal logs).

Pseudostrepula asymmetrica Neckaja; N. Sidaraviciene, Eesti NSV Tead. Akad. Toimetised (Geol), 28 (4), 133, text-figs. 2,
3, 4 (faunal logs).

? Pseudostrepula asymmetrica (& asimmetrica) Neckaja; V. A. Ivanova, Trudy paleont. Inst., 172, 158, 159, 191, pi. 13,
fig. 3.

Pseudostrepula asymmetrica Neckaja; N. Sidaraviciene, Eesti NSV Tead. Akad. Toimetised (Geol.), 29 (4), text-fig. 1
(faunal log).

Explanation of Plate 9, 6

Figs. 1, 2, 9 RV (GPIMH 2475, 870/cm long): fig. 1, ext. lat., fig. 2, ext. vent. obi.
Scale A (100 /cm; x 109), figs. 1, 2.

- -t-r — i
--trH

4_L_)

+ +--t

Stereo-Atlas of Ostracod Shells 9, 7

Braderupia asymmetrica (7 of 8)

Holotype: Vsesojuznyj neftjanoj naucno-issledovatel’skij geologorazvedocnyj institut (VNIGRI) Leningrad,
no. 3- 128, 9 RV.

Type locality: Raion Bolsie Korcany, Leningrad obi., Russia; lat. 59° 33' N, long. 29° 2' E. Viru Series (Middle
Ordovician), Gubkov beds = Schundorov Substage of the Idavere Stage (C3/S).

Figured specimens: Geologisch-Palaontologisches Institut und Museum, University of Hamburg (GPIMH) nos. 2472
(9 LV: PI. 9, 2, figs. 1, 2), 2473 (d'LV: PI. 9, 4, fig. 1), 2474 (juv. car.: PI. 9, 4, fig. 2), 2475 (9 RV:
PI. 9, 6, figs. 1, 2), 2476 (cfRV: PI. 9, 8, figs. 1, 2). From the Upper Viruan (middle Ordovician)
Hornstein erratic boulders no. Sy 52 (2475) and no. Sy 108 (2472, 2473, 2474, 2476) of the
Kaolinsand (Pliocene-Pleistocene), near Braderup, Isle of Sylt (N Frisian Is., N Sea) Germany;
lat. 54° 56' N, long. 8° 21' E; coll, by Ulrich von Hacht in 1978 and 1980.

As for genus.

In the development of puncta in the histiovelar furrow B. asymmetrica resembles Sigmoopsis (S.)
granulata (Sarv) and S. (Sigmoopsoides) sigmoopsoides Schallreuter.

Distribution: NW Russian Platform (Leningrad, Estonia, Latvia, Lithuania): Idavere (C3), Johvi (D,) and
Keila (D2) stages of the Viru Series, middle Ordovician. Rollsteinkalk (= Macrouruskalk) erratic
boulders of Keila age of northern Germany; upper Viruan Hornstein erratic boulders of the
Kaolinsand (Pliocene - Pleistocene) of the Isle of Sylt (N Frisian Is., N Sea), Germany.

Diagnosis:

Remarks:

Explanation of Plate 9, 8

Figs. 1, 2, c/RV (GPIMH 2476, 790/cm long): fig. 1, ext. lat.; fig. 2, ext. vent. obi.
Scale A (100/cm; x 122), figs. 1, 2.

-Tf

•TP

.XL.
I I

lx.

LLL.
I

IT
I I
I I
I I
-14

i-T-r-

Stereo-Atlas of Ostracod Shells 9, 6

Braderupia asymmetrica (6 of 8)

u

r

\-

+-+

Stereo-Atlas of Ostracod Shells 9, 8

Braderupia asymmetrica (8 of 8)

t-r
i i
i i
i i
Li

j±

-4-4

I I I
-414.

Stereo- Atlas of Ostracod Shells 9 (2) 9 - 16 (1982)

595.336.13 (113.312) (492.71 : 161.008.54) : 551.35 + 552.55

Bilobatia serralobata (1 of 8)

ON BILOB ATIA SERRALOBATA SCHALLREUTER

by Roger E. L. Schallreuter
(University of Hamburg, German Federal Republic)

Genus Bilobatia Schallreuter, 1976

Type-species (by original designation): Bilobatia serralobata Schallreuter, 1976

Diagnosis: Median-sized palaeocope, adults 1-2 mm long; domicilium longer in males than females. A well
developed oblong lobe occurs either side the main sulcus (S2), each having a row of short spines
posterolaterally. Behind the strongly developed L3 there is a shallow S3 and a flat, indistinct L4.
Velum in males represented by a row of spines or possibly as a flange. Female has a broad, convex,
flange-like dolon ad jacent to a well developed laterovelar furrow; dolon has a row of long peripheral
spines more of less perpendicular to the contact plane, thus forming a typical wehrliine antrum.
Marginal sculpture formed by a row of spines. Lateral surface more or less distinctly reticulate;
dolonal flange striated.

Remarks: Bilobatia is characterized by its two distinct lobes and its prominent laterovelar furrow. Its phylo-
genetic origin is unknown. Presumably it descended from a smaller quadrilobate ancestor with a
distinct L2. The stratigraphically older Pectidolon is larger, has a strong LI and a relatively weak
L2 and, therefore, could not be its ancestor.

Explanation of Plate 9, 10

Figs. 1, 2, $ LV (GPIMH 2493, 961/xm long excluding posterior spines): fig. 1, ext. lat. ; fig. 2, ext. vent. obi.
Scale A (100p.m; x 92), figs. 1, 2.

-t+-i

+++

444

-|-|-

+ + +

Stereo-Atlas of Ostracod Shells 9, 1 1

Bilobatia serralobata (3 of 8)

Remarks (contd.): The special type of dolon of Bilobatia, with a row of spines forming the outer antral fence, is typical
of the wehrliine type of antral dimorphism which was first described by Schallreuter (Ber. geol. Ges.
D.D.R., 10 (4), 484, 1965). This special kind of antral dimorphism is confined to the subfamily
Wehrliinae and occurs in Rakverella, Wehrlia, Pectidolon and Bilobatia. Kesling (Contr. Mus.
Paleont. Univ. Mich., 12 (13), 1955) first described females of Rakverella ? bonnemai Opik and
noted in his description of the “false pouch of female”: “Each frill consists of 29 or 30 spines, closely
set and apparently fused only along their lines of juncture. . . . The spines composing the false pouch
in Rakverella bonnemai are not as strongly fused as those in Piretella acmaea Opik” (op. cit.,
265-266). The spines do not fuse at all (e.g. Kesling 1955, pi. 1, figs. 1, 2, 5, 6); there is a system of
intervening gaps that is characteristic of wehrliine dimorphism. The function of such a special kind of
a brood pouch is possibly to act like a cage, to hold the brood or eggs together and at the same time
to supply them with fresh water.

Bilobatia serralobata Schallreuter, 1976

1965

1976

? 1976

1979

Ctenonotella bidens Sarv, 1959; R. E. L. Schallreuter, Ber. geol. Ges. D.D.R., 10 (4), 484.

Bilobatia serralobata n. sp. R. E. L. Schallreuter, Palaeontographica (A) 153 (4/6), 205-207, pi. 8, figs. 1, 2, text-fig. 14,
tab. 12 (q. v. for full synonymy).

Ctenonotella bidens (Krause); V. Jaanusson, The Ordovician System (Proc. Palaeont. Assoc, symp. Birmingham Sept. 1974
Ed. M. G. Bassett), text-fig. 1 1 (faunal log).

Ctenonotella bidens (Krause, 1892); V. A. Ivanova, Trudy paleont. Inst. 172, 141-142(pars), pi. 10, fig. 12 (holotype).

Explanation of Plate 9, 1 2

Figs. 1 , 2, 9 RV (GPIMH 2494, 937/xm long): fig. 1, ext. lat.; fig. 2, ext. anterovent. obi.; fig. 3, incomplete 9 RV, ext. posterovent. obi.
(GPIMH 2495, 543/u.m high).

Scale A (100 /urn; x 97), figs. 1, 2; scale B (100/xm; x 85), fig. 3.

.XL.
i l

I I I
Lli.
I I i
I I
I I
I I

■t— H--
I I
I I
I I

Stereo- Atlas of Ostracod Shells 9, 10

Bilobatia serralobata (2 of 8)

-M-

1T

IT

-LLL

-14

I-

-T

r

Stereo-Atlas of Ostracod Shells 9, 12

Bilobatia serralobata (4 of 8)

I I ,
— i — J— !-

i i

i i

i i

i i

1

Stereo- Atlas of Ostracod Shells 9, 13

Bilobatia serralobata (5 of 8)

Holotype: Geologiceskij musej Akademija nauk Estonskoj SSR Tartu no. Os 2210, 9 RV.

Type locality: Alliku, SW of Tallinn, Estonia; lat. 59° 21' N, long. 24° 33' E. Johvi Stage ( D , ), upper Viruan
middle Ordovician).

Figured specimens: Geologisch-Palaontologisches Institut und Museum, University of Hamburg (GPIMH) nos. 2493
(9LV: PI. 9, 10, figs. 1, 2), 2494 (9 RV: PI. 9, 12, figs. 1, 2), 2495 (9RV: PI. 9, 12, Fig. 3), 2496
(d'LV: PI. 9, 14, figs. 1, 2), 2497 (rfLV: PI. 9, 16, fig. l)and 2498 (cfLV: PI. 9, 16, fig. 2). All from
the upper Viruan (middle Ordovician) Hornstein erratic boulder no. Syl08 of the Kaolinsand
(Pliocene - Pleistocene), near Braderup, Isle of Sylt (N Frisian Is, N Sea), Germany; lat. 54° 56' N,
long. 8° 21' E; coll, by Ulrich von Hacht in 1980.

Diagnosis: As for the genus.

Remarks: The holotype of B. serralobata , a female right valve, is 1.25 mm long according to L. I. Sarv (Eesti
NSV Tead. Akad. Geol. Inst, uurimused 4, 73, 1959). The females from boulder Syl08 are much
smaller (0.90-0. 96mm) and, perhaps, this population represents a smaller sub-species. The males
are considerably larger than the females (c. 0.98mm long without velum; domicilium of the females:
0.77-0.80 mm). Therefore, the 3 largest larvae from Backsteinkalk boulders plotted in the diagram
(text-fig. 14) and table 12 of Schallreuter (1976, op. cit.) presumably represent males and not
stage II, as was assumed at that time, when no complete females were available from those boulders.

Explanation of Plate 9, 14

Fig. 1, 2, d LV (GPIMH 2496, 984|U,m long excluding spines): fig. 1, ext. lat.; fig. 2, ext. vent. obi.
Scale A (lOOjum; x 93), figs. 1, 2.

1J_

-FT- (
T + H

J4J

-H-4

.XL.

i I I

Stereo- Atlas of Ostracod Shells 9, 1 5

Bilobatia serralobata (7 of 8)

Remarks (contd.); Based on a supposed d fragment from the Rollsteinkalk boulder Ro2 it was formerly stated (Schall-
reuter 1976, op. cit., 205) that the males possess a velar flange. The figured specimen herein shows
that the velum in males could also be developed only as a row of spines (PI. 9, 14, figs. 1, 2).

The posteriorly directed spines on the top of the central lobes are, on the figured specimens,
distinct only on the posterior lobe, but they also occur on the anterior lobe (see Schallreuter 1976,
op. cit., pi. 41 (8), fig. 1). Sometimes, next to the spines on the top of the lobes a row of fine pores
occurs (PI. 9, 12, fig. 3, PI. 9, 14, fig. 2, PI. 9, 16, fig. 2).

The lectotype of Ctenonotella ? bidens (Krause) is 0.83 mm long without velum (A. Krause,
Z. Deutsch. geol. Ges., 44 (3), 396 1892), and in this respect is of about the same size as the figured
females of B. serralobata. In contrast to B. serralobata the flange in C. ? bidens reaches the cardinal
corner and does not extend peripherally into spines (A. Krause, op. cit., pi. 22, fig. 12). If these
features are the result of incorrect drawing C. ? bidens could possibly be a senior synonym of B.
serralobata. If the lectotype of C.? bidens is not a female valve it is distinguished from the males of
the figured taxon by the velar flange.

Distribution: Northern Estonia; Johvi (D,) Stage and Keila Stage (D2), middle Ordovician. Backsteinkalk
erratic boulders (14B2 type) and Rollsteinkalk (Macrouruskalk) erratic boulders (of Keila age)
of northern Germany. Upper Viruan (middle Ordovician) Hornstein erratic boulders of the
Kaolinsand (Pliocene-Pleistocene) of the Isle of Sylt (N Frisian Is, N Sea), Germany.

Explanation of Plate 9, 1 6

Fig. 1, 9 LV, int. lat. (GPIMH 2497, 927 yu. m long excluding acroidal spine); fig. 2, 9 RV, ext. lat. (GPIMH 2498, 902^im long).
Scale A (100/xm; x 100), figs. 1, 2.

I ' I

I I I

Lli

I i I

I I I

I I I

I I I

1 t" T

I I I

I I I

I I I

I I I

Stereo- Atlas of Ostracod Shells 9, 1 4

Bilobatia serralobatu (6 of 8)

— h i-

i-

r

J_L
I

-14

1-1— -
I
I
I

l_

-+-++-

Stereo-Atlas of Ostracod Shells 9, 1 6

Bilobatia serralobata (8 of 8)

( ' t

Stereo- Atlas of Ostracod Shells 9 (3) 17 - 20 (1982)

595.336.13 (113.313) (261.24 : 161.018.57 + 261.23 : 161.005.54) : 551.35 (26.03)

Byrsolopsina manca (1 of 4)

ON BYRSOLOPSINA MANCA SCHALLREUTER sp. nov.

by Roger E. L. Schallreuter
(University of Hamburg, German Federal Republic)

Byrsolopsina manca sp. nov.

Holotype: Geologisch-Palaontologisches Institut und Museum, University of Hamburg, no. 2519, RV.
[Paratypes: nos. 2520-2525].

Type locality: Ojlemyrflint erratic boulder (no. G30), Isle of Gotland (Baltic Sea), beach opposite the Isle of Lilia
Karlso; lat. 57° 18'N, long. 18° 8' E.; Upper Ordovician .

Derivation of name: Latin, mancus, meaning incomplete, imperfect; alluding to the incompletely reticulated lateral
surface.

Figured specimens: Geologisch-Palaontologisches Institut und Museum, University of Hamburg (GPIMH) nos. 2519
(RV: PI. 9, 18, fig. 1), 2520 (RV: PI. 9, 18, fig. 2), 2521 (LV: PI. 9, 20, fig. 1) and 2526 (LV: PI. 9,
20, fig. 2). From the type locality, boulder no. G30 (nos. 2519-2521), coll, by the author in 1976,
and from Ojlemyrflint erratic boulder no. Sy56 (no. 2526) of the Upper Kaolinsand (Lower
Pleistocene, near Braderup, Isle of Sylt (N Frisian Is, N Sea); lat. 54° 56' N, long. 8° 21' E, coll,
by Ulrich von Hacht in 1977.

Explanation of Plate 9, 1 8

Fig. 1, RV, ext. lat. (holotype, GPIMH 2519, 612|U,m long); fig. 2, RV, ext. lat. (paratype, GPIMH 2520, 639p.m long).
Scale A (100 /Am; x 140), figs. 1, 2.

HJ

T + +

- -t-r — i

- + + +

-L —I

+ + 4

Stereo- Atlas of Ostracod Shells 9, 1 9

Byrsolopsina manca (3 of 4)

Diagnosis: Species of Byrsolopsina with valves up to 0.64mm long and moderately high (length:height ratio
1.75-1.55). Hinge-line short, cardinal angles distinctly greater than 90°. Lateral surface has fine,
scattered puncta; punctate area generally tapering in an anterior direction.

Remarks: Of all the species of Byrsolopsina Swain & Cornell (in Swain et al., J. Paleo. 35 (2), 363, 1961),
B. manca most resembles B. elkrunensis Swain (Ibid. 36 (4), 725, pi. 109, fig. 2, 1962), particularly
in the fine, incomplete punctation of its lateral surface. B. manca is distinguished from#, elkrunensis
mainly by its shorter hinge-line and correspondingly larger cardinal angles.

Compared with/?, manca , B. irregularis (Keenan) (J. Paleo. 25 (5), 562, 1951) has smaller
cardinal angles and a different pattern of distribution of puncta (Keenan 1951, pi. 78, fig. 34;
Copeland, Bull. Geol. Surv. Canada 187, pi. 4, fig. 2, pi. 5, figs. 4, 8, 1970).

Parapyxion sp. of Gailite [ Fauna i stratigrafija paleozoja i mesozoja Pribaltiki i Belorussii
(The Fauna and Stratigraphy of Palaeozoic and Mesozoic of Baltic and Byelorussia) , 63, 1975]
is possibly a congeneric form. Compared to B. manca it is larger, relatively higher, more elliptical
and punctate only in the dorsocentral region.

Distribution: Known from the Ojlemyrflint (Upper Ordovician) erratic boulders of the Isle of Gotland (Baltic
Sea) and of the Upper Kaolinsand (Lower Pleistocene) of the Isle of Sylt (N Frisian Is, N Sea).

Explanation of Plate 9, 20

Fig. 1, LV, ext. lat. (paratype, GPIMH 2521, 640 /mi long); fig. 2, LV, ext. lat. (GPIMH 2526, 545/im long).
Scale A (100 /un; x 145), fig. 1; scale B (100 /im; x 165), fig. 2.

-Tf-

'TP

L-L.L

1 1“ T

Stereo-Atlas of Ostracod Shells 9, 1 8

i i

-U

-i-r

Byrsolopsina mane a (2 of 4)

I— J- L-

I

I

I

I

Stereo-Atlas of Ostracod Shells 9, 20

-14

I

H

T

Byrsolopsina manca (4 of 4)

I I

A-'u

I I I
-xij.

r + -t

Stereo- Atlas of Ostracod Shells 9 (4) 21-26 (1982)

595.337.14 (119.9+ 119.1) (261.27 : 162.006.54 + 420.162 : 001.50) : 551.35

Loxoconcha cuneiformis (1 of 6)

ON LOXOCONCHA CUNEIFORMIS MALCOLMSON

by David J. Horne and Eric Robinson
(City of London Polytechnic and University College, London, England)

Loxoconcha cuneiformis Malcolmson, 1886

1 886 Lopoconcha cuneiformis (sic), n. sp. Brady MS; S. M. Malcolmson, Rep. Proc. Belf. Nat. Fid. Club, appendix 1884-85, 261,
pi. 25, figs. 1 , 2.

Holotype: A male carapace (split into valves and remnants of soft parts dissected); G. S. Brady collection,
Hancock Museum, Newcastle-upon-Tyne, no catalogue number but placed in a separate, labelled
slide.

Type locality:
Figured specimens:

Rockport, County Down, Ireland, approx, lat. 54° 39' N, long. 5° 46' W; intertidal. Recent.

Brit. Mus. (Nat. Hist.) nos. OS 12135 (rfRV: PI. 9, 22, fig. 1), OS12136 (d'LV: PI. 9, 22, fig. 2),
OS 12137 (? car.: PI. 9, 22, fig. 3), OS 12138 (c/LV: PI. 9, 24, fig. 1), OS 12139 (t/RV: PI. 9, 24,
figs. 2, 3, 4). Hancock Museum specimen (holotype: Text-figs. 1, 2). With the exception of the
holotype, all specimens were collected by E. Robinson from Pleistocene (Ipswichian) marine clay in
channels cut into Tertiary deposits on the foreshore at Selsey, Sussex, S England (approx, lat.
50° 47' N, long. 0° 50' W).

Explanation of Plate 9, 22

Fig. l.d' RV, ext. lat. (OS 12135, 550/u.m long); fig. 2,d' LV, ext. lat. (OS 12136, 560 /xm long); fig. 3, d car., ext. 1. lat. (OS 12137,
500 /urn long).

Scale A (100/xm; x 1 10), figs. 1-3.

++-i
+ + H

_l_l _J

-7-n-

+ 4

Stereo-Atlas of Ostracod Shells 9, 23

Loxoconcha cuneiformis (3 of 6)

Diagnosis: Carapace surface finely pitted, with ghost reticulation. Cuneate in dorsal view, with maximum
width in front of mid-length. Strongly dimorphic, male markedly more elongate than female.

Remarks: Malcolmson sent his only specimen of this species to G. S. Brady, in whose collection it remains to
this day. Brady recognised it to be a new species and gave it the manuscript name L. cuneiformis;
it was, however, Malcolmson, not Brady, who published the type description.

Brady and Norman (Scient. Trans. R.Dubl. Soc., ser. 2, 4, 186, 1889) regardedL. cuneiformis
as conspecific with Loxoconcha tamarindus (Jones) (sensu Brady (1868 ) = Lindisfarnia laevata
(Norman) - see Horne & Kilenyi, Stereo-Atlas of Ostracod Shells, 8, 107-116, 1981). From our
examination of the holotype it is clear that L. cuneiformis is indeed a valid species, easily distinguished
from related species by its cuneate outline in dorsal view.

Apart from the presumed Recent holotype, we have only seen Pleistocene specimens of this
species. Malcolmson’s specimen contained a few poorly-preserved remnants of appendages, but
this does not exclude the possibility that it may be a Pleistocene relict; we have found appendages
in undoubted Pleistocene specimens of other species, obtained from borehole material.

Distribution: Pleistocene and Recent(?): British Isles (herein).

Explanation of Plate 9, 24

Fig. l.d'LV, ext. dors. (OS 12138, 540 pim long); figs. 2, 3, cfRV, ant. and post, hinge elements (OS 12139); fig. 4, cfRV, int. muse. sc.
(OS 12139).

Scale A (100/xm; x 1 10), fig. 1; scale B (50/u.m; x 400), figs. 2-4.

—I— LI—

I I I

L-LJ__

— t~t~r-

Stereo- Atlas of Ostracod Shells 9, 22

Loxoconcha cuneiformis (2 of 6)

Stereo-Atlas of Ostracod Shells 9, 24

n .

i i
i i
i i

14

I I
I I
I I
1 I

M-

I I

Loxoconcha cuneiformis (4 of 6)

l i
l I

i

-

r

H

r

t-r

i

i

r

1 1

I I I
. L_Li.
I

' 1 — I-

Stereo- Atlas of Ostracod Shells 9, 25

Loxoconcha cuneiformis (5 of 6)

Text-fig. 1.

a: holotype. d RV, drawn in transmitted light; b-d:
remnants of appendages from holotype : b, c, parts
of copulatory appendage; d, mandible.

i-+-t—

lOOju.m

L-i-L.-

t- + -

Stereo-Atlas of Ostracod Shells 9, 26

Loxoconcha cuneiformis (6 of 6)

-t-r-

-f-i'

i i

i i

tr

i i
i i

Stereo-Atlas of Ostracod Shells 9 (5) 27-32 (1982)
595.337.14 (119.9) (261.2 : 162.017.28) : 551.35

Loxoconcha dimorpha (1 of 6)

T-t-t-

ON LOXOCONCHA DIMORPHA HARTMANN

by David J. Horne

(City of London Polytechnic, England)

Loxoconcha dimorpha Hartmann, 1959

1959

Loxoconcha dimorpha n. sp. G. Hartmann, Zool. Anz., 162 (pars), 1 63-166, text-figs. 12, 13, 16. 17, 1 8 only, (^cm text-figs.

10, 11, 14, 15, 20).

Lectotype: (here designated). A decalcified male carapace with appendages (Paralectotype, a decalcified
female carapace with appendages; syntypes, no. K-28132). All deposited in the Zoologisches
Institut und Zoologisches Museum, Universitat Hamburg.

Type locality .Tenerife, Canary Islands (approx, lat. 28° 15' N, long. 16° 35' W); intertidal. Recent.

Figured specimens: Brit. Mus. (Nat. Hist.) nos. 1982.32 (c/LV: PI. 9, 28, fig. 1; PI. 9, 30, figs 1, 5), 1982.33 (9 LV:
PI. 9, 28, fig. 2; PI. 9, 30, fig. 3), 1982.34 (9 RV: PI. 9, 28, fig. 3), 1982.35 ( c/LV + appendages:
PI. 9, 30, fig. 2; Text-figs, la-e, 2b), 1982.36 (9 LV: PI. 9, 30, fig. 4). Hamburg Museum specimens
(lectotype, cf copulatory appendage: Text-fig. 2a), K-28132 (syntype. d RV: Text-fig. 2c). Nos.
1982.32-36 collected alive from intertidal algae at El Medano, Tenerife, Canary Islands (approx,
lat. 28° 15' N long. 16° 35' W) by M. Linley in 1980. Hamburg Museum specimens from Hartmann’s
syntypic material.

Explanation of Plate 9, 28

Fig. 1, cfLV, ext. lat. (1982.32, 580/zm long); fig. 2, 9LV, ext. lat. (1982.33, 490 pm long); fig. 3, 9 RV, ext. lat. (1982.34, 480 pm long).
Scale A (100 /am; x 110), figs. 1-3.

-tt-i

T-H

+ +--+

Stereo-Atlas of Ostracod Shells 9, 29

Loxoconcha dimorpha (3 of 6)

Diagnosis: Carapace robust, strongly ornamented, with pitting in central area giving way to reticulation in
marginal areas. Sexual dimorphism marked, male more elongate than female. In dorsal view,
greatest width behind mid-length. Male copulatory appendages of typical Loxoconcha shape: basal
part broadly oval and about twice the size of the sub-triangular head-piece which has a convex
ventral margin and a rounded, slightly upturned distal (anterior) corner.

Remarks: A comparison of specimens of two species of Loxoconcha from Tenerife with the original description
and figures of L. dimorpha suggested that Hartmann (op. cit.) had inadvertantly combined the male
of one species with the female of the other and described them as a single species. This view was
confirmed by the original author (Hartmann, pers. comm.) and by a re-examination of his syntypic
material, which was found to include adult male specimens corresponding to the type description
and figures of the male of L. dimorpha, but adult females of two species. One female form, corres-
ponding to Hartmann’s original description and figures of the female of L. dimorpha, belonged to a
species described elsewhere as new (Horn e, Stereo-Atlas of Ostracod Shells, 9 (6), 33-40, 1982); the
other matched those illustrated herein as the true female of L. dimorpha.

A male and a true female of L. dimorpha, selected from the syntypes and dissected, are
designated herein as lectotypes.

Minor differences between the male copulatory appendages of the type specimens and
those of the more recently obtained examples (see text-figs. 2a-b) are regarded as intraspecific
variations, and may even be due to disturbance of the appendages during dissection.

Distribution: Recent: known only from the intertidal zone of Tenerife (Hartmann, op. cit., and herein).

Explanation of Plate 9, 30

Fig. 1 , c/LV, ext. dors. (1982.32, 580/i.m long); fig. 2, cfLV, int. lat. ( 1982.35, 590 p.m long); fig. 3, 9 LV, ext. dors (1982.33, 490 pm
long); fig. 4, 9 LV, int. lat. ( 1982.36, 500 pm long); fig. 5, 6 LV ext. lat., detail of dorsal marginal area showing ornament and sieve
pores, (1982.32).

Scale A (100/u.m; x 110), figs. 1-4; scale B (25/u.m; x 700), fig. 5.

-t-l-
l l
I I
I I

I I I
.Lll.

Stereo- Atlas of Ostracod Shells 9, 28

i

J

i-

Loxoconcha dimorpha (2 of 6)

l-t-

hi--!--

— J_ I —
I

f- +

Stereo-Atlas of Ostracod Shells 9, 30

-14

Loxoconcha dimorpha (4 of 6)

t -r

i—

r

I 1 I
I I I
. Lii.

I I I

I I I
I I I
I I I

"+T"

Stereo- Atlas of Ostracod Shells 9, 3 1

Loxoconcha dimorpha (5 of 6)

-t- +

-+-•+ + -

Stereo- Atlas of Ostracod Shells 9, 32

Loxoconcha dimorpha (6 of 6)

Text-fig. 2. a: d copulatory appendage (lectotype; Hamburg Museum specimen); b: cfcopulatory appendage (1982.35); c: cfRV
(decalcified) seen in transmitted light (syntype, Hamburg Museum no. K-28132).

-t-r-

- -I-L— I —
I I I

J-L.

Stereo-Atlas of Ostracod Shells 9 (6) 33-40 (1982)

595.337.14 (119.9) (261.2 : 162.017.28+ 162.017.32) : 551.35

Loxoconcha linleyi (1 of 8)

ON LOXOCONCHA LINLEYI HORNE sp. nov.

by David J. Horne

(City of London Polytechnic, England )

? 1855

1911

1959

Loxoconcha linleyi sp nov.

Cythere maculata sp. nov. S. Fischer, Abh. bayer. Akad. Wiss., 7, 656-658, PI. 2, figs. 9-12 ( = nomen dubium).
Loxoconcha impressa (Baird); G. S. Brady, Proc. zool. Soc. Lond., 27, 595 (pars).

Loxoconcha dimorpha sp. nov. G. Hartmann, Zool. Anz., 162, (pars), 163-166, text-figs. 10, 11, 14, 15, 20 only; ( non
text-figs. 12, 13, 16, 17, 18).

Holotype: A male carapace + appendages (split into valves and dissected), Brit. Mus. (Nat. Hist.) 1982.37.

[Paratypes: a female carapace, split into valves, 1982.38; and a male carapace + appendages,
split into valves and dissected, 1982.39].

Type locality: El Medano, Tenerife, Canary Islands, approx, lat. 28° 15' N, long. 16° 35' W; intertidal, Recent.

Derivation of name: After Mike Linley, who collected the sample in which this species was first recognised.

Figured specimens: Brit. Mus. (Nat. Hist.) nos. 1982.37 (holotype, d car.; RV: PI. 9, 34, figs. 1, 3; LV, PI. 9, 38, fig. 3),
1982.38 (9 car.; RV: PI. 9, 34, figs. 2, 4; LV: PI. 9, 38, fig. 2, PI. 9„40, fig. 1), 1982.39 (cfappendages:
Text-figs, la-e, 2a), 1982.40 (9 RV: PI. 9, 36, figs. 1. 2), 1982.41 (c/RV: PI. 9, 36, fig. 3), 1982.42
(?LV: PI. 9, 38, fig. 1), 1982.43 (9LV: PI. 9, 40, fig. 3), 1982.44 (c/RV: PI. 9, 40, fig. 2), 1982.45
(cf copulatory appendage: Text-fig. 2b). Nos. 1982.37-42 were collected alive from intertidal algae
at the type locality by M. Linley in 1980. Nos. 1982.43-45, taken from slide no. 1911.11.8. M3454
in the Norman Collection at the Brit. Mus. (Nat. Hist.), were collected by A. M. Norman between
tidemarks on the island of Madeira in spring, 1897, (approx, lat. 32° 45'N, long. 17° 00' W).

Explanation of Plate 9, 34

Figs. 1, 3, <3 RV (holotype, 1982.37, 560 ^m long), fig. 1, ext. lat.; fig. 3, ext. dors.; figs. 2, 4,9 RV (paratype, 1982.38, 510/um long),
fig. 2, ext. lat.; fig. 4, ext. dors. Scale A (100/xm; x 120), figs. 1-4.

I i i

. -L J l

r + -t

-r -l

+ +

-7-1-1-

- + + 4

Stereo- Atlas of Ostracod Shells 9, 35

Loxoconcha linleyi (3 of 8)

Diagnosis: Carapace rather quadrate in lateral view. Dimorphic, male more elongate and with a straighter
dorsal margin than the female. Finely pitted ornament becoming coarser posteriorly and developing
marginally into reticulation, especially in the posterodorsal region. In dorsal view, greatest width
at about mid-length. Male copulatory appendages of typical Loxoconcha shape: the broadly
triangular head-piece about two-thirds the size of the basal part, with an acutely rounded posterior
corner and a rounded, tooth-like process at the anterior (distal) corner.

Remarks: Female specimens of L. linleyi were erroneously described and figured by Hartmann (op. cit.) as
the female of Loxoconcha dimorpha Hartmann 1959 (see Horne 1982, Stereo-Atlas of Ostracod
Shells, 9, (5) 27-32, 1982). Specimens from Madeira in the Brit. Mus. (Nat. Hist.), collected by
A. M. Norman and identified by G. S. Brady as Loxoconcha impressa (Baird), include both Loxo-
concha rhomboidea (Fischer) (see Athersuch & Whittaker, Stereo-Atlas of Ostracod Shells, 3 (17),
81-90, 1976) and L. linleyi. The two species are indeed very similar, but L. linleyi is easily distin-
guished from L. rhomboidea by its more quadrate outline, its posterodorsal reticulation, and by
the detailed shape of the male copulatory appendages.

Cythere maculata Fischer 1855, described from Madeira, might possibly be conspecific with
L. linleyi. Unfortunately Fischer’s illustrations are so poor as to make a confident identification
impossible, and in the absence of any type specimens it must be regarded as nomen dubium.
Specimens of L. linleyi from Madeira are somewhat thicker-shelled, and the females have less
strongly arched dorsal margins, than those from the type locality; minor differences are also
apparent between the male copulatory appendages of specimens from the two islands (see text-figs.
2a-b). These variations are presently regarded as intraspecific.

Distribution: Recent: Tenerife and Madeira, intertidal (herein).

Explanation of Plate 9, 36

Figs. 1, 2, 9 RV (1982.40, 530^m long), fig. 1, int. lat.; fig. 2, int. muse, sc.; fig. 3, / RV, int. hinge (1982.41).
Scale A (lOO/^m; x 120), fig. 1; scale B (25/u.m; x 440), fig. 2; scale C (50^,m; x 240), fig. 3.

r

.iL.
I I

I I I

I I I
L-LX
I l I
I l I
I I
I I
l"T
I
I

Stereo-Atlas of Ostracod Shells 9, 34

J-i-

-i-r

Loxoconcha linleyi (2 of 8)

IT
I I
I I
I I
-14

1-1-
!- + +-•

■f+f

Stereo-Atlas of Ostracod Shells 9, 36

Loxoconcha linleyi (4 of 8)

l l
i l
H
I I
I I

r

V- |

. *

9

— I — J— !-

i i i
— L _L _1 .

Ti-“t

Stereo- Atlas of Ostracod Shells 9, 37

Loxoconcha linleyi (5 of 8)

Explanation of Plate 9, 38

Fig. 1, 9 LV, int. lat. ( 1982.42, 520^m long); fig. 2, 9 LV, ext. lat. (paratype, 1982.38, 520 /am long); fig. 3, cfLV, ext. lat. (holotype,
1982.37, 560 /am long).

Scale A (100 /am; x 120), figs. 1-3.

--t-t-i

--t-M

JiJ

4

Stereo-Atlas of Ostracod Shells 9, 39

Loxoconcha linleyi (7 of 8)

Text-fig. 2. cfcopulatory appendages, a: (paratype, 1982.39), from Tenerife; b: (1982.45) from Madeira.

Explanation of Plate 9, 40

Fig. 1, 9 LV, ext. lat., detail of central region showing sieve-pores (paratype, 1982.38); fig. 2, d RV, ext. lat. (1982.44, 580/am long),
fig. 3, 9 LV, ext. lat. (1982.43, 550/am long).

Scale A (10/am; x 950), fig. 1; scale B (100/am; x 120), figs. 2, 3.

—

,1L.
i I I

I I
I I

,L±j
I l
I I
I I

I 1 1

I I
I I
I I
I I
I I
I I
I
I
I
I

Stereo-Atlas of Ostracod Shells 9, 38

t-r-

Loxoconcha linleyi (6 of 8)

l-l-
— h 1-

- 1 — I— J —

— 1 — I — I —

Stereo- Atlas of Ostracod Shells 9, 40

Loxoconcha linleyi (8 of 8)

1 _1.

Stereo-Atlas of Ostracod Shells 9 (7) 41-46 (1982)

595.337.14 (1 18.21) (931-13 : 163.170.45) : 552.51

Waiparacythereis joanae ( 1 of 6)

ON WAIPARACYTHEREIS JOANAE SWANSON

by Kerry Swanson

(University of Canterbury , New Zealand )

Waiparacythereis joanae Swanson, 1969

1969

1979

1979

Waiparacythereis joanae K. M. Swanson, Trans. Roy. Soc. N.Z., Earth Sci. 7(3), 41, pi. 2, figs. 36-38.
Waiparacythereis joanae K. M. Swanson, N.Z. Oceanographic Inst. Memoir 78, 28, fig. 33.
Waiparacythereis joanae K. M. Swanson, N.Z. Journ. Marine & Freshwater Res. 13(1), 160, figs. 40-p.

Holotype: New Zealand Geological Survey no. TO 1009.

Type locality: M Waipara sequence, Canterbury Province, New Zealand; lat. 43° 0.5' S, long. 172° 35' E., 80ft
below the top of the Gowan Hill Sandstone. Grid. ref. S68 991121. L Miocene.

Explanation of Plate 9, 42

Fig. 1, RV, ext. lat. (UCF 1337.1, 880 /am long); fig. 2, LV, ext. lat. (UCF 1337.1, 900 long).
Scale A (200 /am long; x 97), fig. 1; scale B (200 /am long; x 93), fig. 2.

"t-r

■ J4J

-7-N-

- + H--t

Stereo- Atlas of Ostracod Shells 9, 43

Waiparacythereis joanae (3 of 6)

Figured specimens: University of Canterbury (Geol Dept) nos. UCF 1337.1 (d car.; LV and RV: PI. 9, 42, figs. 1-2),
UCF 1337.4 (o' car.; LV: PI. 9, 44, figs. 1-3), UCF 1337.2 ( d appendages: Text-figs, la-c, 2a),
UCF 1337.1 (d appendages: Text-figs. 2b-d). All from dredge sample ( 17.7m), Otago Shelf, South
Island, New Zealand; lat. 45° 55' S, long. 170° 36'E.

Diagnosis: Carapace large, elongate, strongly inflated posteriorly. Very heavily calcified. Surface ornamentation
subdued, especially medially.

Remarks: In my original description ( 1969), on the basis of carapace morphology (in particular muscle scars),
I felt Waiparacythereis to be closely related to Urocythereis. This decision is confirmed by the soft
parts illustrated herein.

Explanation of Plate 9, 44

Fig. 1 . FV, int. lat. (UCF 1337.4, 880 /on long); fig. 2, LV, int. muse. sc. and ant. hinge (UCF 1337.4, 880/am long); fig. 3, LV, int. post,
hinge (UCF 1337.4, 880 /am long).

Scale A (200 /am long; x 95), fig. 1; scale B (100/am long; x 215), fig. 2; scale C (50/am long; x 515), fig. 3.

■Tf-

•rr-

,1L.

J.J

I I I

1 LJ-.

I l
I I
l I
I I

tff-r-

i

Stereo-Atlas of Ostracod Shells 9, 42

W aiparacythereis joanae (2 of 6)

-I— M~

— IT

I I I
I I I

-Ltl

I I I
I I I
I I I
I I I

+--M-

I I I

Stereo-Atlas of Ostracod Shells 9, 44

i i i

W aiparacythereis joanae (4 of 6)

t-r

—IT

— I_L

.

I I I
. J L J..

I I

"+T-

Stereo-Atlas of Ostracod Shells 9, 45

Waiparacythereis joanae (5 of 6)

Text-fig. 1. Appendages: a: thoracic legs; h: antenna and antennule;
c: mandible.

-1-1-1-

-Li

- -I h

Stereo-Atlas of Ostracod Shells 9, 46

Waiparacythereis joanae (6 of 6)

-t-r-

Text-fig. 2. Appendages: a: copulatory apparatus; b: posterior of body;
c: brush organ; d: maxilla

--L1J

Stereo- Atlas of Ostracod Shells 9 (8) 47-50 (1982)
595.337.14 (1 18.21) (567 : 161.043.36) 551.35

Acanthocythereis dohukensis (1 of 4)

ON ACANTHOCYTHEREIS DOHUKENSIS KHALAF sp. nov.

by Saleh K. Khalaf

( University of Hull, England and University of Mosul, Iraq)

Acanthocythereis dohukensis sp. nov.

Holotype: University of Hull Coll. no. HU.275.T.1, d car.

[Paratype: HU.275.T.2.5].

Type locality: Dohuk anticline. Southern limb, 10 km NE Dohuk City N. Iraq. Lower Fars Formation bed no. 12;
lat. 36° 54' N, long. 43° 01' E; M Miocene.

Derivation of name: From Dohuk City, which gave its name to the Dohuk anticline, where the species was found.

Figured Specimens: University of Hull Coll. nos. HU.275.T.1 (d car.; LV: PI. 9, 48, fig. 1; PI. 9, 50, figs. 2, 3), HU.275.T.2
(9 car.: PI. 9, 48, fig. 2; PI. 9, 50, fig. 1).

Diagnosis: Species of Acanthocythereis with well-developed surface reticulation and strong blunt spines.
Carapace subrectangular with dorsal and ventral margins converging slightly posteriorly.

Explanation of Plate 9, 48

Fig. 1, cfcar., ext. It. lat. (holotype, HU.275.T. 1, 836/i.m long); fig. 2, 9 car. ext. rt. lat. (paratype, HU.275.T.2, 788/u.m long).
Scale A (200/xm, x 115), fig. 1; scale B (200|U,m; x 122), fig. 2.

--t-t-i
- + + -!

-T-n-

--H-4

Stereo-Atlas of Ostracod Shells 9, 49

Acanthocythereis dohukensis (3 of 4)

Remarks: Anterior margin broadly rounded with row of small tubercles, posterior end subtriangular. Sexual
dimorphism marked, presumed males are longer and less wide than the females.

Acanthocythereis hystrix differs in having much blunter spines and the posterior margin is
more subtriangular. Trachyleberis (Acanthocythereis) procapsus Siddiqui, 1971 is narrower and
less high, has a slightly concave ventral margin anteroventrally and differs in detail of reticulation.

Distribution: This species has been found in the L Fars Formation of the Bashiqa, Sheikhan and Dohuk sections,
N Iraq.

Explanation of Plate 9, 50

Fig. 1, 9 car., dors, (paratype, HU.275.T.2, 788/xm long); figs. 2, 3, dear. (holotype, HU.275.T.1, 836Ju,m long): fig. 2, dors.; fig. 3.
ext. rt. lat.

Scale A (200p.m; x 79), figs. 1, 2; scale B (200yu.m; x 133) fig. 3.

I I I
LJ-J.

I

I

I

Stereo- Atlas of Ostracod Shells 9, 48

i

J

i

j-

-i-r

Acanthocythereis dohukensis (2 of 4)

1-1-
h- M--

■-11

t-r

-4— M--

Stereo-Atlas of Ostracod Shells 9, 50

-i

Acanthocythereis dohukensis (4 of 4)

f-

T

I

-4-4

Stereo-Atlas of Ostracod Shells 9 (9) 51-54 (1982)
595.337.14 (118.21) (567 : 161.043.36) 551.35

Actinocythereis iraqensis (1 of 4)

ON ACTINOCYTHEREIS IRAQENSIS KHALAF sp. nov.

by Saleh K. Khalaf

( University of Hull, England and University of Mosul, Iraq)

Actinocythereis iraqensis sp. nov.

Holotype:
Type locality:

University of Hull Coll. no. HU.275.T.6, dear.

[Paratypes: HU.275.T.7-12].

Sheikh Ibrahim anticline, southern limb, NW Iraq, 45km west of Mosul. Lower Fars Formation
bed No. 50 M; lat. 36° 18' N, long. 42° 39' E; Miocene.

From its abundant occurrence in the M Miocene of Iraq.

University of Hull Coll. nos. HU.275.T.6 (<S car.; RV: PI. 9, 52, fig. 1), HU.275.T.7 (9 car.; RV:
PI. 9, 52, fig. 2; PI. 9, 54, fig. 2), HU.275.T.8 (9 LV, int. lat.: PI. 9, 54, fig. 1).

Diagnosis: A thick-shelled species of Actinocythereis, subrectangular in lateral view with greatest height at the
eye tubercle; surface strongly ornamented with different sized tubercles, well developed marginal
rim with small tubercles.

Derivation of name:
Figured specimens:

Explanation of Plate 9, 52

Fig. l.c/car., ext. rt. lat. (holotype, HU.275.T.6, 1092/u.m long); fig. 2, 9 car., ext. rt. lat. (paratype, HIJ.275.T.7, 916/xm long).
Scale A (200 p.m; x 90), fig. 1; scale B (200p,m; x 103), fig. 2.

tt
■ rr

nj

T+--t

4 4

-H-4

Stereo- Atlas of Ostracod Shells 9, 53

Actinocythereis iraqensis (3 of 4)

Remarks: The present species shows affinities vj'xth Actinocythereis tumefacentis (Lyubimova and Guha, 1960)
but differs in that the ventral margin of the latter species is strongly concave medially and the
posterior end is more rounded than that of A. iraqensis. This species differs from Trachyleberis
(Actinocythereis) birmanica pyawbwe Gramann, 1975 which is narrower posteriorly.

This species occurs abundantly and shows many of the features typical of Actinocythereis.
The dorsal and ventral margins are nearly straight and subparallel, the anterior end rounded with
distinct marginal rim and the posterior end is subrectangular. The surface is coarsely ornamented
with different sized tubercles, and the distinctive median row starts anteroventrally and runs in a
dorsally convex arch to the posteroventral part of the valve. This is a very common species in the
M Miocene of Iraq and is often accompanied by Hermanites transversicostata and Chrysocythere
naqibi.

Distribution: A. iraqensis is very common in the M Miocene of Iraq in the Bashiqa, Sheikhan, Dohuk, Sheikh
Ibrahim, Tel. Hajer, Kirkuk and Hamerin sections.

Explanation of Plate 9, 54

Fig. 1, $ LV, int. lat. (paratype, HU.275.T.8, 956/u.m long); fig. 2, cfcar., ext. dors, (paratype, HU.275.T.7, 916^m long).
Scale A (200 p,m; x 102), fig. 1; scale B (200 p,m; x 100), fig. 2.

.XL.

I I I
.Lll.
■ I
I
I
I

i Stereo- Atlas of Ostracod Shells 9, 52

Actinocythereis iraqensis (2 of 4)

rr

— t- i —

4- +

LJ-L-
I
I
I
I

+ -

T

I I
I I
I I
-14
I I
I I
I I
I I

-rr

Stereo-Atlas of Ostracod Shells 9, 54

Actinocythereis iraqensis (2 of 4)

t-r

Stereo- Atlas of Ostraeod Shells 9 (10) 55-58 (1982)
595.337.14 (118.21) (567 : 161.043.36) 551.35

Ckrysocythere naqibi (1 of 4)

ON CHRYSOCYTHERE NAQIBI KHALAF sp. nov.

by Saleh K. Khalaf

(University of Hull, England and University of Mosul, Iraq)

Ckrysocythere naqibi sp. nov.

Holotype: University of Hull Coll. no. HU.275.T.13, 9 car.

[Paratype: HU.275.T. 14, d'car.].

Type locality: Sheikh Ibrahim anticline, southern limb, NW Iraq, 145 km west of Mosul, Lower Fars Formation,
bed no. 50; M Miocene: lat. 36° 18' N, long. 42° 39' E.

Derivation of name: In the honour of the late Iraqi geologist, Khorshid M. Al. Naqib, in recognition of his extensive
contributions to the Iraqi geology.

University of Hull Coll. nos. HU.275.T.13 (holotype, 9 car.: PI. 9, 56, fig. 1; PI. 9, 58, fig. 1),
HU.275.T. 14 ( d car.: PI. 9, 56, fig. 2; PI. 9, 58, fig. 2).

Three longitudinal ridges, the dorsal and median are connected by a short curved transverse ridge
at about one third length from anterior end. The median ridge runs in an elegantly curved, convex
upward arc from middle of anterior margin towards middle of posterior end. The rest of the surface
is ornamented with thick, short transverse ridges.

Explanation of Plate 9, 56

Fig. 1. 9 car., ext. rt. lat. (holotype, HU.275.T.13, 904^01 long); fig. 2, d car., ext. It. lat. (paratype, HU.275.T.14, 992/u.m long).

Scale A (200/u.m; x 105), fig. 1; scale B (200/i.m; x 93), fig. 2.

Figured specimens:
Diagnosis:

-Lia

Ti--t

-tt-l
ft— I

JiJ

-M--+

Stereo- Atlas of Ostraeod Shells 9, 57

Ckrysocythere naqibi (3 of 4)

Remarks: The species is characterised by a distinctive type of ornamentation, the thick transverse ridges which
connect the median and ventral ridges forming distinctive reticulae; two rounded reticulae alternate
in the anterior part of the median ridge. Sexual dimorphism is very marked, the presumed male
being longer and narrower than the female.

The present species differs from Chrysocythere paradisus Doruk, 1973, which is more
elongate and differs in details of the reticulum.

C. naqibi is more elongate than C. cataphracta, Ruggieri, 1962, and differs in detail of the
reticulation.

The new subspecies of C. cataphracta described from the M Miocene in the Mersin of Turkey
by Bassiouni, 1980 shows some similarity in ornamentation but the present species differs in the
development of an additional longitudinal ridge below the ventral ridge which is joined to the
latter and follows a curved path in the posterior two-thirds of the shell.

Distribution: C. naqibi occurs in the M Miocene sections of Sheikh Ibrahim anticline, NW Iraq where it is
associated with Hermanites transversicostata Khalaf (Stereo-Atlas of Ostraeod Shells, 9, 59-62,
1982) and Actinocythereis iraqensis Khalaf (Ibid., 51-54).

Explanation of Plate 9, 58

Fig. 1, 9 car., ext. dors, (holotype, HU.275.T.13, 904/aiti long); fig. 2, cfcar., ext. dors, (paratype, HU.275.T.14, 992^i,m long).
Scale A (200 ^im; xl09), fig. 1; scale B (200/a.m; x 102), fig. 2.

-Tf

-Tf-

r-r.
i— m~

i i
i i
L-L.L
i i

i

i

i

1- + T
I
I
I

I

Chrysocy there naqibi (2 of 4)

Stereo- Atlas of Ostraeod Shells 9, 56

L-J-L

-14

t-

•(--(- +

Stereo-Atlas of Ostraeod Shells 9, 58

Chrysocythere naqibi (4 of 4)

i

-4-1-1

.J.1J

Stereo-Atlas of Ostracod Shells 9 (1 1) 59-62 (1982)
595.337.14 (118.21) (567 : 161.043.36) 551.35

Hermanites transversicostata (1 of 4)

ON HERMANITES TRANSVERSICOSTATA KHALAF sp. nov.

by Saleh K. Khalaf

(University of Hull, England and University of Mosul, Iraq)

Hermanites transversicostata sp. nov.

Holotype:
Type locality:

University of Hull Coll. no. HU.275.T. 15, cf car.

[Paratype: HU.275.T. 16-25],

Sheikhan anticline, northern limb, 27 km NE of Mosul City, Northern Iraq, Lower Fars Formation,
bed no. 10; M Miocene; lat. 36° 42' N, long. 43° 25' E.

Derivation of name: From the characteristic vertical transverse rib running from the posterodorsal margin about one-
eighth the length from the posterior margin and sub-parallel to the latter.

University of Hull Coll. nos. HU.275.T.16 (9 car.: PI. 9, 60, fig. 1; PI. 9, 62, fig. 1), HU.275.T.15
(holotype, d car.: PI. 9, 60, fig. 2; PI. 9, 62, fig. 2).

A species of Hermanites with curved dorsal ridge joined posteriorly with a short, vertical transverse
ridge and ending in a small node. Surface ornamented with strong reticulation, well-developed eye
tubercle and steeply inclined posterior margin.

Figured specimens:
Diagnosis:

Explanation of Plate 9, 60

Fig. 1,9 car., ext. It. lat. (paratype, HU.275.T.16, 824/xm long), fig. 2, d car., ext. It. lat. (holotype, HU.275.T.15, 992^tm long).
Scale A (200 jam; x 107), fig. 1; scale B (200/xm; x 94), fig. 2.

Ti-t

-t-f-f
-t + H

44

-7-rr

+ -!-

Stereo- Atlas of Ostracod Shells 9, 61

Hermanites transversicostata (3 of 4)

Remarks: Surface ornamented with well-developed reticulae, dorsal ridge joined by the short posteriorly
transverse ridge. Anterior end of the ventral ridge is joined to the sub-central tubercle by a short
curved ridge. The reticulae at the anterior margin are sub-rectangular.

Hermanites transversicostata shows affinities with Hermanites crucens Siddiqui 1971 but,
in the present species, the dorsal margin is humped and the ventral margin strongly concave just
behind the anterior margin. Furthermore, the posterior margin has the steeply inclined, short
posteriorly transverse ridge not seen in Siddiqui's species. H. transversicostata differs from H. sp.
cf H. paijenborchianus (Khosla 1978) in the shape of the posterior margin which is slightly concave
posterodorsally and more rounded posteroventrally; the present species is also much longer.

Distribution: The species is found in the M Miocene of Tel. Hajer, Sheikh Ibrahim, Sheikhan. Bashiqa, Kirkuk
and Hamerin sections.

It is very abundant and associated species are A. dohukensis, A. iraqensis and Chrysocythere
naqibi (see Khalaf, Stereo-Atlas of Ostracod Shells, 9, 47-50, 51-54, 55-58, 1981).

Explanation of Plate 9, 62

Fig. 1, 9 car., ext. dors, (paratype, HU.275.T.16, 824/xm long), fig. 2, d car., ext. dors, (holotype, HU.275.T.15, 992pm long).
Scale A (200/a.m; x 114), fig. 1; scale B (200/xm; x 86), fig. 2.

■TT‘

-Tf"

-ll_.
I I

I 1 I

I I 1
Lii

' i

I

J

I

— I — I —

— J_ J —

Stereo-Atlas of Ostracod Shells 9, 60

Hermanites transversicostata (2 of 4)

t-r

t~n--

Stereo-Atlas of Ostracod Shells 9, 62

Hermanites transversicostata (4 of 4)

:

k

!r

I I ,

-4 — L-

Stereo- Atlas of Ostracod Shells 9 (12) 63-68 (1982)
595.337.14 (119.9) (261.26 : 161.009.58) : 551.351

Propontocypris trigonella (1 of 6)

ON PROPONTOCYPRIS TRIGONELLA (SARS)

by John Athersuch and John E. Whittaker
(B.P. Research Centre, Sunbury and British Museum (Natural History), London )

Genus Propontocypris Sylvester-Bradley, 1947
Type species: (designated by Sylvester-Bradley, 1947) Pontocypris trigonella Sars, 1866

1866

1912

1923

1947

1969

1969

1969

Pontocypris gen. nov. G. O. Sars (pars), Forh.VidenskSelsk.Krist., 1865, 13.

Pontocypris Sars; G. W. Muller, Tierreich, 31, 109.

Pontocypris Sars; G. O. Sars, An Account of the Crustacea of Norway , vol. 9, Ostracoda, Bergen Museum, parts 3, 4, 47.
Propontocypris gen. nov. P. C. Sylvester-Bradley, Ann. Mag. nat.Hist., ser. 11. 13, 193.

Propontocypris (Propontocypris) Sylvester-Bradley; R. F. Maddocks , Smithson. Contrib. Zook, 7, 11.

Propontocypris ( Ekpontocypris ) subgen. nov. R. F. Maddocks, ibid., 27.

Propontocypris (Schedopontocypris) subgen. nov. R. F. Maddocks, ibid., 37.

Diagnosis: Carapace trigonal, longest ventrally, moderately compressed; RV larger than LV; marginal flanges
weak or absent; marginal serrations absent. Greatest height and inflation at, or slightly in front of
midpoint. Adductor muscle-scars form cluster of five arranged in three near-horizontal rows or in a

Explanation of Plate 9, 64

Fig. 1 .d'LV, ext. lat. ( 1982.25, 620 /am long); fig. 2,9 LV, ext. lat. ( 1982.26, 650 /rm long); fig. 3,9 car., ext. dors. (1982.27, 650/xm long).
Scale A (200 /um; x 89), figs 1-3.

I i i
.414

--rt-t

- -t-T — I
--+ + H

44

-y-n-

Stereo-Atlas of Ostracod Shells 9, 65

Propontocypris trigonella (3 of 6)

Diagnosis (contd.)/ rosette. First antenna 8-jointed; male legs symmetrical or nearly so. Terminal pectinate seta of third
leg exceeded in length by one other seta. Subterminally, furca bears two stout setae of medium
length flanked by two unequal, slender setae. Testes positioned posteriorly, curving forward
ventrally. S-shaped ovaries positioned posteriorly. Large eyes present.

Remarks:

Maddocks (1969) recognised three subgenera of Propontocypris based mainly on carapace curvature
and details of the muscle-scar patterns. The appendages of all 3 subgenera would appear to be
very similar, except for the fine structure of the genetalia.

In our opinion the criteria used to separate these taxa are rather subjective and we prefer
not to subdivide the genus.

Propontocypris may be distinguished from Pontocypris both on carapace and appendage
characters. Pontocypris has five muscle-scars arranged in three horizontal rows, two in each of the
lower rows, and one centrally above; the greatest inflation and height coincide well in front of the
midpoint and the carapace is strongly elongated posteriorly; the dorsal margin is angular and the
posteroventral margin of the RV is serrate. Maddocks (1969) records a number of striking
differences between the soft parts of these two genera. Pontocypris is blind, has a 7-jointed first
antenna and differs from Propontocypris in the detailed anatomy of the legs, the furcal setae and
the genetalia.

The diagnosis of the appendages of Propontocypris is based on our own observations together
with those recorded by Sars, 1923 and Maddocks, 1969.

Explanation of Plate 9, 66

Fig. 1 . c(RV, ext. lat. ( 1982.28, 630 yam long); fig. 2, 9 RV, ext. lat. ( 1982.29, 650 /xm); fig. 3, 9 LV, int. lat. ( 1982.30, 650 yum long).
Scale A (200/am; x 89), figs. 1-3.

H-l

l-t-l-

4L.
I I I

|T

i I

I I
I I

Lii

I I
I I
I I

I I

— f-T

i ! Stereo-Atlas of Ostracod Shells 9, 64

i i
i i

-1-J-

-i-r

Propontocypris trigonella (2 of 6)

t-r

■h-M-

-l_ J_l_-

Stereo-Atlas of Ostracod Shells 9, 66

__n_

-44-

I

Propontocypris trigonella (4 of 6)

i -r

i i
i i

r!'

4.L
I I

_ L-LX.

1-i-

l~1-t-

I--M-

Stereo- Atlas of Ostracod Shells 9, 67

Propontocypris trigonella (5 of 6)

Propontocypris trigonella (Sars, 1866)

1866 Potocypris trigonella sp. nov. (sic) G. O. Sars, Forh.VidenskSelsk.Krist., 1865, 16 (lapsus calami).

1868 Pontocypris trigonella Sars; G. S. Brady, Trans, Linn. Soc.Lond., 26, 387, pi. 25, figs. 31-34; pi. 38, fig. 3.

1923 Pontocypris trigonella Sars; G. O. Sars, An Account of the Crustacea of Norway , vol. 9, Ostracoda, Bergen Museum, 48. pi. 20.

1947 Propontocypris trigonella (Sars); P. C. Sylvester-Bradley, Ann. Mag. nat. Hist., ser. 11, 13, 193 (new combination).

1957 Pontocypris trigonella Sars; A. P. C. de Vos , Arch.Zool. exp. gen., 95, 10, pi. 4, figs. la-i.

Type specimens: Several dried and unrecognisable specimens remain in the Sars Collection, Zoological Museum,
Oslo, Norway, no. F 1532. However, Sars’ (1923) illustrations and description adequately define
this species.

Type locality: Risor, Norway, approx, lat. 58° 44' N, 09° 15' E. Recent.

Diagnosis: Carapace of medium size (c. 0.60-0.70 mm), dimorphic. Greatest height slightly anterior to mid-
point. with broadly rounded dorsal margin; more tapered posteriorly; rounded at both ends. In
dorsal view, carapace moderately inflated, widest in front of midpoint, rounded posteriorly,
acutely tapering anteriorly. Slight marginal flange anteriorly, and to a lesser extent, posteriorly,
in both valves. Male carapace proportionately more elongate than female. Furcae and male
copulatory appendages distinctive.

Figured specimens: Brit. Mus. (Nat. Hist.) nos. 1982.25 (cfLV: PI. 9, 64, fig. 1), 1982.26 (?LV: PI. 9, 64, fig. 2), 1982.27
( 9 car.: PI. 9, 64, fig. 3), 1982.28 (rfRV: PI. 9, 66, fig. 1), 1982.29 (? RV: PI. 9, 66, fig. 2), 1982.30
(9 LV: PI. 9, 66, fig. 3), 1982.31 ( cTappendages: Text-fig. 1). 1982.25, 26, 28-30, ex Norman
Collection no. 1911.11.8.M3019 from “oyster ooze” at Stranraer, SW Scotland (lat. 54° 55' N,
long. 05° 00' W); 1982.27 ex Scott Collection from Loch Fyne, W Scotland, collected 1896;
lat. 56° 00' N, long. 05° 25' W. 1982.31, ex Norman Collection, no. 1900.3.6.150, from Firth of
Clyde, W Scotland, collected July 1885.

+--M-

Stereo- Atlas of Ostracod Shells 9, 68 Propontocypris trigonella (6 of 6)

Remarks: For a comparison with Propontocypris pirifera (G. W. Muller) with which P. trigonella has frequently
been confused, see Athersuch & Whittaker, (Stereo-Atlas of Ostracod Shells, 9, 69-76, 1982).

No males could be found from our British collections which were well enough preserved to
allow dissection of the male copulatory appendages, although one specimen still had a furca intact
(Text-fig. 1). Text-fig. 4 is therefore redrawn after Sars (1923, pi. 20) for comparison of the copulatory
appendages with those of P. pirifera (see Athersuch & Whittaker, op.cit., text-figs. 2, 4, 6); a further
drawing, again after Sars (1923), is finally appended (Text-fig. 2) in order to compare the furca of
the Norwegian specimen with that from Britain.

Distribution: A NW European species; records from Scandinavian and British coasts can be confirmed; Medi-
terranean records need to be checked. Found in small numbers on a variety of substrates in sub-littoral,
marine environments.

Text-figs. 1-4, Appendages of P. trigonella, M. Fig. 1, Furca (1982.31). Firth of Clyde, Scotland; figs. 2-4, Furca, Zenker's organ and
copulatory appendage, Norway (after Sars, 1923, pi. 20). Scale lOOyum; fig. 1. Figs. 2-4, no scale.

t -r

-4-U

I I I

Stereo-Atlas of Ostracod Shells 9 (13) 69-76 (1982)

595.337.14 (119.9) (261.26 + 261.27 + 261.28+ 262 : 161.014.40) : 551.313.1 + 551.351

Propontocypris pirifera (1 of 8)

ON PROPONTOCYPRIS PIRIFERA (G. W. MULLER)

by John Athersuch and John E. Whittaker
(B.P. Research Centre, Sunbury and British Museum < Natural History), London)

Propontocypris pirifera (G. W. Muller, 1894)

1889

1894

1957

1969

1969

1976

Pontocypris trigonella Sars; G. S. Brady & A. M. Norman (pars), Scient. Trans. R.Dubl.Soc., 4, 109. pi. 22, figs. 1 8-25; pi. 23,
fig. 6 (non P. trigonella Sars, 1866).

Pontocypris pirifera sp. nov. G. W . Muller, Fauna Flora GoIf.Neapel, 21, 247, pi. 10, figs. 1-3, 1 8-20. 22-24; pi. 38. fig. 52.
Pontocypris pirifera Muller; A. P. C. de Vos . Arch. Zool. exp. gen., 95, 12, pi. 5, figs. la-1.

Propontocypris pirifera (Muller); 1. Yassini, Bull. Inst. Geol.Bassin Aquitaine. 7, 29. pi. 15.

Propontocypris ( Ekpontocypris ) pirifera (Muller); R. F. Maddocks, Smithson. Contrib. Zool., 7, 27 (no description).
Propontocypris pirifera (Muller); G. Bonaduce, G. Ciampo & M. Masoli .Pubbl.Staz. zool. Napoli, 40, pi. 9, figs. 5, 6 (no description).

Type specimens: Several specimens remain in the Muller Collection at the Zoologischen Institut, Greifswald and at
the Institut fur Spezielle Zoologie und Zoologisches Museum der Humboldt Universitat zu Berlin,
E Germany.

Type locality: Bay of Naples, W Italy, approx, lat. 40° 50' N, long. 14° 15' E. Recent.

Explanation of Plate 9, 70

Fig. 1, d car., ext. rt. lat. ( 1982.1 1, 970/u.m long); fig. 2, ? car., ext. rt. lat. (1982.12, 830 /rm long); fig. 3, 9 car., ext. It. lat. (1982.13,
790yu,nr long).

Scale A (250/un; x 65), figs. 1-3.

-r-i

-t--t

JiJ

+ +--4

Stereo- Atlas of Ostracod Shells 9, 7 1

Propontocypris pirifera (3 of 8)

Diagnosis: Carapace large, strongly dimorphic. Greatest height at midpoint; marked by distinct cardinal angle,
particularly in RV. Anterior margin broadly rounded, posterior slightly more tapered. In dorsal
view, carapace moderately and evenly inflated, widest at midpoint, rounded at both ends. Marginal
rim anteriorly in RV. Males more trigonal than females. Furcae and male copulatory appendages
distinctive.

Figured specimens: Brit. Mus. (Nat. Hist.) nos. 1982. 1 1 (d'car.: PI. 9, 70, fig. 1 ; PI. 9, 74, fig. 3), 1982.12 (9car.: PI. 9, 70,
fig. 2), 1982.13 (9car.; PI. 9, 70, fig. 3), 1982. 14 (d car.: PI. 9, 72, fig. 1), 1982.15 (9car.: PI. 9, 72,
fig. 2), 1982.16 (d'LV: PI. 9, 72, fig. 3), 1982. 17 (A-l car.: PI. 9, 74, fig. 1), 1982.18 (9car.: PI. 9, 74,
fig. 2), 1982.19 (cfRV and appendages: PI. 9, 76, fig. 1), 1982.20 (9 RV: PI. 9, 76, fig. 2), 1982.21
(cfRV: PI. 9, 76, fig. 3), 1982.22 (c/appendages: Text-figs. 1, 2), 1982.23 (cf appendages: Text-figs.
3, 4), 1982.24 (d appendages: Text-figs. 5, 6).

1982.11-14, 17-19, 21, 22, were collected alive by J. E. Whittaker from various stations in
East Fleet, Dorset, S England (lat. 50° 36' N, long. 02° 28' W), between August 1968 and August
1969, on Zostera, green-algae and Laminaria holdfasts; salinities varied between 31 and 35°/oo,
and water temperature 7 to 21°C, depth 0.5-3m. 1982.16, 20, 23, are from Cyprus and were
collected alive by J. Athersuch, in November 1973. 1982.16, 23, are from S of Cape Greco (lat.
33° 55' N, long. 34° 10' E), water temperature 21.5°C, depth 8m on algae; 1982.20 is from
Famagusta Bay (lat. 35° 07' N, long. 33° 56' E), water temperature 22°C, depth 15 m, in fine sand.
Salinity in both localities c. 39°/oo. 1982.15, 24 (ex slide no. 1972.3.2.2) collected by K. G. McKenzie
from Lago del Fusaro, near Naples, W Italy; approx, lat. 41° 40' N, long. 14° 04' E.

Explanation of Plate 9, 72

Fig. 1, d'car., ext. It. lat. (1982.14, 940 /urn long); fig. 2, d'car., ext. It. lat. (1982.15, 860^01 long); fig. 3, d'car., ext. It. lat. (1982.16,
830/u.m long).

Scale A (250/u.m; x 65), figs. 1-3.

•rr

■TP-

.XL.
I I

.L_Ll-

•t— t-T'

Stereo-Atlas of Ostracod Shells 9, 70

Propontocypris pirifera (2 of 8)

i-r

t— t~t~

LL.

+--M-

Stereo- Atlas of Ostracod Shells 9, 72

-44-

Propontocypris pirifera (4 of 8)

t-r

i i
i i

T-t-t

Stereo- Atlas of Ostracod Shells 9, 73

Propontocypris pirifera (5 of 8)

Remarks: P. pirifera differs in size and shape from P. trigonella (Sars) with which it has often been confused.

P. trigonella is smaller and more elongate, particularly posteriorly, than P. pirifera. Furthermore,
the greatest height and inflation is further forward in P. trigonella. The furcae and copulatory
appendages of each species are also very distinctive (cf. Text-figs. 1-6, herein and Stereo-Atlas of
Ostracod Shells , 9, 63-68, text-figs. 1, 2, 4). Both species are sexually dimorphic but in P. pirifera
this is very pronounced; the male is the larger of the two sexes in P. pirifera , but the smaller dimorph
in P. trigonella. The male carapaces from the population in The Fleet are considerably larger (c.

O. 95mm) than those from the Mediterranean (c. 0.85mm), whilst their copulatory appendages
are virtually identical in size. The pear-shaped sperm heads (after which the species is named) are
well seen in PI. 9, 76, fig. 1.

Distribution: Associated with algae and sea-grass in brackish and marine littoral and shallow sublittoral environ-
ments. Some records from Scotland, S England, the Atlantic coast of France and the Mediterranean,
as far E as Cyprus, can be confirmed. Many specimens from British coasts, examined by the authors
in the Brit. Mus. (Nat. Hist.) and Hancock Museum, Newcastle-upon-Tyne, purporting to be

P. trigonella, are referable to P. pirifera. Our investigations show that P. pirifera has a wider
geographical distribution outside the Mediterranean than previously recognised.

Explanation of Plate 9, 74

Fig. 1, A-l car., ext. It. lat. ( 1982.17, 750 /im long); fig. 2, 9 car., ext. dors. (1982.18, 850yum long); fig. 3. d'car., ext. rt. lat. showing
normal pores and setae (1982.11).

Scale A (250^tm; x 65), figs. 1, 2; scale B (10/xm; x 1,650), fig. 3.

tt

-rr

-t-t-i

-t-r-i

U

jjj-

-t--+ —

Stereo-Atlas of Ostracod Shells 9, 75

Propontocypris pirifera (8 of 8)

Text-figs. 1-6. Appendages of P. pirifera, dd. Figs. 1, 2, Furca and copulatory appendage (1982.22), The Fleet, S England; figs. 3, 4,
ditto (1982.23), Cyprus; figs. 5, 6, ditto (1982.24), Naples. Scale 100 pm: x 200. After drawings by D. J. Horne.

Explanation of Plate 9, 76

Fig. 1, d, detail of sperm (1982.19); fig. 2, 9 RV, int. lat. (1982.20, 710pm long); fig. 3, c/RV, int. lat., muse. sc. (1982.21).
Scale A (25/u.m; x 520), fig. 1; scale B (250/u.m; x 65), fig. 2; scale C (50/xm; x 330). fig. 3.

■ t r ■
-tt

-i -i

I L l_ -

i-r

i i i

i i i

. J L.L-

i i i
i i i
i i i
i i i
' 1 T “
I I I
I I I
I I I
I I I
I I
I I
I I

Stereo- Atlas of Ostracod Shells 9, 74

Propontocypris pirifera u>

LJ_

-+~f-

Stereo- Atlas of Ostracod Shells 9, 76

Propontocypris pirifera (8 of 8)

Stereo-Atlas of Ostracod Shells 9(14) 77-84 (1982)

595.337.14 (1 19.9) (261.26 : 162.005.045- 161.020.57) : 551.351

Bonnyannella roberisoni (1 of 8)

-Tl"T

ON BONNYANNELLA ROBERTSON I (BRADY)

by John Athersuch

(B.P. Research Centre, Sunbury-on-Thames, England)

Derivation of name:
Diagnosis:

Genus BONNYANNELLA gen nov.

Type species: Cythere robertsoni Brady, 1868

After my wife. Dr Anne Bonny, in appreciation of her encouragement of my ostracod studies.

Genus of the Loxoconchidae with small (400-500/u.m long) reticulate carapace; elongate and
quadrate in lateral view; in dorsal view evenly inflated, broadly rounded anteriorly, obtusely tapered
posteriorly. Large smooth eye spots. Conjunctive, rimmed normal pores. Hinge gongylodont,
median element smooth, posterior tooth of right valve curved around socket. Adductor muscle scars
decrease in size from top to bottom; upper three scars elongate, lower scar rounded. Frontal scar
U-shaped. Fulcral notch present. Antennula and antenna stout with long strong setae; antennula
six-jointed. Respiratory plate of maxillula bears a single reflexed seta. Male copulatory appendage
subovate with an inconspicuous rounded terminal lappet; ductus ejaculatorius conspicuous and
simply coiled.

Explanation of Plate 9, 78

Fig. 1, ? LV, ext. lat. (lectotype, Hancock Museum specimen A, 480 /urn long); fig. 2, 9 car., ext. rt. lat. ( 1982.1, 460 /cm long); fig. 3,
dCV, ext. lat. (1982.2, 450/u.m long).

Scale A (lOOyum; x 127), figs. 1-3.

-rr

--tt -i-
--t-i-H-
I
I
I

-J4J.

Stereo-Atlas of Ostracod Shells 9, 79

Bonnyannella robertsoni (3 of 8)

Remarks: Bonnyannella differs from Loxoconcha in having antennulae with six, instead of five, podomeres and
in possessing a fulcral notch and a smooth median hinge element. It may be distinguished from
Cytheromorpha by its smaller size, by weaker sexual dimorphism, by less pronounced subdivision of
the posterior tooth/socket and by a less prominent fulcral notch. The two genera differ in the general
form of the copulatory appendages and, as far as I am able to determine, Cytheromorpha lacks an
aberrant reflexed seta on the respiratory plate of the maxillula. The type species of Bonnyannella,
B. robertsoni, closely resembles Sagmatocy there, typified by S. napoliana (Puri) in the general
structure of the hinge, except that in Bonnyannella the terminal elements are less curved and are not
as distinctly lobed as in Sagmatocy there. However, the two genera differ in a number of other
important respects. Unlike Sagmatocy there, Bonnyannella does not have sub-parallel dorsal and
ventral margins, a conspicuous marginal flange or pore conuli. In addition, the reticulum of
Bonnyannella is far less pronounced with a tendency towards celation of the fossae. In contrast to
Sagmatocy there, neither sex is highly inflated and the male does not have a post-ocular depression.
The four distal podomeres of the antennulae in Bonnyannella are noticeably stouter and bear
relatively much longer and more conspicuous setae than those of Sagmatocythere. Furthermore, the
distal seta of the second podomere of the third leg is longer than the third podomere, whereas in
Sagmatocythere it is shorter. The length of this seta has been used in the diagnosis of another
loxoconchid genus , Lindisfarnia (Horne & Kilenyi, Stereo-Atlas of Ostracod Shells, 8, 107, 1981),
and may prove to be of some taxonomic significance. The other appendages of Bonnyannella and
Sagmatocythere are similar.

Explanation of Plate 9, 80

Fig. 1, dear., ext. dors. (Hancock Museum specimen B, 450|U,m long); fig. 2, 9 car., ext. dors. (Hancock Museum specimen C, 470/am
long); fig. 3, d RV, int. lat. ( 1982.3, 450/u.m long); figs. 4, 5, c/RV, int. lat. terminal hinge elements (1982.4, 440 /am long); figs. 6, 7,
c/LV, int. lat., terminal hinge elements (1982.4).

Scale A (100/u.m; x 127), figs. 1-3; scale B (25/um; x 470), figs. 4-7.

-_LL.

I

iffli ■

I I
I I
I I
I I

■ t-+T-
I I
I
I

Stereo- Atlas of Ostracod Shells 9, 78

Bonny annella robertsoni (2 of 8)

i-r

h-M-'

-U.L.

-h-M--
I

Stereo-Atlas of Ostracod Shells 9, 80

-n-

Bonnyannella robertsoni (4 of 8)

t-r

±_1.

Stereo- Atlas of Ostracod Shells 9, 81

Bonny annella robertsoni (5 of 8)

Bonnyannella robertsoni (Brady, 1868)

1868

1969

Cythere Robertsoni sp. nov. G. S. Brady , Ann.Mag.nat. Hist., ser. 4, 2, 33, pi. 4, figs. 5, 8-10.

Cytheromorpha robertsoni (Brady); I. Yassini, Bull. Inst. Geol.Bassin Aquitaine. 7, 111, pi. 18, fig. 3; pi. 19, fig. 20.

Type locality:
Diagnosis:

Lectotype: In the Brady Collection, Hancock Museum, Newcastle-upon-Tyne, 9 LV. No registration number,
but housed in a separate, labelled slide.

Drobak, Oslofjord. S Norway (lat. 59° 40' N, long. 10° 40' E); from 30- 35 fathoms (55-65 m); Recent.

Carapace with numerous small, sub-rounded, deep fossae; evenly inflated with slight posterior
protuberances dorsally and ventrally; parallel-sided in dorsal view.

Figured specimens: Hancock Museum specimens (no catalogue numbers, but placed in separate labelled slides) A
(lectotype, 9 LV: PI. 9, 78, fig. I), B (d'car.: PI. 9, 80, fig. 1), C (9 car.: PI. 9, 80, fig. 2).

Brit. Mus. (Nat. Hist.) specimen nos. 1982.1 ( 9car.: PI. 9, 78, fig. 2), 1982.2 (d'car.: PI. 9, 78,
fig. 3), 1982.3 (cfRV: PI. 9, 80, fig. 3; Text-fig. 2). 1982.4 (dTV & RV: PI. 9, 80, figs. 4-7), 1982.5

(9car. and appendages: Text-figs. 3a-d, 4b-d), 1982.6 (d'car. and appendages: Text-fig. 4a), 1982.127
(9 RV: Text-fig. 1).

Hancock Museum specimens A, from Drobak, S Norway, B and C from 4 miles off Hawthorn,
Yorkshire, NE England (ex Brady slide N3), depth 20 fathoms (36m).

Brit. Mus. (Nat. Hist.) specimens 1982.1 and 1982.2 were collected by T. Scott from the Firth of
Clyde, SW Scotland. 1982.3, from beach sand at Cemaes Bay, Anglesey, N Wales (lat. 53° 25' N,
long. 04° 30' W), was collected by C. P. Palmer. 1982.4, 6 and 127 were found by the author in beach
sand: 1982.4 and 127 at Robin Hood’s Bay, Yorkshire, NE England (lat. 54° 25' N, long. 00° 35' W)
and 1982.6 from Colwyn Bay, N Wales (lat. 53° 17' N, long. 03° 44' W). 1982.5, collected by
D. J. Horne, was living amongst Laminaria holdfasts at low-water mark. Gore Point, Porlock,
SW England (lat. 51° 14' N, long. 03° 37' W).

TT

■rr

--t-r

i — i —

i • i“

Stereo-Atlas of Ostracod Shells 9, 83

Bonnyannella robertsoni (7 of 8)

Remarks: Cytheromorpha exigua Wouters, 1978 (Een Systematische, Biostratigrafische en Paleobiologische
studie van de Ostracoda uit Miocene afzettingen in Noord-Belgie, Leuven, 2, 25, pi. 4, figs, la, lb;
pi. 34, figs. 7a, 7b). from the Egedem and Antwerp Sands (Miocene) of Belgium may be referred
to Bonnyannella. It differs from B. robertsoni in dorsal view in being more inflated and rounded
posteriorly. It also lacks any posterodorsal protuberance. I am unaware of any other species that
may be assigned to Bonnyannella.

Distribution: Recorded from the lower littoral and sublittoral zones of the coasts of Britain, mainly in the south,
to a depth of about 70 m. Known also from the Atlantic coasts of France, Germany, S Norway
and the Baltic.

v Ik

b

Text-fig. 1

50/i.m
Text-fig. 2

Text-fig. 1, 6 RV int. seen in transmitted light (1982.127).
Text-fig. 2, cfRY int., muscle-scars (1982.3).

— I — L L_ .

. L-L

t--r

Stereo-Atlas of Ostracod Shells 9, 82

Bonny annella robertsoni (6 of 8)

-i-ri—

-h+t--

. J-L.

Text-fig. 3, 9 appendages: a, antennula; b, antenna; c, maxillula; d, mandible (1982.5). (Kindly drawn by D. J. Horne.)

4- + -

~n~

Stereo-Atlas of Ostracod Shells 9, 84

Bonnyannella robertsoni (8 of 8)

100/xm

Text-fig. 4, a, d copulatory appendage (1982.6); b-d, 9 legs (1982.5). (Kindly drawn by D. J. Horne.)

-t-r-

- — i -i — i —
1 1 i

t — * — — — — >

Palaeontological

microslides

Cardboard slides in aluminium holders

Glass or thick clear acetate coverslides. Single, 4-celled orfaunal (32 or64 cell divisions)

Plastic slides

Single, double, 3 or4-celled orfaunal (32or64celldivisions)

All slides sold completely assembled
For air- freight, sold without glass coverslides if required

Cotton and plastic sample collecting bags and paper storage packets

EKHull Microslide Co

24, Lynmouth Gardens, Perivale, Middlesex UB6 7HR, England
Telephone 01-935 3777

V,

Stereo-Atlas of Ostracod Shells: Vol. 9, Part 1

CONTENTS

9 (1) 1- 8 On Braderupia asymmetrica (Neckaja); by R. E. L. Schallreuter (£2.80)

9 (2) 9-16 On Bilobatia serralobata Schallreuter; by R. E. L. Schallreuter (£2.80)

9 (3) 17-20 On Byrsolopsina manca Schallreuter sp. nov.; by R. E. L. Schallreuter (£1.40)

9 (4) 21-26 On Loxoconcha cuneiformis Malcolmson; by D. J. Horne & E. Robinson (£2.10)

9 (5) 27-32 On Loxoconcha dimorpha Hartmann; by D. J. Horne (£2.10)

9 (6) 33-40 On Loxoconcha linleyi Horne sp. nov.; by D. J. Horne (£2.80)

9 (7) 41-46 On Waiparacythereis joanae Swanson; by K. Swanson (£2.10)

9 (8) 47-50 On Acanthocythereis dohukensis Khalaf sp. nov.; by S. K. Khalaf (£1.40)

9 (9) 51-54 On Actinocythereis iraqensis Khalaf sp. nov.; by S. K. Khalaf (£1.40)

9 (10) 55-58 On Chrysocythere naqibi Khalaf sp. nov.; by S. K. Khalaf (£1.40)

9 (11) 59-62 On Hermanites transversicostata Khalaf sp. nov. (£1.40)

9 (12) 63-68 On Propontocypris trigonella (Sars); by J. Athersuch & J. E. Whittaker (£2.10)

9 (13) 69-76 On Propontocypris pirifera (G. W. Muller); by J. Athersuch & J. E. Whittaker (£2.10)

9 (14) 77-84 On Bonnyannella robertsoni (Brady); by J. Athersuch (£2.80)

Prepaid annual subscription (valid for Volume 8, 1981)

Individual subscription £20.00 or US $ 50.00 for 2 parts (post free)

Price per Part: £20.00 or US $ 50.00
Institutional subscription £30.00 or US $ 75.00 for 2 parts (post free)

Price per Part: £30.00 or US $ 75.00

Back volumes: Vol. 1 (4 Parts): £20.00; price per Part: £5.00
Vol. 2 (4 Parts): £28.00; price per Part: £7.00
Vol. 3 (2 Parts): £24.00; price per Part: £12.00
Vol. 4 (2 Parts): £30.00; price per Part: £15.00
Vol. 5 (2 Parts): £32.00; price per Part: £16.00
Vol. 6 (2 Parts): £40.00; price per Part: £20.00
Vol. 7 (2 Parts): £40.00; price per Part: £20.00
Vol. 8 (2 Parts): £60.00; price per Part: £30.00

Prices of Numbers as shown in Lists of Contents for each Part
Postage extra in sales of all back Numbers and back Parts
No trade discount is allowed on the subscription rate

Orders should be addressed to: Dr. C. W. Haskins,

Robertson Research International Limited,

‘Ty’n-y-Coed’, Llanrhos, Llandudno, Gwynedd LL30 ISA.

Cheques should be made payable to Robertson Research Co. (Stereo-Atlas Account)

SPECIAL OFFER

50% off all back part prices if
you become a subscriber to the Atlas