Washington Park
Arboretum Bulletin
Published by the Arboretum Foundation
for the University of Washington
Vol. 49, No. 4, Winter 1986
$2.50
STAFF
Harold B. Tukey, Jr.
Director of Arboreta
Brian O. Mulligan
Director Emeritus
Timothy Hohn
Curator, Plant Collections
J. A. Wott
Professor, Continuing
Education
OFFICERS OF THE
ARBORETUM FOUNDATION
Mrs. C. Edwards Simons, Jr.
President
Mr. Richard Doss
1st Vice-President
Mrs. David Taft
2nd Vice-President
Mr. Paul Thienes
3rd Vice-President
Dr. Alan Adams
4th Vice-President
Mrs. J. Newton Morris
Secretary
Mrs. Frank Thome
Treasurer
Col. Leroy P. Collins
Immediate Past-President
BULLETIN EDITORIAL
BOARD
Nancy Pascoe
Editor
Nancy Ballard
James Clark
Col. Leroy P. Collins
Rosamund P. Engle
S.P. Gessel
Timothy Hohn
Tina Kuhnle
B.J.D. Meeuse
Brian O. Mulligan
Jan Pirzio-Biroli
Ruth E. Vorobik
Concerning This Issue...
This issue of the Bulletin contains many
articles of international interest — starting with Ward
Horn's report about the University's involvement in
the international seed exchange. In addition we have
two fun articles about British gardens; one by Dennis
Thompson, plant expert, and the other by Sue
Buckles, who has a particular interest in those gardens
around the Welsh border. A little closer to home is an
article about the University of British Columbia's
plant introduction program by Bruce Mcdonald. So as
not to give short shrift to the gardens of the United
States, Virginia Morrel has written about Wave Hill
in the Bronx. Although technically not international,
the University of Washington's Herbarium draws
together the plant knowledge from countries all over
the world. Melinda Denton, Curator of the Herbarium
has written an interesting article about the role of
herbaria in general and the University's Herbarium in
specific.
This is a good time to sit by the fire and do
some armchair garden touring!
Nancy Pascoe
Editor
The ARBORETUM BULLETIN is published quarterly, as a bonus of membership, by the Arboretum
Foundation, a non-profit organization to further the development of the Washington Park Arboretum. Information
regarding membership in the Foundation may be obtained by writing to the Arboretum Foundation, University of
Washington XD-10, Seattle, WA 98195 or by calling (206) 325-4510. Articles on botany and horticulturally-related
subjects written by professional and amateur botanists, horticulturists, educators and gardeners are welcome. No part of
the BULLETIN may be reprinted without the authority of the Arboretum Foundation. Typesetting and design by Nancy
Pascoe, lithography by United Graphics Printers.
©copyright 1986, Arboretum Foundation
UW Arboretum Bulletin
The Washington Park
Arboretum Bulletin
VOLUME 49, NUMBER 4, WINTER 1986
TABLE OF CONTENTS
The International Seed Exchange, by Ward Horn 2
The University of British Columbia Botanical Garden's
Plant Introduction Scheme — An Innovative Program for
the Introduction of New and Recommended Plants,
by Bruce Macdonald 5
In The Arboretum, by Barbara Engler 9
Small British Gardens — Some Personal Favorties, by
Dennis Thompson 10
Gardens of the Welsh Border Country, by Sue Buckles...
Wave Hill, by Virginia Morrel 20
A Collection of Botanical Treasures — The University
of Washington Herbarium, by Melinda F. Denton 22
Arboretum Foundation Book Fair 25
The Donald G. Graham Visitors Center 26
The Arboretum Foundation Spring Garden Tour 27
A Letter to the Editor 27
COVER
Idesia polycarpa, a well-known
feasture of the gardens at Wave Hill.
From the book leones Plantarum
Sinicarum, edited by Hsen-Hsu Hu &
Woon- Young Chun, published by the
Commercial Press, Shanghai, China.
The International Seed Exchange
WARD HORN
We call it the "International Seed
Exchange", which is an imposing name for
something with no charter or by-laws, no
offices or committees, no dues and, perhaps
best of all, no meetings. Nevertheless, it
exists on a world-wide basis and is a major
source of plant material for botanical gardens
and arboreta. It's operations are simple; partic-
ipating institutions collect seeds and offer them
to each other on a no-charge basis. Our own
Arboretum sends its "Index Seminum" (another
imposing name, which freely translated, means
"seed list") to about 550 correspondents in 50
countries. They, in turn, send their Index Sem-
ina to us. This article will be limited to a dis-
cussion of our distribution of seeds to other
institutions.
Our Index Seminum of seed collected
in 1985 was sent out in January, 1986 and list-
ed 203 items. Fifty-one of these were collected
from the wild in California, Idaho, Oregon and
Washington.1 The other 153 items were from
the Arboretum or, in a few instances, came
from private gardens. All but three of the
items were woody plants.
Items listed in the Index Seminum
change from year to year depending on seed
availability and on the demand or lack of de-
mand in previous years. The list is limited to
Collectors were Director-Emeritus Brian Mulligan
and his wife Margaret, Arboretum naturalist JanPirzio-
Biroli and her husband Giacomo, graduate student
Clayton Antieau, A.L. Jacobson, E. and M. Mason
and Ward Horn.
about 200 items. We usually collect more than
that number and thus difficult decisions must
be made as to what to exclude. This will not
be a problem for the 1987 list; many plants
did not set seed because of unfavorable weather
in 1986 at flowering time.
With few exceptions we offer seeds of
species only, and exclude cultivars and hybrids,
with the exception of an occasional item which
may be useful for breeding purposes and is not
widely available. This means that many of the
nearly 5,000 different kinds of woody plants in
the Arboretum will not be included on our list.
In addition, some species may be excluded be-
cause of the possibility of cross-pollination.
For example, we usually do not list oaks, pines
and some other wind-pollinated genera. How-
ever, in 1986 we did send out acorns from the
Quercus vacciniifolia which grows in a location
remote from other oaks — it was apparently
remote from the squirrels as well! We look for
seed plants which are well separated from close-
ly related species or which bloom at different
times.
Through September 30, 1986 we had
received requests ("desiderata" is the technical
term) for seeds from 284 institutions in 41
countries. We had sent out 4,483 packets of
seeds and, had we not run out of seeds, could
have sent 898 more. The following table
shows the number of correspondent institutions
in several countries and the number of such
institutions which requested seeds.
2
UW Arboretum Bulletin
Pulpy seeds are soaked in water in plastic bags for a week or two, then photo: Joy Spun-
put in a blender for a second to break down the pulp. Photo is of the author.
Request Table
The first figure represents the number of
Arboretum Index Semina sent out and the second
figure the number of requests received from each
country.
Argentina
2
1
Australia
5
3
Belgium
13
11
Great Britain
47
32
France
25
21
East Germany
9
8
West Germany
26
19
Iceland
3
2
Hungary
12
7
Japan
10
5
Monaco
1
1
Norway
4
3
Poland
9
7
Russia
51
30
The largest request we received was for 133
items and came from Hungary. Forty-seven
requests were for more than 50 items. Eight
requests were for one item only, and seven
asked for two items. We sent ten items, all
wild-collected, to the Royal Botanical Gardens
at Kew and received a "thank-you" note from
Queen Elizabeth.
We are often asked which items are the
most popular. There is always a healthy
demand for things collected from the wild. For
example, our 1986 list included the Oregon
white oak, Quercus garryana from two stations
in Washington and one in Oregon; many
correspondents wanted seed of all three. As
another example, we had 54 requests for devil's
club, Oplopanax horriduml Unfortunately, or
fortunately according to one's view, we did not
have enough seed to fill all of these requests.
With regard to seeds collected in the Arboretum,
records from the past five years show that
Winter 1986 (49:4)
3
For three years, Ward Horn has been the stalwart leader of volunteers working on the
International Seed Exchange. Although the distribution of seed had long been carried out by
Arboretum Foundation members, those of us who became responsible for collecting and cleaning it
started from scratch after the death of Joe Witt and the unexpected retirement of Pablo Abellera, who
for many years performed these important aspects of the project.
Ward stepped in with his almost daily explorations of the Arboretum in search of plants
that were bearing good seed. In addition, he proved to be very inventive when it came to processing
seed. He purchased a rolling pin and strainers; he scrounged a hair drier (ideal for blowing away the
chaff) and a homemade food drier for initial dehydration of ripe fruits. His small bonehandled
pocket knife is in frequent use to cut open fruits or seed to determine their viability.
He is the leader of collecting forays into the Arboretum and has passed on to a host of
fellow workers his knowledge of the correct methods for handling the varied array of seeds that we
work with. Ward and his cohorts deserve the highest praise for their dedication to a project that
keeps us in touch with similiar institutions around the world. Jan Pirzio-Biroli
species of Acer, Alnus, Magnolia, Sorbus and
Stewartia are much in demand. However not
all species of a genus are equally popular, for
example we had 32 requests for Mahonia ner-
vosa, but only 6 for M. aquifolium , our two
native Oregon grapes. There was a minimum
demand for some other items, as for example 3
for Koelreuteria paniculata and 4 for Photinia
villosa. In 1986 we had heavy requests for the
following:
Dipteronia sinensis
Chamaecyparis thyoides
Viburnum cassinoides
Daphniphyllum macropodum
Sy cop sis sinensis
Corylopsis sinensis
Embothrium coccineum
Podocarpus nivalis
Sciadopitys verticillata
Paulownia lilacina
Until 1984 most of the seeds from the
Arboretum were collected, cleaned and processed
by an employee, Pablo Abellera. Until 1983
the seeds were packaged and distributed by the
Arboretum Foundation units 25 and 66. Since
then this work has been done by other volun-
teers — last year 37 volunteers spent 627 hours
in collecting, cleaning and distributing seeds.
During the 1985 and '86 collecting seasons
volunteers met at 10 a.m. on Monday morn-
ings. We have a saying, "It never rains in the
Arboretum on Monday mornings!", because
this last year good weather has been with us
consistently. Normally collecting begins in
June and extends into early November with
most of the items collected in September and
October. Decisions on what to collect are made
by the Arboretum staff. This has been a
popular program for our volunteers since it
offers a unique way to become better acquainted
with the Arboretum.
Seed cleaning and processing begins
about the first of October and continues until
late in December. The basic process is screen-
ing the fruits to extract the seed. We use a hair
dryer to winnow some seed. Fleshy fruits may
be crushed and allowed to ferment for a few
days in order to break down the pulp, which is
then washed away over screens. We sometimes
use a kitchen blender (its blades covered with
plastic tubing) to break down fleshy fruits.
The cleaned seed is stored in a refrigerator at
about 40e F to preserve viability. Our Index
Seminum is mailed early in January and we
begin to send out seeds about the third week of
February. In most years the desiderata will
have been filled by the end of April, although a
scattering of requests will drift in for several
more months. In each of the past five years
(and probably for many years previously) over
40% of our correspondents have requested seeds.
Some of the Index Semina we receive contain
more than a thousand items. Our own is
limited to about 200 items for reasons noted
»
above and by cost considerations. However,
even though it is a small list we try to make it
a distinguished one which will reflect credit on
the Arboretum, the Center for Urban Horti-
culture and the University of Washington. The
figures sited above indicate that we have
achieved this objective.
4
UW Arboretum Bulletin
The University of British Columbia
Botanical Garden's Plant Introduction Scheme
An Innovative Program for the Introduction
of New and Recommended Plants
BRUCE MACDONALD
Bruce Macdonald is Acting Director of the University of
British Columbia Botanical Garden, Vancouver, B.C.
It was during 1980 that a group of
nurserymen and landscape architects met with
Dr. Roy Taylor, the Garden's past Director
(currently Director of Chicago Botanic Garden),
and his staff to investigate the feasibility of
formulating a program by which some of the
plant collections within our different Garden
components could be selected for commercial
use. As the meetings progressed, we became
increasingly enthused by the long-term benefits
that a plant introduction program of new and
recommended plants could provide for the nur-
sery and landscape industries of British Colum-
bia, and subsequently for other areas within
Canada and the United States. Six years later,
with over a million plants of the first six intro-
ductions being commercially produced by the
participator nurseries, the program has exceeded
our original expectations.
The aim of this article is to outline the
structure of the program, the impact it has
made, and some brief details of the plants cur-
rently publicly released.
The executive committee of the Plant
Introduction Scheme of the University of Brit-
ish Columbia Botanical Garden (PISBG) con-
sists of representatives from the British Colum-
bia Nursery Trades Association (BCNTA), B.C.
Society of Landscape Architects (BCSLA) and
Garden staff. Before recommending procedures
for plant introduction, the committee reviewed
many other plant introduction programs in
different areas of the world and tried to establish
the reasons for their success and failure. The
major mistakes which had been made were poor
communication with the nursery trade, too
many plants introduced at any one time, with
little or no obligation from the nursery and
Winter 1986 (49:4)
5
Mother plants of Anagallis monellii 'Pacific Blue' ready for distribution to the photo: author's
participator nurseries.
landscape industries, too few plants available at
the date of public release, and poor promotion
and publicity. Subsequently, great effort has
been make to rectify these problems, for exam-
ple, a contract is signed between the Garden and
each participator nursery and only one to three
plants will be introduced each year through the
PISBG program.
The initial selection is made by a 35-
member invited evaluation panel which in-
cludes representatives from wholesale and retail
nurseries, landscape contractors, and parks
boards. Some twelve to fifteen plants out of
the Garden's different 16,000 accessions are eval-
uated annually. The final selection is made by
a five-member introduction and release sub-
committee. It is the industry itself which
makes the final choice, not the Botanical
Garden.
The next phase of the program is at
the Botanical Garden nursery where the staff
propagate and grow-on to produce some 500-
1 ,000 one- or two-gallon container plants.
These stock plants are then sold at a premium
price in lots of fifty to the participator
nurseries. The revenue obtained is returned
back into the PISBG program. At this stage, a
contract is signed for subsequent payment of
royalties on the number of cuttings stuck by
the nurseries. The revenue received for the
royalties is also returned to the program.
New plants are registered through the
Canadian Ornamental Plants Foundation
(COPF), while recommended plants are process-
ed through the Garden. COPF, based in Ontar-
io, is an organization which registers new and
improved ornamental plant material in Canada.
A number of nurseries in the United States and
Europe also belong to COPF.
Plants are also sent for testing across
North America. There are currently seven test
sites in Canada, e.g., in Alberta and Ontario,
while there are five in the United States, e.g.,
in Illinois and California. The information
received from the different geographical loca-
tions is then communicated back to the nursery
and landscape industries in British Columbia.
The program was very fortunate to
receive funding from both the Science Council
of British Columbia in Burnaby and the
6
UW Arboretum Bulletin
Wm
Devonian Group of Charitable Foundations in
Calgary. This not only helped to provide staff,
but ensured that our nursery could be further
expanded to handle efficiently the production of
the stock plants.
Publicity and industry involvement
has been a major priority of the PISBG pro-
gram. Colored information sheets are produced,
detailing the culture, nursery propagation and
sales potential of the selected introductions. A
colorful picture tag label has been designed
which must be attached to PISBG plants for
retail sales — this helps the public to identify
the plant with the program. We actively partic-
ipate in nursery trade shows to display and in-
form the industry about the introductions.
Besides radio, television has been a very useful
asset through the popular Canadian Broad-
casting Company (CBC) garden program "West-
ern Gardener", co-hosted by the Garden's educa-
tion coordinator, David Tarrant. The British
Columbia landscape industry has helped to
ensure that the plants have been sited in high
profile areas — for example, Expo 86 and the
recently constructed light rapid transit system
or "Skytrain".
The effectiveness of this promotion
and industry involvement has meant that sales
have been good, with plants being exported to
six different countries. A respected economist
hired by the Science Council of British
Columbia found that the combined value of
wholesale sales for the six introductions in
1985 was just under $600,000, while a figure
of $1.2 million is estimated for 1986 and with
further increase to $1.9 million for 1987. At
the commencement of the program we had ten
participator nurseries — that number has risen
to twenty-three at the present time.
The following are brief descriptive
notes on our first six introductions.
Genista pilosa ’Vancouver Gold'
An outstanding ground cover or
specimen plants given to the Garden by the late
E.H. Lohbrunner of Vancouver Island. A mass
of intense golden-yellow flowers are borne dur-
ing May on the undulating mounds of plants.
Virtually no seeds are produced and the dead
flowers are quickly hidden by the new growth
that occurs shortly after flowering. Hardy down
to Zone 5.
Container plant of Viburnum plicatum 'Summer
Snowflake . ( Registered COPF cultivar).
photo: author’s
Arctostaphylos uva-ursi 'Vancouver
Jade'
There was a defined local need for a
clone of Arctostaphylos uva-ursi which could
be propagated readily from cuttings, flowered
well and was vigorous in habit with a low sus-
ceptibility to some of the foliar diseases. This
clone was selected and named by the PISBG
program because it met these requirements. The
demand for this plant is by the landscape indus-
try and one participator nursery is now produc-
ing over 150,000 plants annually for this
particular market. Hardy down to Zone 4.
Rubus calycinoides 'Emerald Carpet'
This selection was collected some
9500 feet above sea level in Taiwan and was
subsequently found to be sufficiently distinct
from existing clones to be given a cultivar
name. It is a vigorous, evergreen ground cover,
with a particularly attractive marbled-textured
leaf, that provides a useful alternative to ivy on
specific landscape sites. Hardy down to Zone 7.
Viburnum plicatum 'Summer
Snowflake'
A very attractive compact clone named
by the Garden. A mass of "lace-cap" white
flowers occur in May with recurring flushes of
flowers continuing through into October. The
flowering is followed by fall leaf color of plum-
red shades. This deciduous shrub has been very
popular for retail sales. Hardy down to Zone 6.
Winter 1986 (49:4)
7
Container plant o/Rubus calycinoides 'Emerald Carpet. Registered CORF
cultivar. photo: author's
Anagallis monellii 'Pacific Blue'
This selection has been very popular
with visitors to the Garden. A carpet of eye-
catching gentian-blue flowers occurs during
sunny weather from mid-May through to early
October. Although it is a short-term perennial,
it has performed well as a bedding plant and is a
useful addition for hanging baskets in sunny
locations. Hardy down to Zone 8a.
Microbiota decussata UBC clone
#12701
This was introduced to the British
Columbia nursery industry as a recommended
plant. Originally found in Siberia, it was sub-
sequently introduced to nurseries in Europe.
Our clone was obtained from the Royal Botan-
ical Gardens, Edinburgh, Scotland, but was not
found to be sufficiently distinct to warrant a
clonal name. We have an excellent highway
planting of 400 plants on the outside perimeter
of the Garden. It is becoming more accepted in
British Columbia as a useful alternative to juni-
pers, particularly for shipping to the prairies of
eastern Canada. Hardy down to Zone 2.
The committee is reviewing ideas and
projections to ensure the future success of the
PISBG program. There are sufficient plants for
introduction during the next three years and
work is being undertaken to select others. Var-
ious species are being propagated at the nursery
to evaluate their performance as suitable con-
tainer-grown plants for a typical nursery
production schedule. Plants which grow well
in the Garden could well present difficulties in
production — one example we experienced was
with a weeping form of Paxistima myrsinites
which is susceptible to Phytophthora and Py-
thium when overhead irrigation was used.
We have an open ground nursery eval-
uation and display area where some 230 plants
are established for clonal selection and eval-
uation by the nursery and landscape industry.
This area is also used to show good introduc-
tions from other parts of the world which local
nurseries should consider for inclusion in their
catalogues. Some breeding work is being
initiated — for example, Dr. Gerald Straley, the
Garden research scientist and taxonomist, is
working with some specific perennial plants,
e.g., Meconopsis, Schizostylis and Alstroe-
meria — he is projecting the breeding of a
hardy compact form of the latter.
In conclusion, we hope that the
PISBG program will encourage other botanical
gardens and arboreta to use their collections in a
similar manner. We are pleased that the Royal
Botanical Garden, Hamilton, Ontario, and the
Chicago Botanical Garden have now started sim-
ilar programs. Considerable interest has been
stimulated in Washington State, Oregon and
North Carolina. We will always be very pleas-
ed to share our experiences.
Visitors are always welcome to the
Garden where many of our introductions,
identified by a distinctive blue label, are located
in the different components. Opening hours for
the Garden can be obtained by phoning (604)
228-3928 or (604) 228-4208.
8
UW Arboretum Bulletin
In The Arboretum
For the past month. Grounds Super-
visor, Richard Hart, has been recovering at
home from quadruple-bypass heart surgery.
Last report is that he is making rapid progress,
walking two miles or more each day. We
expect Dick's return in early 1987. Meanwhile,
Phil Renfrow is doing a great job as acting
supervisor.
Two new faces appeared in the Visitors
Center in October. We welcome Laurence A.
Smith, "Smitty", a retired pipefitter/merchant
engineer, who now works as part-time
custodian here, as well as at the Center for
Urban Horticulture. The new program assistant
for the Visitors Center, replacing Eric Nelson,
is Dan Dewald. Dan, his wife and two chil-
dren, recently moved to Seattle from Idaho. He
comes from a forestry and wildlife management
background. Dan says he is finding his new
job working with all three organizational
bodies of the Arboretum: the City, the Uni-
versity, and the Foundation both interesting and
challenging.
The bulk of work performed by the
Arboretum grounds crew has involved reno-
vation of various areas by pruning or removal
of overgrown and invasive native species. Take
a look at the newly exposed rockery along Lake
Washington Boulevard across from the Emper-
ors' Gate to the Japanese Garden. This project,
spearheaded by Phil Renfrow and assisted by
Bob Hilzinger, has brought delight to those
onlookers who are (re)discovering the existence
of this landscape feature. Future plans are in
the works to highlight this area with appro-
priate new plantings. Not far from the rockery,
a complete renovation of the southeastern-most
bed of Azalea Way has been undertaken. John
Hushagen along with Phil Renfrow detail-
pruned the larger species, such as Acer
monspessulanum. David Zuckerman and Bob
Hilzinger did a fine job reshaping the azaleas
and other shrub materials in this bed as well as
in other beds farther north along Azalea Way.
Fred Mauch has been concentrating on pruning
out the native materials south of the Look-out
Some of the unusual conifer specimens in addi-
tion to Daphniphyllum macropodum are now
exposed to a better view. Thanks are extended
to Paul Wiltberger, who donated a day's work to
the Arboretum. Assisted by David Zuckerman,
Paul's service included pruning of Douglas fir
trees that were impacting the Stewartia area of
the Camellia section. All of these projects
have brought about an improvement to the
Arboretum's appearance.
The interior of the Visitors' Center has
taken on a new look as well with the recent
addition of unusal plants brought over from the
Arboretum's greenhouse by Dean Powell. Dean
is happy that some of the more interesting
plants such as the Dioon edule and the Solandra
maxima have been accepted into this new
home where all Arboretum visitors may appre-
ciate them. The greenhouses are close to being
empty now in order that renovation can take
place. By the way, the oldest member to leave
the greenhouse was a 47-year old Pyrostegia
venusta
Dean also cleared out a new nursery
bed just north of the existing ones with the
assistance of Fred Hoyte and his tractor from
CUH. This has allowed me to begin the vast
movement of plants from coldframes to lath
beds or nursery area. The result is that the lids
have been placed on top of the coldframes,
winterizing them. Now with my wrist in a cast
for six weeks, I've turned my efforts towards
inventorying the lath house along with Jan
Pirzio-Biroli.
Looking ahead to winter, we all hope
that the excessive fall rains do not continue,
otherwise the trails — especially Azalea Way
— will become impassable without use of high
rubber boots or amphibious vehicles.
Barbara Engler
Gardener in the
Arboretum
Winter 1986 (49:4)
9
Small British Gardens
Some Personal Favorites
DENNIS THOMPSON
To the British SMALL and GARDEN
are almost contradictory terms. They seem to
believe anything under 10 acres is a small
garden. Definitely, any garden under three acres
is small. The American small, MY small, is a
garden that would fit on a Seattle lot. For the
British garden enthusiast, this is nearing
minute. The gardens I present in this article are
not ones that appear in the tour guides, yet
many are impressive, unrecognized national
treasures. They fall into almost every
gardening category: public gardens, commercial
gardens and private gardens.
Public Gardens
Old London has many tiny landscapes
in hidden comers. Several of my favorites are
in conjunctions with churches designed by
Christopher Wren, architect for 53 parish
churches in the area after the Great Fire in 1666
One such is a small courtyard of raised beds at
St. Anne and St. Agnes which adjoins the
crumbling remains of the city's outer wall.
The carefully planted beds meld into the
wildlings scrambling in the old wall. Annual
bedding plants blend into butterfly-bush,
kenilworth ivy, English ivy, wallflowers.
plantains and ferns. Several other Wren
churches have sheltered churchyards or entries.
Trees and shrubs, green elegance against old
stonework; noontime parks where area workers
eat quiet lunches.
The most exciting small garden in Old
London is a small park built around a Wren
church that did not survive the blitz of World
War II. The tower and steeple of St. Dunstan-
in-the-East created furor from the beginning.
Radical design variations brought prophecy of
impending doom. In a biography of Wren
written in 1920 architectural critic, Lawrence
Weaver, notes:
The tower of St. Dunstan-in-the-East is unique
for its date, and of quite extraordinary interest:
any idea of removing it should be resisted with
vehemence. But why anyone should be the least
concerned with the disappearance of the body of
the the church is known only to those who
detect beauty in the Gothic adventures of
Mssrs. Laing and Tite in the Year 1810.
What the author was unable to accomplish with
his prose in 1920, the Germans accomplished
1 Weaver, Lawrence. Sir Christopher Wren:Scientist,
Scholar and Architect. Offices of "Country Life",
London, 1923.
10
UW Arboretum Bulletin
St. Dunstan-in-the-East.. photo: author's
in the early 1940's. The tower and walls were
left standing but the interior was gutted. Old
London no longer needed "twenty-eight
churches. ..within the square mile of the City",
and chose not to rebuild the chapel. In the
1970's a designer with the City conceived an
award-winning plan for the old structure. The
tower was turned into a museum and the body
of the church into a park.
My first glimpse of the garden was of
a large blue Ceanothus impressus jutting
through the stone arches of an empty Gothic
window frame. It is hard to describe the
impact. Romantic. Breath-taking. Wrought
iron gates open into the small, tree-sheltered
north yard. New office buildings soar over the
churchyard making the area seem even smaller.
Immediately next to the stone walls is an area
paved with round cobbles in cement with a
narrow fringe of bracken frothing from a chink
between wall and paving. A path bends as it
enters the chapel through an old door arch.
Winter 1986 (49:4)
11
Perennial summer phlox. Phlox paniculata.
Inside the bombed-out church is a somewhat
anachronistic lane that winds through a small
cr
lawn with bedding plants and leads to a cobbled
seating area with a fountain. The garden is de-
signed to be viewed from this seating area and
from the small paths that meander around the
walls and through arches at both ends of the
chapel. This area is planted with an amazing
variety of vines that cling to ruined walls and
drape over and tangle through the many win-
dows. Clematis, Hydrangea. Ampelopsis,
Parthenocissus , Campsis, Lonicera, Jasminum,
Schizandra, Polygonum, Rhodochiton, Vitis
and Rosa mingle in almost indistinguishable
confusion; distinct only in bloom. Other
climbers exhibit their uniqueness by extra-
ordinary foliage like the lusty leaves of Schizo-
phragma hydrangeoides and the opulent, velvety
green-black dissected leaves of Parthenocissus
henryana w ith their silver veins and flashes of
pink. The warm gray stone w alls provide a
variety of exposures and backgrounds. For a
melange of material, the overall effect is
amazingly unified. Perhaps it is scale and
intimacy that allows the success with so much
variety. Birds as well as people seem attracted
to the site, an oasis in an urban desert.
Commercial Gardens
There are a number of commercial
businesses that have pleasant small gardens.
Nurseries frequently include display gardens and
two small nurseries that have interesting little
gardens are County Park Nursery at Hornchurch
east of London and Edrom Nursery in Scotland.
County Park is entered from a street
w hich gives no hint that either nursery or gar-
den exist. Graham Hutchins, the proprietor,
has a mania for New Zealand plants. Stepping
through the gate is like entering the Twilight
Zone. The grotesqueries of the plant world
seem to have gathered for a meeting. The leaf-
less Clematis afoliata looks like a climbing
Scotch broom. The leafless, brown-felt stems
of Chordospartium stevensonii growing in a
raised bed resembles a Daliesque weeping wil-
low. Pots of Coprosma look like baby's-tears
under vegetable marbles. Beds, shelves and
houses are filled w ith similar rarities — Hebe,
Helichrysum, Aciphylla, Carex, Cotula and
Raoulia. There are examples of award-winning
miniatures (barely portable) exhibited at the
Chelsea Flower Show. Mr. Hutchins has devel-
oped a variation on hypertufa (concrete-peat
containers) to create tall, totem-like peatbeds
for acid loving plants.
Edrom Nursery is much more tradition-
al. The display garden is composed of stone
troughs located behind the sales area. It con-
sists of a wide variety of alpines, for sale in
frames, w hich have been grown to perfection in
the sinks. Large troughs of select Gentiana,
Primula, Dianthus, Phlox. Lewisia, Androsace
and Saxifraga make barbaric displays, each in
its season.
Other commercial ventures with
gardens include hotels and crafts shops. The
Swan in Lavenham is an excellent example of
gardens as outdoor rooms. The hotel is assem-
bled from four 15th century housed joined
together in the 17th century. It includes a small
interior courtyard and a comfortable patio gar-
den as a portion of the bar. The gardens do not
contain unusual materials, but these green
rooms enhance the charm of the hotel. The
quietness of the gardens is a marvelous comple-
ment to the half-timbered and plaster residence.
Private Gardens
It is difficult to choose among small
private gardens, but those of four gardening
families would always make my list of favor-
ites, no matter what larger gardens could be
added. They are the gardens of the Becketts, the
Leeds, the Maules and the Fishers.
Kenneth Beckett's home in King's
12
UW Arboretum Bulletin
Alpine house, rock wall and border in Mrs. Maule's garden.
photo: B.O. Mulligan
Lynn, Norfolk, is a 2-story warm brick with a
very simple front entry of tucked flowers. The
low beam in the old house necessitates an
occational duck. The Dickensian book-lined
library /workroom opens through French doors
onto a crazy-paved, sunken patio with planted
urns and cracks. Great tumbling shrubbery
rolls across the garden to the hedge which
allows only a glimpse of the flint church in
back. A small greenhouse at the side of the lot
holds collected treasures — blue Tropaeoleum
and rare bulbs. The garden’s fauna include
Kenneth’s son's guinea pigs. Sitting in the
library, looking into the garden creates the
warm feeling of being wrapped in peaceful,
protective green.
Chestnuts is the home of Jane and
Rodney Leeds in Sudbury, Suffolk. The view'
from the road through a tall shrub border pre-
sents a comer view of the thatch roof and cream
stucco reminiscent of a Helen Allingham’s
painting. The lawrn path winds between serpen-
tine beds and shrubs, perennials and alpines
which grow in front of the house. The Leeds
cultivate, in the ground and in containers, a
wide variety of plants from seeds. At the far
end of the bouse is a small greenhouse and
paved patio filled with perennials in containers
and troughs of alpines. The rear garden is the
largest and includes a sunny lawn with fruit
trees. It is surrounded by beds including Marta-
gon lilies and Cardiocrinums. A large lattice,
partially covered with vines, screens the vege-
table garden and more perennials. A cluster of
cold frames just outside the vegetable garden
protects seedlings and tender perennials. Behind
the frames, a large compost pile is bed to one
of the garden cats. This garden, as with many
British gardens, is ruled and guarded by a rank
of garden cats. They are very polite, enticing
people about the garden for appreciation and
photos. Chestnuts is among the best of the
miniaturized "estates".
Sheila Maule's Hannahfield Quarry
Home does not technically fit into this group of
gardens. It is in an old five acre stone quarry
just outside of Edinburgh, but less than half of
the area is under cultivation; the rest is segre-
gated by rabbit fencing to keep these marauders
back. I justify its inclusion by the fact that
there was no soil in the area when the Maules'
began working with the garden. Sheila hauled
all of the soil into the area in the "boot"
of an old car. "Ruined the coupe", she always
Winter 1986 (49:4)
13
photo -.author's
Eleanor Fisher's garden Colthurst.
giggles. Much of the area is paved with stone
in order to hold enough soil in to be able to
garden. The lawn sits on bare inches of soil
and berms appear in areas where deeper roots
had to be accommodated.
The wood frame cottage, an anomaly
amid the stone and plaster of Scotland, faces a
one acre pond (deeper quarry hole) and steep
cliffs of the outer quarry wall. The cracks in
the stone have been planted with alpines and the
rock chip talus with perennials. Masses of
native primroses and naturalized daffodils wel-
come spring. Raised bulb frames shelter plants
collected in Spain, North Africa and the Near
East. Cold frames and alpine houses protect
buns and polesters from around the world as
well as masses of new seedlings. Among the
naturalized plants in the garden are Asiatic and
alpine Primula , Kirengoshoma, Morisia,
Tropaeolum polyphyllum , Japanese orchids,
Trillium, Lewisia and many other small exo-
tics. Sheila is one of those growers who seem
able to manage almost anything without spe-
cialized modem equipment. She is the only
person I know to have ever grown our Saxif-
raga tolmiei. Her propagation chamber for
cuttings is an old traveling trunk, half filled
with sand and covered with a windshield glass.
No heat. No mist. Only skill. The side patio
is primarily a display area for alpine troughs.
In front of the house is a table made of an old
stone grinding wheel, and besides the small
formal pool is a finial from John's ancestral
home which serves as garden sculpture. A
gardener’s garden, the effects of loving care are
obvious. As a fiftieth anniversary gift, John
gave Sheila another 200 feet of rabbit fencing!
Colthurst is the second garden of
Eleanor Fisher. Besides caring for two teenage
sons and a husband, plus volunteering weekly
at the Harlow Car Botanic Garden, she main-
tains one garden at home and a second at their
country house. The latter was converted from
the old stables of the existing estate, thus the
name colthurst. The stables and the adjoining
walled kitchen garden have been modified as a
unit to provide house and garden. The house is
a mellow stone and wraps around two sides of
the garden. Eleanor is known as a master
plantswoman, specializing in perennials and
companion plants. The color and form
combinations she has assembled are breath-
14
UW Arboretum Bulletin
Colthurst, green and white color combination. photo: author's
taking. She plans the garden to progess
through the seasons with new color schemes
against a background planting. Among my
favorite groupings are Otto Luyken laurel in
bloom with white lily-flowered tulips. Mid-
night purple and white Rembrandt tulips and
purple wallflowers growing through purple-
leaved sage produce a striking package, as do
the soft pink peonies and shrub roses. When it
was time to move the greenhouse out of the en-
closed garden, the foundation was not easily
disassembled, so the door was sealed up and the
foundation became a raised garden pool. Few
gardens, no matter how large, have the stun-
ning staging or effective plant groupings of
Colthurst.
It is difficult not to continue on,
considering the "slightly larger" small gar-
dens; Jenny Robinson's Chequers , Dily
Davies' Greyrigg , Jean and Jack Elliot's
Coldham, Kenneth Armistead and Jim Ellis'
Bell Cottage and numerous others. But gar-
dens are like pictures of grandchildren and
summer vacations — best presented in small
doses in hopes of another invitation!
Winter 1986 (49:4)
15
Gardens of the Welsh Border Country
SUE BUCKLES
Sue Buckles is a gardener on the grounds of Childrens' Hospital and
Medical Center. As one of a crew of five, she is involved in
all aspects of the garden's maintenance , but is particularly interested
in perennials.
The Powis Castle Gardens have been
the subject of talks I have given since returning
from a three month working period there in
1984. A group from the Northwest visited the
gardens last year, as part of a garden tour led by
city arborist Marvin Black, so it is one Nation-
al Trust garden that is familiar to many Arbor-
etum Bulletin readers.
Rather than describe Powis again, I
would like to describe some its workings, and
to suggest other gardens that can be seen in the
area of England's western counties and the mid-
Welsh border country in which Powis stands.
Powis castle is a suitable central point for any
tour, standing high on its rocky ridge, and over-
looking the town of Welshpool in the Welsh
county of Powys.
Practices in propagation and main-
tenance carried out at Powis may be of interest
to gardeners here. No irrigation is available in
the gardens, and the many large stone ums plan-
ted with annuals are watered by hand only occa-
sionally. The beds, therefore, are mulched
heavily in spring with compost which contains
only partially decomposed organic matter
(small sticks, leaves, pieces of bark, etc.).
Their compost comes from large compost
piles, about 15 x 20 feet, into which is thrown
all garden clippings, leaves, straw, loam and
manure, but no weeds. This compost is not
used for potting soil, which comes from ster-
ilized loam, peat and sand. The potting soil
varies in the balance of its components accord-
ing to need. There are set proportions for cut-
tings, seeding, potting on and for the contain-
ers, which are lined with a coarse peat and in-
clude a great deal of coarse humus matter,
which helps to retain water over long periods.
The perennial borders have many new
plants put in every year. I suppose these would
be called bedding plants as many are tender and
have been propagated by cuttings and held over
the winter in the greenhouse. While I was
there many new plants were used to fill new
beds which had been made the previous year and
prepared for the spring planting. The old estab-
lished beds are filled with new cuttings and divi-
sions from the established beds plants. Annual
cuttings are taken from plants such as Cheiran-
thus 'Wenlock Beauty' and Erysimum 'Bowles
Mauve', Anthemis cupaniana, Chrysanthemum ,
Helianthemum, Achillea and Dianthus. The
16
UW Arboretum Bulletin
The Ideal English Gardener
The head gardener at Powis Castle is Mr. Jimmy Hancock. He has just been awarded the
Royal Horticultural Society Associateship of Honour, given to "persons who have rendered
distinguished service to horticulture in the course of their employment." This is a well-deserved and
appropriate award. Mr. Hancock has a permanent staff of about five, and a very large garden to
maintain. His main activities are carried out in the greenhouses and borders, he propagates plants
for the beds and containers as well as for the popular plant sale held every day the garden is open.
He has an under-gardener whom he has trained to be in charge of the greenhouse area. In the area of
the glasshouses are various sheds and a large polythene covered tunnel which gives frost protection
to container plants. Mr. Hancock is often to be seen there late at night, working at the propagation
bench with a flashlight fixed above his head. He was there when I arrived at Powis, and was taken
to the greenhouses to meet him, and he was there when I left. I expect to find him there again when
I return on a visit this year. His large family would know where to find him and would send down
the youngest boy to tell him, "Dad, your supper's ready". Fifteen minutes later the next youngest
would come, "Dad, Mum says you've got to come for your supper". Finally, the eldest daughter, as
she passes by on her way out- "Dad, you've missed your supper, G'bye."
Jimmy has various students from horticultural colleges sent to do their time as working
apprentices. Then there are young people working for a year on one of the government training
schemes for unemployed youth, and older people on similar re-training programs. They have to
attend classes one day a week at the nearest community college. To all of them, skilled or
unskilled, keen or dispirited, bright or not so bright, Jimmy encourages, teaches, helps, kids along,
inspires those who have found gardening to be a sudden wonderful new occupation.
I was included in this grand mixture, put to work straight away, and offered ready answers
to all my questions. Those who know something of the chauvinistic nature of British head
gardeners, and the conservative character of the horticultural profession, will realize that Jimmy's
approach is rare and valuable in one who employs so many young gardeners.
tender plants included many of the Salvias —
S. patens , S. hians , S. uliginosa and S.farin-
acea , Mimulus glutinosus, M. luteus and M.
guttatus, Felicia amelloides, Osteospermum ,
Convolvulus cneorum , Jasminum primulinum
and Solanum jasminoides. Plants which pro-
vide the beddings familiar to us here are used in
groupings and for containers — Nicotiana,
Pelargonium, Fuchsia and Helichrysum.
The plants at Powis are extremely
varied. Half-hardy and even more tender spe-
cies can be used because the gardens are protect-
ed. They face, on the whole, south-east, and
are situated on a hill from which the frost drops
away. Not so the gardens at Burford House,
attached to John Treasure's nursery in Tenbury
Wells, an hour's drive from Powis.
The gardens at Burford House sit in the
frost hollow of a river valley and are frost free
for only 3 or 4 months of the year. Here you
may see unusual and adventurous gardening.
The national collection of Clematis is here, and
the visitor sees this lovely vine used in many
ways — threading through shrubs (dark red
Clematis 'Niobe' in variegated dogwood) and
carpeting the ground between perennials, the
garden contains many old roses, and I found
particularly interesting the use of stream-side
plants and moisture lovers. Varieties of wil-
lows line the stream on the far side of the gar-
den and give a background to Thalictrum ,
Rheum and Primula.
Burford House is a privately owned
garden, open to the public most afternoons, but
the attached nursery is open all day (with the
ubiquitous tea-room!) Unfortunately, Amer-
icans cannot take plants back to the States
without a good deal of trouble, but the nursery
is still a wonderful place for looking and learn-
ing. It can get very crowded, especially on
weekends — mornings are best, and then one
can spend the afternoon in the garden. Most
Winter 1986 (49:4)
17
Above: Helichrysum
Right: Anthriscus
other gardens in the area are only open in the
afternoon as well, making it difficult to com-
bine several visits in one day. However the
surrounding countryside is beautiful. There are
many areas which deserve a walking tour rather
than a driving through and many mornings can
be spent walking the lanes or across low hills.
Another hour from Powis, half an
hour from Burford House, is the lovely garden
at Hergest Croft at Kington, Herefordshire,
which belongs to the Banks family. This is
also a private garden, open in the afternoons,
and best seen in spring for there are many rhodo-
dendrons and azaleas. Here are the national col-
lections of maples and birches, and with these
are many other beautiful trees.
Besides these three gardens, which pro-
vide a tremendously wide range of styles and
plants, there are cottage gardens, scenic views,
and wild flowers in the hedgerows all around
Powis and Hergest Croft . Halfway between
them is a mediaeval manor house, Stokesay,
worth a visit if only for the wild flowers grow-
ing in the moat that surrounds the fortified
house. These flowers have been encouraged,
judiciously thinned and weeded by the custo-
dian, a gardener. Here you can see the master-
wort, Astrantia major , which is native to
Britain and Europe but very rare now in the
former. In the lanes about Powis can be found
the lovely cow parsley, Anthriscus sylvestris ,
Smyrnium olusatrum or Alexanders, not often
seen inland, and Aconitum anglicum , also rare-
ly seen now as a true native. Around Hergest
Croft, in amongst the grasses of the ditches one
can see Squinancywort ( Asperula cynanchica )
and the yellow pimpernel ( Lysimachia
nemorum).
There are as many good things to see
on the northern and western sides of Powis.
Chirk Castle has a good garden, and Erddig a
fascinating conservation project — lam just
giving you a good start. Stay in the little town
of Welshpool, by Powis, or Kington, by
Hergest Croft, and you will be able to say to
yourself, "If it's Tuesday, this must be the
same place I was in yesterday, and I'll still be
here tomorrow — Joy!"
18
UW Arboretum Bulletin
Top: View from RA. Banks' house. Bottom: Mr. John Treasure's garden. photos: B.O. Mulligan
Winter 1986 (49:4)
19
Wave Hill
VIRGINIA MORREL
On a visit to New York City we
expect to be caught up in the hustle and bustle
of this huge city's life, and so need even more
the sanctuary of a delightful and alluring park
away from the crowds and noise. One such
place is Wave Hill in the Riverside section of
the Bronx, overlooking the Hudson River and
the Palisades. It was an estate owned by
George W. Perkins, whose widow gave twenty-
eight acres and two mansions to the city of
New York for a city park and cultural center in
1960. The garden is comprised of fifteen acres,
and eighteen acres remain wooded. Former resi-
dents of Wave Hill include Mark Twain and
Arturo Toscanini.
Shortly after Mr. Perkins bought the
property in 1903 he hired Albert Villard, a
royal landscape gardener from Vienna to design
and develop a garden. Today an innovative and
imaginative series of gardens have been created
within the basic framework of the original
garden by Marco Polo Stufano, Director of
Horticulture, and John Nally, Curator, and their
staff. Upon arriving at the gate you step back
in time to a Flower Garden filled with flowers
popular in 1910 and 1920. There are numerous
dianthus, peonies, clematis, iris, pulmonarias,
lilies and artemisias, among others.
In front of the conservatory is a garden
of all gray foliage plants, said to be the largest
collection of gray plants in the eastern United
States. One greenhouses is filled with a cactus
collection, while another houses tropical
plants. A new solar greenhouse contains a
large collection of rare alpine and rock garden
plants. This is the only alpine collection open
to the public in the East. In the foundation
ruins of an old greenhouse there is the Herb
Garden with over 100 different herbs. A Dry
Garden has been started within another old
stone greenhouse foundation. Many half-hardy
plants, such as zauschnerias, phlomis, euphor-
bias, origanums and rosemary are being grown
there.
Up a steep slope, further along the
path from the Dry Garden, is the Wild Garden.
It is "wild" only in the use of the plant mater-
ial. Around each curve in the winding path up
to the summerhouse is another exciting visual
treat. Crambe maritima , a frothy white blos-
soming wild kale from Britain grows happily
in an environment far different from its native
habitat. Yucca glauca grows along with Asclep-
ias tuberosa (self-seeded). Verbascum bomby-
ciferum, Zauschneria calif ornica and Glauciums
share space with a number of Carex. Interming-
ling are Iris pseudacorus 'Bastardii' and I.
chrysographes 'Black Form' with Eremurus ,
small flowered clematis, Arctostaphylos uva-
ursi , Pinus aristata and cacti. Plants which
20
UW Arboretum Bulletin
The Wave Hill herb garden.
have never been associated intermingle in an
exciting experiment. The Wild Garden is an
area where Stufano and Nally dare to do the
unusual, and get away with it.
Christopher Lloyd wrote "What a gem
their pond garden is", to which he must have
referred to the textures and forms of the grasses,
reeds, water lilies and arums used. This garden
is located at the flat area at the base of the Wild
Garden. It is a peaceful spot where an occation-
al heron comes to fish, and people sit and relax,
enjoying the tranquility.
A new Monocot Garden in the curve
by the pond near the pergola is planned for the
spring of 1987. There will be mainly grasses,
bamboos, lilies, species Iris and the like. On
the pergola grow a most imaginative selection
of vines such as gourds, morning glories,
moonflowers and cardinal climbers.
A Symmetrical Garden has been
created for displaying interesting color from
early spring until fall. This garden is made of
shrubs and roses, as Stufano says "in a Sissing-
hurstesque manner".
Great expanses of lawn and huge old
photo: Timothy Hohn
trees separate these gardens. There are oaks,
maples, magnolias, elms, parrotias and pines,
as well as several Idesia polycarpa with its
interesting fruit, a tree for which Wave Hill is
particulary well known.
Incorporated in all this, scattered about
on the grounds, are large pieces of sculpture, on
loan from the artists, which are changed from
time to time. On going exhibits of garden de-
sign, horticultural displays and musical pro-
grams are presented in Wave Hill House, along
with a variety of educational classes, workshops
and training classes for elementary school
teachers. The collection and memorabilia of
Arturo Toscanini, and 137 tapes of the broad-
casts the maestro led with the NBC Symphony
Orchestra are housed in the Wave Hill Library.
The entire garden is an imaginative,
fascinating, daring creation. The great care
given to color, form and texture reflects
Stufano's artistic background. The continual
seeking out, and use of unusual plant material
makes this garden a plant lovers delight. Wave
Hill has become an inspirational, exciting, yet
relaxing sanctuary in this country's largest city.
Winter 1986 (49:4)
21
A Collection of Botanical Treasures —
The University of Washington Herbarium
MELINDA F. DENTON, CURATOR
Early in man's recorded history, the
search for information concerning plant distri-
butions and diversity was based on a need to
know more about drug, medicinal, or food
plants. The Chinese, for example, had floral
pharmacopoeias as early as 5000 B.C. Later,
the Hebrews, Babylonians, and Greeks produced
herbals that dealt with medicinal substances de-
rived from plants. Given the concern for medi-
cal applications, it is no surprise that physi-
cians were the first taxonomists (persons who
study the kinds of organisms to determine their
identity and extent of variation and to arrange
them in an orderly sequence or classification).
Their medical skill depended in large measure
on an ability to know which plants could be
used to treat various illnesses; their gardens
were their laboratories. By the early 1700s,
physicians and estate-holders were cultivating
many kinds of plants and subsidizing exped-
itions of adventurous collectors who would
bring back herbarium specimens, seeds, and
exotic plants from remote parts of the world.
Plants were preserved for identification pur-
poses or for general reference by making dried
pressed specimens, a technique developed in
Italy in the 1500s. The earliest herbarium
specimens were bound into books; later, single
herbarium specimens were filed systematically.
The increase in knowledge of plants only
increased the appeal of unusual ornamentals
as well as the desire to acquire more of the food
and drug plants that grew naturally elsewhere.
It soon became evident that classifications were
necessary to provide some framework for depict
ing plant relationships so that identifications
could be made readily.
In 1623, Caspar Bauhin published
Pinax Theatri Botanici , which was a compil-
ation of information concerning all plants
known to exist at that time. By the mid-
1700s, Carolus Linnaeus constructed an artifi-
cial system of classification of flowering
plants, based primarily on the number, arrange-
ment, and fusion of male and female reproduc-
tive parts of the flow'ers; this was a landmark
contribution in that it arranged the plants
known to him in an easily interpretable se-
quence. Linnaeus and other taxonomists of the
17th and 18th centuries usually did not see
more than one or a few specimens of a given
species and did not realize the extent of varia-
tion that usually exists within a species. Even
though their taxonomic decisions were not
based on an appreciation of the processes of
evolution or populational variation of species,
the framework for modem studies in plant
systematics was laid by these early taxono-
mists.
Today, plant systematics (encompas-
sing taxonomy) aims at understanding the
nature of plant variation and phylogeny
22
UW Arboretum Bulletin
Workroom for receiving and fumi gating plant specimens.
(pertaining to evolutionary' relationships within
and between groups) of taxa (whether species,
genera, families, orders, or classes). The inten-
sive study of a group of related species provides
an understanding of the w av those organisms
C J
likely have evolved and documents the nature of
their phylogenetic relationships. One might ask
the value of know ing if plant species A is relat-
ed to plant species B. The answer becomes
especially apparent when plant species A has an
important medicinal, nutritional, or ecological
property. Plants that are phylogenetically
closely related tend to be similar in their chemi-
cal and physiological properties. The resources
for an important product can be increased if
more than one species w ith the property can be
obtained. Such information usually is avail-
able only if intensive monographic studies that
evaluate the phylogenetic relationships of
plants and the processes that have led to diversi-
fication have been made. In a monographic
study, all aspects of the growth and develop-
ment of the included species should be invest-
igated to provide a full understanding of the
way in which the species have adapted (changed
genetically) and evolved.
Systematic studies rely on examin-
photo: author's
ation of numerous plant specimens made of the
species being studied. To be thorough, speci-
mens that have been collected throughout the
known range of geographic distribution should
be examined. Plant specimens, housed in her-
baria or plant museums, document the existence
of particular kinds of organisms or species at a
given time in a specific place; they are vouch-
ers. The preserv ation of vouchers in a herbar-
ium allows researchers, present and future, to
study the extent of species variations as well as
to check or confirm the results of others. The
continuing refinement of scientific techniques
enables researchers to use herbarium specimens
for a w ide variety of biological studies that seek
answers to diverse questions or hypotheses.
For example. DNA from plants preserved as
herbarium specimens now' can be extracted and
used for comparative analyses. In some species,
seeds can be removed from herbarium speci-
mens. germinated, and used for a particular
study w here live materials are needed for chro-
mosomal, ecological, genetic, or chemical anal-
yses. In all cases, examination of a represen-
tative set of herbarium specimens for a given
species can readily depict the range of morphol-
ogical variation and distribution pattern.
Winter 1986 (49:4)
23
Plant specimens representing a holotype(the specimen
on which a name of a plant is based; in this case, the
type o/Sedum laxum subspecies flavidum.
photo: author’s
Since rare and endangered plant species
are now of state and federal concern, the deter-
mination of the status of a plant species (rare,
endangered, or threatened) depends on the
thoroughness with which plant exploration and
collecting have been carried out in a region. If
collecting has been extensive in all areas, for
example in Washington state, then one can ade-
quately assess the frequency and locations of
populations of the native species. Upon exam-
ination of the collections kept in herbaria, the
geographic distribution is determined. If all or
nearly all potential habitats have been explored,
then one can assume that the distribution pat-
tern of a given species determined by the exam-
ination of herbarium specimens is accurate.
While obtaining distribution data, the identifica-
tion of the specimens examined always should
be checked to make certain that errors of identi-
fication or filing were not made previously.
Such verifications provide a clearly documented
basis of the distribution pattern of a plant
species.
For the state of Washington, when a
flowering plant species is known to be rare or
endangered, the location of each population is
noted from the herbarium specimen and the
information sent to the Natural Heritage Pro-
gram, now affiliated with the Washington State
Department of Natural Resources in Olympia,
Washington. The locations and sizes of the
populations of these species are currently moni-
tored by the Heritage Program. Information on
the status of rare and endangered species and of
other native species that are dominant in particu-
lar plant communities is used by the Washing-
ton State Natural Areas council and groups
such as the The Nature Conservancy to deter-
mine priorities for habitat preservation and land
management and acquisition.
Once the accumulation of specimens
in a herbarium for a given region is represen-
tative of the existing plant life, floras or man-
uals can be written. These volumes are essen-
tial for general plant identification of the local
plant life and for documenting the geographic
occurrences of plants. (The term flora can be
used to refer either to the plant life of a given
area or to a published work describing the plant
life of a given area.) Consultation of available
floras or manuals allows one to determine or
estimate the overall distribution pattern of
particular species, which, in turn, provides data
for understanding the phytogeographical rela-
tionships of plants. Floras, however, are never
really complete. They require periodic revision
to account for newly found species as well as
those that disappear either by natural or man-
made means (usually by alteration or elimina-
tion of a habitat). In most floras, introduced
plants are not included unless they have become
naturalized. In the Pacific Northwest, we have
a model flora (The five-volume illustrated
Vascular Plants of the Pacific Northwest by
C.L. Hitchcock, A. Cronquist, M. Ownbey,
and J.W. Thompson) and a manual ( Flora of
the Pacific Northwest , by C.L. Hitchcock and
A. Cronquist). The collections that document
these major works are housed in the University
Herbarium.
The University Herbarium was founded
in 1879 by the Young Naturalists Society in
Seattle, an organization that was active up until
1905. All of the early collections were incor-
porated into the holdings of the Washington
State Museum (now known as the Thomas
Burke Memorial Washington State Museum)
located on the University campus. Many
collections of native plants were made by
24
UW Arboretum Bulletin
early members of the society and by faculty
members at that time. By the early 1900s, the
Herbarium had started to expand and to be used
as a resource for research and teaching. The
Herbarium consisted of about 30,000 speci-
mens in 1937 when it was transferred to the
Department of Botany where the faculty, main-
ly C.L. Hitchcock, could use the specimens
more intensively in research and teaching. As
Curator for 35 years (1937-1972), Hitchcock
devoted most of his energies to building the
Herbarium through an active collecting and
exchange program. By the time of his retire-
ment, the Herbarium had grown to about
400,000 collections.
As Curator since 1972, 1 have devoted
most of my efforts to strengthening the hold-
ings of certain plant groups or families, and
improving the physical layout and accessibility
of the collections. Since 1980, we have had an
assistant curator (Bonnie Tucker, 1980-82;
Anna Zeigler, 1982-present) and undergraduate
hourly support to help with daily herbarium
operations. Over the years, we also have had
invaluable assistance provided by volunteers.
We now have over 510,000 collections of
which 60% is representative of Pacific North-
west taxa. Included in these are over 1,500
type specimens; each serves as the type of a
scientific name. Considering all groups of
plants, we believe that we have the best repre-
sentation of native plants from the region that
can be found in any museum or herbarium. In
brief, the composition of our holdings is:
324,000 vascular plants (ferns and allies,
gymnosperms, and flowering plants), 72,000
bryophytes (mosses), 16,000 hepatics
(liverworts and allies), 21,000 algae, 1 1,000
lichens, and 50,000 fungi. These specimens
were collected by many important collectors,
the more notable being W.H. Baker, W.C.
Cusick, W. J. Eyerdam, T.C. Frye, C.L.
Hitchcock and C.V. Muhlick, G.E. Howard,
D.E. Stuntz, W.N. Suksdorf, and J.W.
Thompson. Today, our acquisition policy
emphasizes plant species that occur in remote
areas of the Pacific Northwest and western
North America in general.
In 1983, the University Herbarium
was moved from Johnson Hall to the new
biological sciences building on campus, named
C. Leo Hitchcock Hall. Along with the move
to new facilities, we were able to consolidate all
holdings and provide for a 35% increase in
storage space. The improvements in physical
layout and access to the collections enable us to
handle special requests and accommodate the
needs of visitors more readily than we could
previously. With improved representation of
plant groups, the value of the herbarium is
increased for purposes of teaching, basic
research, and for obtaining information useful
in land management and conservation.
The Arboretum Foundation
Used Book Sale
at the
Donald G. Graham Visitors Center
Take this opportunity to stock up on spring & summer reading!
Friday, March 6th — Arboretum Foundation members' Preview Sale 5:00-8:00 p.m.
Saturday, March 7th & Sunday March 8th — Public Sale 10:00 a.m. to 4:00 p.m.
Books you wish to donate may be brought to the Visitors Center on Wednesday, March 4th and
Thursday, March 5th.
Winter 1986 (49:4)
25
The Donald G. Graham Visitors Center
At the request of the Arboretum
Foundation, the Seattle Parks Department has
officially named the new building in the Wash-
ington Park Arboretum the Donald G. Graham
Visitors Center. The Arboretum Foundation
feels that this is a most fitting recognition of
Mr. Graham's devotion to horticultural causes,
dedication to the establishing and development
of the Arboretum, and the generous financial
support that has continued from its beginnings
in the 1 930’s to the building of the Visitors
Center itself.
Donald G. Graham was bom in Fort
Worth, Texas in 1884, and came to Seattle to
practice law in 1921, following his marriage in
1919 to Juanita Fisher. Donald Graham's
horticultural interests were well recognized by
1934, when minutes of the September 6th
meeting of the Board of Park Commissioners
record his appointment to the temporary com-
mittee "to make recommendations as to financ-
ing and plans" for the "Arboretum Project".
Also in 1934, Donald Graham was appointed
chairman of an "Arboretum Advisory Council".
One of this Council's first actions was to estab-
lish the Arboretum Foundation — to serve as a
repository for funds and to raise revenues for
the Arboretum. At this time Juanita Graham
was instrumental in the formation of 18 auxil-
iary support units to the Foundation and served
as the Unit Council President from 1938-1941.
Unit 1 is named the Juanita Graham unit, and
after fifty years, Mrs. Graham continues her
involvement and support of the 75 active and
productive units.
In 1938 and 1939 there were difficul-
ties in resolving and financing the fledgling
Arboretum. Thanks to the efforts of a group
of dedicated people, the Arboretum survived,
"Chief among these perhaps, was Donald
Graham, who donated a prodigious amount of
time and unusual knowledge and talent to the
enterprise at a time when it sorely needed such
services." ( The Long Road Traveled , p. 204) In
1943, Donald Graham was significantly involv-
ed in the lobbying that led to a legislative ap-
propriation for support of the Arboretum. In
1963, he was appointed to the "City-University
Arboretum Committee", fonned as a result of
the receipt of Thompson Expressway funds.
He was president of the Arboretum Foundation
in 1941-42 and again in 1962-64.
Donald Graham was a frequent writer
for this publication and a major disseminator of
fine plant material throughout this area. He
donated many rare plants to the Arboretum
which had been brought to this country by his
efforts. He was one of the very few men to be
made an honorary member of the Seattle Garden
Club in recognition of his service to the city.
In 1971 he was awarded the Bronze Medal by
the Seattle Chapter of the American Rhodo-
dendron Society, a chapter he was directly
responsible for organizing. In 1972, the year
of his death, he was to have received the
Amateur Citation Award of the American
Horticultural Society.
26
UW Arboretum Bulletin
The Arboretum Foundation
Invites you to join their
Spring Garden Tour of
South Carolina, North Carolina & Virginia
April 11-25, 1987
Here is an opportunity to see Southern gardens in full bloom. Starting in Charleston,
South Carolina you will see magnificent plantings of evergreen azaleas, effective use of camelias,
magnolias, bald cypress and native plants of the Eastern United States. Public gardens, arboreta
and private gardens will be seen in an area rich in American history. You will be escorted by
Barbara Keightley and Mary Ellen Mulder.
Private gardens from the Charleston House and Garden Tour and the Virginia Historic
Garden Tour will be included. Some of the public gardens that will be seen are Magnolia
Plantation with its 300 year old garden in a superlative water setting and Brookgreen Garden with
its sculpture collection among the native plants. Tyron Palace Restoration shows the transition
in garden design from 17th century Dutch English School to the natural landscaping of the 18th
century. The beautiful North Carolina Botanical Gardens, historic Williamsburg, Monticello,
and a garden designed by Salvatore Dali are some of the highlights.
The motorcoach tour will carry you through the Shenandoah Valley and the Blue Ridge
Mountains to the 12,000 acre Biltmore Estate in Asheville, North Carolina before returning to
Charleston for the return to Seattle.
The cost is $1,920 per person, based on sharing a twin room and a minimum of 20
passengers. A donation to the Arboretum Foundation of $75 for members and $100 for non-
members is required.
For further information please contact: Barbara Keightley, 232-3556; Mary Ellen
Mulder, 232-81 19; Heather at Travel Professionals, 236-0990; or the Arboretum Foundation
office, 325-4510.
A Letter to the Editor
Dear Editor,
I noticed an article in the Fall, 1985
issue of the Arboretum Bulletin that attributed
the design of the south portion of the Japanese
Garden to Richard Yamasaki. To lay to rest
this subject about which there has been con-
fusion for some years I would like to add the
following.
When Juki Iida returned to Japan after
the completion of most of the Garden, it was
apparent to Fred Mann (then University Arch-
itect) that most visitors to the Garden would
walk almost the length of the Garden, from the
parking lot to the south, before they could enter
and the suggestion was made that another en-
trance should be designed at the south end. The
University found funds for this project and ask-
ed me to work directly with Richard Yamasaki
(who build the original gate and teahouse) and
William Yorozu (who set the original rocks and
planting under Mr. Iida's direction). Conse-
quently a collaborative effort was begun to
complete the south portion of the Garden in
sympathy with the major portion of the Garden
already in place.
Richard Yamasaki designed and build
the new entrance gate. The dry stream, general
grading, paths and rock placement were laid out
to my plan and the detailed rock placement and
some of the planting was done by William
Yorozu. Further planting was completed by
Brian Mulligan.
Since the general character of the
Garden designed by Juki Iida was eclectic, that
is combining several styles of Japanese gardens,
it was assumed that the addition was not out of
place although it was not originally designed by
Mr. Iida. -Eric Hoyte
University Landscape Architect
Winter 1986 (49:4)
27
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UW Arboretum Bulletin
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Winter 1986 (49:4)
29
Cleaned seeds ready for the International Seed Exchange at the Center for
Urban Horticulture. photo: Joy Spurr
Published by the
Arboretum Foundation
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University of Washington XD-10
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